Cooperative Fish and Wildlife Research Units Program: Maine
Education, Research and Technical Assistance for Managing Our Natural Resources

Maine Education Activities


Jordan Pond, Acadia National Park.

As part of the mission of the Cooperative Research Units program, the unit scientists participate in graduate education at the host university. Unit scientists supervise graduate student thesis and dissertation research, participate in graduate student advisory committees as chair, co-chair, or committee member, teach graduate level courses, and provide instruction for our cooperators as needed.


Title Instructor(s) Start Date End Date
Wetland and Waterbird Conservation and Management Krohn 2002-09-01 2002-12-30
Waterbird and Wetland Ecology Loftin 2002-09-03 2002-12-14
WLE 650 Graduate Seminar – Fish Physiology Zydlewski 2013-01-01 2013-05-15
WLE 650 Graduate Seminar - Movements and Migrations Zydlewski 2013-01-01 2013-05-15
WLE 591 Movements and Migrations Zydlewski 2012-09-01 2012-12-15
WLE 490 Special Projects in Fisheries Zydlewski 2012-09-01 2012-12-15
Special Problems: GIS Loftin 2002-01-06 2002-05-16
Special Problems in Wildlife Ecology (Independent Study) Zydlewski 2015-01-15 2015-05-15
Seminar Zydlewski 2007-09-01 2007-12-01
Quantitative Freshwater Fisheries Zydlewski 2010-09-01 2010-12-31
Quantitative Freshwater Fisheries Zydlewski 2008-09-01 2008-12-31
Movements and Migrations Zydlewski 2019-09-01 2019-12-31
Movements and Migrations Zydlewski 2017-09-01 2017-12-31
Movements and Migrations Zydlewski 2014-09-01 2014-12-15
Movements and Migration Zydlewski 2009-09-01 2009-12-15
Movements and Migration Zydlewski 2005-09-01 2005-12-15
Modeling Wildlife Habitat Relationships: Current Issues Krohn 2004-01-01 2004-06-15
Modeling Wildlife Habitat Relationships Krohn 2008-01-01 2008-05-09
Integration of remote sensing and geographic information systems in natural resources management Loftin 2013-09-01 2013-12-20
Integration of remote sensing and geographic information systems in natural resources management Loftin 2012-09-01 2012-12-20
Integration of remote sensing and geographic information systems in natural resources management Loftin 2011-09-01 2011-12-20
Integration of remote sensing and geographic information systems in natural resources management Loftin 2010-09-01 2010-12-20
Integration of remote sensing and GIS in natural resources management Loftin 2009-09-01 2009-12-19
Integration of remote sensing and GIS in natural resources management Loftin 2015-09-01 2015-12-15
Integration of remote sensing and GIS in natural resources management Loftin 2014-09-01 2014-11-15
Integration of GIS and remote sensing data in natural resources applications Loftin 2008-09-05 2008-12-15
Integration of GIS and remote sensing data in natural resources applications Loftin 2007-09-05 2007-12-15
Integration of GIS and remote sensing data analysis in natural resource applications Loftin 2020-01-20 2020-05-08
Integration of GIS and remote sensing data analysis in natural resource applications Loftin 2018-09-04 2018-12-14
Integration of GIS and remote sensing data analysis in natural resource applications Loftin 2017-08-29 2017-12-18
Integration of GIS and remote sensing data analysis in natural resource applications Loftin 2016-08-29 2016-12-18
Integration of GIS and remote sensing data analysis in natural resourc Loftin 2004-08-30 2004-12-17
Integration of GIS and Remote Sensing Data in Natural Resource Applica Loftin 2005-09-06 2005-12-23
Graduate Seminar – Fish Physiology Zydlewski 2016-01-01 2016-05-15
Ecology, Evolution, and Complex Systems McKinney 2013-09-03 2013-12-20
Ecology, Evolution, and Complex Systems McKinney 2013-01-05 2013-05-05
Bayesian Study and Methods McKinney 2013-01-05 2013-05-05
Applications of GIS and remote sensing techniques in natural resources Loftin 2001-09-03 2001-12-19
Training Provided Instructor(s) Completion Date
