West Virginia Cooperative Fish and Wildlife Research Unit Staff Member
Stuart A Welsh
Assistant Unit Leader
Phone: (304) 293 - 5006
Email: swelsh@usgs.gov
Faculty Email: swelsh@wvu.edu
Education
- Ph D West Virginia University 1997
- MS Frostburg State University 1994
- BS West Virginia University 1990
Biography
Stuart serves as Assistant Unit Leader at the West Virginia Cooperative Fish and Wildlife Research Unit, West Virginia University, Morgantown, WV, where he conducts research involving graduate students, with an emphasis on mentorship and development of future fisheries professionals.
His research informs management decisions of the state
cooperator (West Virginia Division of Natural Resources), where project results have provided population dynamics and management implications for Blue Catfish, Channel Catfish, Flathead Catfish, Smallmouth Bass, Yellow Perch, and Walleye.
Research projects on fishes or crayfishes also focus on systematic studies for biodiversity recognition, habitat use, movement ecology, species distribution modeling, endangered species, and invasive species. Biodiversity studies have included new species descriptions for four freshwater fishes and six crayfishes. Movement ecology studies have addressed dam passage of the American eel. Species distributions have been modeled for approximately 190 species for an upcoming book on West Virginia fishes. Research on the endangered Candy Darter and Diamond Darter have focused on habitat use and the impacts of invasive species.
Stuart serves the Atlantic States Marine Fisheries Commission with analyses of tagging data relative to management of Atlantic Coast Striped Bass. International research has focused on fishes of Australia, including descriptions of two new species of eel-tailed catfishes.
Areas of Expertise
Aquatic Ecology, Biodiversity, Fisheries Management, Habitat Management, Invasive Species, Movement Ecology, Population Dynamics, Population and Community Ecology, Predator-Prey Dynamics, Species Distribution Modeling, Species Management, Species Status Assessments, Statistics and Modelling, Stream Ecology, T&E Species Management
Taxon Groups Studied
Crayfish, Freshwater Fishes, Gamefish, Invertebrates/Insects, Nongame Fish/Wildlife, Species of Greatest Conservation Need
| Research Publications | Publication Date |
|---|---|
| Zimmerman, J. and S. Welsh. 2012. Prevalence of Anguillicoloides crassus and growth variation in migrant yellow-phase American eels of the upper Potomac River drainage. Diseases of Aquatic Organisms 101:131-137. | November 2012 |
| Zimmerman, J. L., and S. A. Welsh. 2008. Pit tag retention in small (205-370 mm) American eels, Anguilla rostrata. Proceedings of the West Virginia Academy of Sciences 79(2):1-8. | October 2008 |
| Whitaker, J.M., A.B. Welsh, D.W. Hondorp, J.C. Boase, G.T. Merovich, S. Welsh, and C. Krueger. 2018. Variation in DNA methylation is associated with migratory phenotypes of lake sturgeon Acipenser fulvescens in the St. Clair River, MI, USA. Journal of Fish Biology 93:942–951. DOI: 10.1111/jfb.13804 | September 2018 |
| Welsh, S.A., Z.J. Loughman, and T.P. Simon. 2010. Concluding remarks: A symposium on the conservation, biology, and natural history of crayfishes form the southern United States. Southeastern Naturalist 9(Special Issue 3):267-269. | June 2010 |
| Welsh, S.A., K.J. Zipfel, A.W. Peters, D.C. Hoffman and C.M. Layne. 2024. Paddlefish movement and dam passage in the Ohio and Kanawha Rivers, West Virginia. West Virginia Academy of Science 96:12-19. | October 2024 |
| Welsh, S.A., J.L. Aldinger, M.A. Braham, and J.L. Zimmerman. 2016. Synergistic and singular effects of river discharge and lunar illumination on dam passage of upstream migrant yellow-phase American eels. ICES Journal of Marine Science 73:33-42. https://doi.org/10.1093/icesjms/fsv052 | January 2016 |
| Welsh, S.A., D.R. Jerry, D.W. Burrows. 2014. A New Species of Freshwater Eel-tailed Catfish of the Genus Tandanus (Teleostei: Plotosidae) from the Wet Tropics Region of Eastern Australia. Copeia 2014:136–142. https://doi.org/10.1643/CI-13-067 | January 2014 |