Motorboat Operator's Certification Course (MOCC), U.S. Department of the Interior, Oneida Lake, NY Fisher, Parrish, Zydlewski 2011-05-17
Motorboat Operator's Certification Course (MOCC), U.S. Department of the Interior Zydlewski 2010-05-28
Motorboat Operator's Certification Course (MOCC), U.S. Department of the Interior Zydlewski 2008-05-07
Motorboat Operator's Certification Course (MOCC), U.S. Department of the Interior Zydlewski 2007-05-01
Motorboat Operator's Certification Course (MOCC), U.S. Department of the Interior Zydlewski 2006-05-25
Motorboat Operator Certification Course at Cornell Biological Field Station on Oneida Lake, Bridgeport, NY Zydlewski 2011-05-18
MOCC Boat Safety Certification Course Zydlewski 2013-06-08
Instructor for the Mandela Washington Fellowship Institute, a national leadership program for young African leaders. Zydlewski 2016-07-20
Instructor for the Mandela Washington Fellowship Institute, a national leadership program for young African leaders Zydlewski 2017-07-20
Instructor for federal Motorboat Operator Certification Course at USFWS Craig Brook National Fisheries Program Office. Zydlewski 2017-06-05
Instructor for federal Motorboat Operator Certification Course at USFWS Craig Brook National Fisheries Program Office. Zydlewski 2015-06-05
Instructor for federal Motorboat Operator Certification Course at Acadia National Park. Zydlewski 2014-06-05
Instructor for USGS Over-the-water training at University of Maine Zydlewski 2015-06-01
Instructor for USGS Over-the-water training at University of Maine Zydlewski 2014-06-01
Title and Location Speaker Date
Working in the field of Wildlife Ecology
Presented to: UNiversity of Maine-Expanding Your Horizons Conference
Loftin 2002-03-14
W. B. Krohn, and D., J. Harrison, "Efforts by the University of Maine to Integrate Wildlife Habitats into Land Conservation Planning: A Brief History." Interagency Steering Committee of Maine?s ?Beginning With Habitat? Program,? University of Maine, Orono.
Presented to: n/a/
Krohn 2008-10-17
Using historic wildlife information: promises and pitfalls
Presented to: University of Maine, Wildlife Ecology Seminar Series
Krohn 2004-12-08
Snyder, S., C.S. Loftin, and A.S.Reeve. 2020. Identification, characterization and threat assessment of groundwater dependent ecosystems in the northeastern United States with an integrated GIS-and field survey-based approach. presented in the Wildlife, Fisheries, and Conservation Biology seminar series
Presented to: University of Maine, Orono, ME. web format.
Snyder, Loftin 2020-05-07
Schilling, E.G., C.S. Loftin, and A.D. Huryn, Landscape attributes and invertebrate communities of fishless lakes in Maine.
Presented to: University of Maine, Graduate Limnology class
Loftin 2003-11-06
Lewis, M., C.S. Loftin, and D. Hayes. 2020. Evaluating the Use of Unmanned Aerial Vehicles in Colonial Bird Population Studies; Presentation in the Wildlife, Fisheries, and Conservation Biology seminar series
Presented to: University of Maine
Lewis, Loftin 2020-04-28
Krohn, W. B., D. J. Harrison, and L. L. Robinson. ?Modeling Canada lynx occurrences at three scales across northern Maine.? Meeting of the Advisory Committee, Maine Cooperative Forestry Research Unit, University of Maine, Orono.
Presented to: n/a
Krohn 2008-10-04
Krohn, W. B. The Life and Times of Manly Hardy (1832-1910): Maine Naturalist and Forest Explorer, Maine Forest and Logging Museum, Brewer, Maine.
Presented to: Maine Forest and Logging Museum
Krohn 2007-05-14
Krohn, W. B. Manly Hardy (1832-1910): Maine's first wildlifer? Department of Wildlife Ecology's Noontime Seminar Series, University of Maine, Orono.
Presented to: University of Maine
Krohn 2006-04-24
Krohn, W. B. Manly Hardy (1832-1910): Early Maine Fur-buyer, Hunter, and Naturalist, Brewer Historical Society, Brewer, Maine.
Presented to: n/a
Krohn 2007-05-01
Krohn, W. B. Joshua Gross Rich (1820-1897): Western Maine Pioneer and Wildlife Writer, Bethel Historical Society, Bethel, Maine.
Presented to: Bethel Historical Society
Krohn 2007-09-13
Krohn, W. B. An overview of Maine's largest terrestrial predators: historical patterns, habitat relationships, and current management issues. Two lectures to Senior College, University of Maine, Orono. March 14 and April 4, 2008.
Presented to: University of Maine
Krohn 2008-03-14
Krohn, W. B. "Historical Ecology of the White-tailed Deer in Maine." Munsungan Workshop on Deer Wintering Areas in Maine, Maine Cooperative Forestry Research Unit, Stillwater, ME.
Presented to: n/a
Krohn 2008-12-10
Krohn, W. B. Use of GIS in biodiversity studies for land conservation planning in Maine. Invited lecture and discussion in Integration of GIS and Remote Sensing in Natural Resource Applications (INT 527). University of Maine, Orono, ME.
Presented to: University of Maine
Krohn 2005-09-21
Krohn, W. B. Manly Hardy: the life and writing of a 19th century Maine businessman and naturalist. Invited slide presentation and discussion at the Author's Series, Bangor Public Library, Bangor, ME.
Presented to: Bangor Public Library
Krohn 2005-10-22
Krohn, W. B. Manly Hardy: early sportsman and naturalist from Brewer, Maine. Invited slide presentation to the Penobscot County Conservation Association, Brewer, ME.
Presented to: Penobscot County Conservation Association
Krohn 2005-10-06
Krohn, W. B. Harvest-induced changes in forest age, structure, and composition: effects of recent forestry trends on wildlife habitats in Maine. Invited presentation to the New England Chapter of The Society of American Foresters. Orono, Maine.
Presented to: New England Chapter of The Society of American Foresters
Krohn 2005-10-11
Krohn, W. B. An overview of Maine's largest terrestrial predators: historical patterns, habitat relationships, and current management issues. Two lectures to Senior College, University of Maine, Orono. March 12 and 24, 2006.
Presented to: University of Maine
Krohn 2006-03-12
Kline, L., C.S. Loftin, and D.J.Hayes. 2020. • Using Unmanned Aerial Vehicles and Automated Detection Processes to Monitor Colonial Birds in Maine. Presented at the Wildlife, Fisheries, and Conservation Biology Seminar Series
Presented to: University of Maine
Kline, Loftin 2020-04-14
Joshua Gross Rich (1820-1897): Western Maine Pioneer and Wildlife Writer
Presented to: University of Maine
Krohn 2008-11-17
Human perceptions and ecological effects of fire in Okefenokee Swamp
Presented to: University of Maine
Loftin 2011-02-14
Hoving, C. L., D. J. Harrison, and W. B. Krohn. Canada lynx habitat in eastern North America.
Presented to: University of Maine, Alice Steward Lecture Series
Krohn 2002-10-31
Hepinstall, J. A., D. J. Harrison, and W. B. Krohn. Spatially explicit wildlife habitat modeling: case studies from Maine.
Presented to: University of Maine, Wildlife Ecology Seminar Series
Krohn 2002-04-08
Harrison, D. J., W. B. Krohn, S. Scott, A. K. Fuller, L. Robinson, and C. L. Hoving. Stand-scale management to increase hare and lynx use of managed forests.? Invited presentation at a workshop on Bridging Science and Stewardship sponsored by the Maine Cooperative Forestry Research Unit, Brewer, ME.
Presented to: n/a
Krohn 2008-05-14
Harrison, D. J., W. B. Krohn, A. K. Fuller, J. A. Homyack, and L. L. Robinson. "Temporal and spatial variation in snowshoe hare densities in eastern North America: relationships to lynx and forest management." Meeting of the Advisory Committee, Maine Cooperative Forestry Research Unit, University of Maine, Orono. (Harrison presented).
Presented to: n/a/
Krohn 2008-10-04
GIS Modeling in Landscape Limnology
Presented to: University of Maine Landscape Limnology Class
Loftin 2003-03-06
Conserving freshwater systems in Acadia National Park: an opportunity for reconciliation ecology?
Presented to: Acadia National Park Science Symposium
Loftin 2012-10-23
Combining quantitative approaches and expert opinion to identify Priority Amphibian and Reptile Conservation Areas (PARCAs) and evaluate vulnerability.
Presented to: University of Maine
Loftin 2017-11-15
Combining quantitative approaches and expert opinion to identify Priority Amphibian and Reptile Conservation Areas (PARCAs) and evaluate vulnerability.
Presented to: University of Maine
Loftin 2017-02-13
Combining quantitative approaches and expert opinion to identify Priority Amphibian and Reptile Conservation Areas (PARCAs) and evaluate vulnerability.
Presented to: University of Maine
Loftin 2016-11-16
Changing wildlife populations: the interactions of multiple factors at multiple scales.
Presented to: Wildlife-Habitat Relationships Class(WLE 450), University of Maine
Krohn 2005-04-21
Changes in vegetation distributions in the lower Savannah River tidal marsh following removal of a tidal flap gate
Presented to: University of Maine
Loftin 2004-03-29
Bank, M.S., C.S. Loftin, T.A. Haines, and R.E. Jung.2003. Effects of watershed heterogeneity on mercury bioaccumulation in two-lined salamanders.
Presented to: University of Maine, Mercury Biogeochemistry course
Loftin 2003-11-04
Bank, M.S., C.S. Loftin, T.A. Haines, and R.E. Jung. Mercury, salamanders and watershed condition.
Presented to: Harvard University Forest
Loftin 2003-11-14
Application of Maine Gap Analysis and GIS to land conservation planning
Presented to: University of Maine
Krohn 2005-09-28
An overview of Maine's largest terrestrail predators
Presented to: Senior College, University of Maine
Krohn 2005-04-01
*Zydlewski, J., C.M. Holbrook, D. Gorsky and M.T. Kinnison (2007) Atlantic salmon in the Penobscot River.. Veazie Salmon Club. March 22.
Presented to: Veazie Salmon Club. March 22.
Zydlewski, Holbrook 2007-01-01
*Zydlewski, J., C.M. Holbrook, D. Gorsky and M.T. Kinnison (2007) Atlantic salmon in the Penobscot River. Penobscot Fly Fishing Club. March 7.
Presented to: Penobscot Fly Fishing Club. March 7.
Zydlewski, Holbrook 2007-01-01
Title Presenter Date
Using modeling and GIS in studies of aquatic systems Loftin 2007-02-27
Using ArcGIS in Hydrological Applications Loftin 2003-10-22
Using ArcGIS Geostatistical Analyst in Natural Resources Applications Loftin 2003-10-22
Understanding species-habitat relationships: the importance of a long-term perspective. Invited lecture in WLE 650, Wildlife Habitat Relationships, University of Maine, Orono. Krohn 2010-02-25
Responsible conduct in research: Animal welfare. Loftin 2015-11-04
Responsible conduct in research: Animal welfare Loftin 2019-11-13
Responsible conduct in research: Animal welfare Loftin 2018-11-06
Responsible conduct in research: Animal welfare Loftin 2017-11-07
Responsible conduct in research: Animal welfare Loftin 2016-11-02
Maine’s forest carnivores: historical status, habitat relationships, and current management issues. Senior College, University of Maine, Orono. Krohn 2010-03-19
Landscape Attributes and Invertebrate Communities of Fishless Lakes in Maine Loftin 2005-11-07
GIS Applications in Wetland Ecology Loftin 2019-11-22
GIS Applications in Wetland Ecology Loftin 2018-11-07
GIS Applications in Wetland Ecology Loftin 2017-11-21
Applications of GIS in management of public lands Loftin 2001-04-03
An overview of the University of Maine freshwater mussel research program Loftin 2006-01-21
Student (Degree) Advisor Start Year End Year
York, Geneva (MS) Zydlewski 2015 2017
Woolard, Tyler (MS) Loftin 2018 Present
Wilson, Kristin (PhD) Loftin 2006 2010
Weil, Kristen (MS) Loftin 2013 2015
Wegener, Matthew (MS) Zydlewski 2010 2012
Watson, Jonathan (MS) Zydlewski 2014 2017
Turinetti, Lauren (MS) Zydlewski 2019 Present
Trinko, Tara (MS) Zydlewski 2006 2008
Steinman, Medea (MS) Loftin 2003 2005
Ryan, Kevin (PhD) Zydlewski 2008 2014
Robertson, Ellen (MS) Loftin 2009 2012
Ramberg-Pihl, Nicole (PhD) Zydlewski 2016 Present
Raber, Jen (MS) Loftin 2013 2014
Queheillalt, Dianna (PhD) Loftin 2003 2006
Powell, Luke (MS) Loftin 2005 2008
Popescu, Viorel (PhD) Zydlewski 2007 2011
Politi, Natalia (PhD) Krohn 2004 2008
Perry, Dakota (MS) Loftin 2019 Present
Perkins, Dustin (PhD) Loftin 1999 2004
Peckenham, John (PhD) Loftin 2003 2006
Patrick, David (PhD) Krohn 2004 2007
Palacios-Bianchi, Pilar (PhD) Loftin 2009 2012
Ness, Kirsten (MS) Loftin 2002 2006
Michaud, Corrine (PhD) Loftin 2008 2010
Mansfield, Margo (MS) Loftin 2010 2013
Mallett , David (MS) Krohn 2009 2011
Leppold, Adrienne (PhD) Zydlewski 2014 2016
LeClair, Greg (MS) Loftin 2019 Present
Kramer, David (PhD) Loftin 2005 2006
Knurek, Emily (MS) Loftin 2007 2010
Kneeland, Stephen (MS) Loftin 2003 2006
Kiraly, Ian (MS) Zydlewski 2009 2011
Kelly, Morgan (MS) Loftin 2002 2004
Jordaan, Adrian (PhD) Loftin 2003 2006
Jones, Mitchell (PhD) Zydlewski 2015 2016
Johnston, Catherine (MS) Zydlewski 2013 2016
Jin, Suming (PhD) Krohn 2001 2005
Jackson, Casey (MS) Loftin 2006 2009
Holmes, Vaughn (MS) Zydlewski 2018 Present
Heath, Shane (MS) Loftin 2002 2004
Hearn, Brian (PhD) Krohn 2000 2008
Hayashi, Rei (PhD) Loftin 2011 2016
Hall, Monica (MS) Loftin 2009 2011
Halfyard, Edmund (PhD) Zydlewski 2014 2014
Hachey, Michael (MS) Zydlewski 2005 2007
Gonnerman, Matthew (PhD) Zydlewski 2018 Present
Fuller, Angella (PhD) Krohn 2001 2006
Freeman, Robb (PhD) Loftin 2005 2010
Franzoi Dri, Gabriela (PhD) Loftin 2019 Present
Foss, Carol (PhD) Krohn 1999 2005
Flye, Melissa (MS) Zydlewski 2017 Present
Fleming, Sharon (MS) Zydlewski 2005 2008
Fish, Alex (PhD) Zydlewski 2018 Present
Fernandes, Stephen (MS) Zydlewski 2005 2008
Evans, Bryn (PhD) Loftin 2016 Present
Erdman, Bradley (PhD) Zydlewski 2017 Present
Erbland, Patrick (PhD) Zydlewski 2009 2013
Elsner, Rick (MS) Zydlewski 2017 2017
Duffy, Michelle (MS) Loftin 2019 Present
Douglas, Luke (MS) Loftin 2018 Present
Dionne, Phillip (MS) Zydlewski 2007 2011
Cunningham, Jesse (MS) Loftin 2000 2004
Cline, Brittany (PhD) Zydlewski 2010 2014
Call, Erynn (PhD) Loftin 2009 2015
Call, Erin (PhD) Zydlewski 2009 2013
Boxler, Brandon (MS) Zydlewski 2017 Present
Bolinikjar, Amay (MS) Loftin 2019 Present
Boenish, Robert (PhD) Zydlewski 2015 2018
Bloomquist, Sean (PhD) Loftin 2004 2008
Blomquist, Sean (PhD) Zydlewski 2005 2009
Bailey, Michael (PhD) Zydlewski 2005 2009
Bailey, Jacolyn (PhD) Loftin 2009 2012
Ashe, Wesley (MS) Zydlewski 2009 2012
Ames, Catlin (PhD) Zydlewski 2016 Present
Altenritter, Matthew (PhD) Zydlewski 2010 2015
Allen, Brian (MS) Zydlewski 2015 2017