| Welsh, S.A., D.R. Jerry, D.W. Burrows, and M.L. Rourke. 2017. A new species of freshwater eel-tailed catfish of the genus Tandanus (Teleostei: Plotosidae) from coastal rivers of mid-northern New South Wales, Australia. Copeia 105:229-236. https://doi.org/10.1643/CI-13-067 | June 2017 |
| Welsh, S.A., D.M. Smith, and N.D. Taylor. 2013. Microhabitat use of the diamond darter. Ecology of Freshwater Fish. doi: 10.1111/eff.12062 | April 2013 |
| Welsh, S.A., D.A. Cincotta, and R.L. Raesly. 2014. First Record of the Bigeye Shiner (Notropis boops) from West Virginia. American Midland Naturalist 172:372–375. https://doi.org/10.1674/0003-0031-172.2.372 | April 2014 |
| Welsh, S.A., D. A. Cincotta, and W.C. Starnes. 2013. First records of Nocomis biguttatus (Hornyhead Chub) from West Virginia discovered in museum voucher specimens. Northeastern Naturalist. Volume 20; DOI 10.1656/045.020.0412. | October 2013 |
| Welsh, S.A. and Z.L. Loughman. 2015. Physical habitat and water quality correlates of crayfish distributions in a mined watershed. Hydrobiologia 745:85–96. https://doi.org/10.1007/s10750-014-2095-y | January 2015 |
| Welsh, S.A. and Z.J. Loughman. 2015. Upstream dispersal of an invasive crayfish aided by a fish passage facility. Management of Biological Invasions 6:287-294. https://doi.org/10.3391/mbi.2015.6.3.07 | July 2015 |
| Welsh, S.A. and Z.J. Loughman. 2015. Upstream dam passage and use of an eel ladder by the common watersnake (Nerodia sipedon). Herpetological Review 46(2):176-179. DOI number not applicable | April 2015 |
| Welsh, S.A. and J.L. Aldinger. 2014. A semi-automated method for monitoring dam passage of upstream migrant yellow-phase American Eels. North American Journal of Fisheries Management 34:702–709. http://dx.doi.org/10.1080/02755947.2014.910580 | July 2014 |
| Welsh, S. and H. Liller. 2013. Environmental correlates of upstream migration of yellow-phase American eels in the Potomac River drainage. Transactions of the American Fisheries Society 142:483-491. | November 2012 |
| Welsh, S. A., and S. A. Perry. 1999. Spatial scale and its influence on estimates of substrate use in stream fishes. North American Journal of Fisheries Management 18:954-959. | July 1999 |
| Welsh, S. A., and S. A. Perry. 1998. Habitat partitioning in a community of darters in the Elk River, West Virginia. Environmental Biology of Fishes 51:411-419. | June 1998 |
| Welsh, S. A., and S. A. Perry. 1997. Acidification and fish occurrence in the upper Cheat River drainage, West Virginia. Journal of the American Water Resources Association 33:423-429. | July 1997 |
| Welsh, S. A., and R. M. Wood. 2008. Crystallaria cincotta, a new species of darter (Teleostei: Percidae) from the Elk River of the Ohio River drainage, West Virginia. Zootaxa 1680:62-68. | January 2008 |
| Welsh, S. A., and D. A. Cincotta. 2004. Natural hybrids of the madtoms Noturus flavus and Noturus insignis from the Monongahela River drainage, West Virginia. Northeastern Naturalist 11:399-406. | October 2004 |
| Welsh, S. A., S. M. Eyler, M. F. Mangold, and A. J. Spells. 2002. Capture locations and growth rates of Atlantic sturgeon in the Chesapeake Bay. Pages 183-194 in W. Van Winkle, P. J. Anders, D. H. Secor, and D. A. Dixon, editors. Biology, management, and protection of North American sturgeon. American Fisheries Society Symposium 28, Bethesda, Maryland. | June 2002 |
| Welsh, S. A., R. M. Wood, K. R. Sheehan. 2009. Threatened fishes of the world: Crystallaria cincotta Welsh and Wood 2008 (Percidae). Environmental Biology of Fishes 84:191-192. | January 2009 |
| Welsh, S. A., M. F. Mangold, J. E. Skjeveland and A. J. Spells. 2002. Distribution and movement of shortnose sturgeon (Acipenser brevirostrum) in the Chesapeake Bay. Estuaries 25:101-104. | February 2002 |
| Welsh, S. A., M. F. Mangold and S. M. Eyler. 2001. Movements of shortnose sturgeon in the upper Chesapeake Bay, Maryland. Proceedings of the Southeastern Association of Fish and Wildlife Agencies 55:298-303. . | August 2001 |
| Welsh, S. A., J. T. Anderson, and D. A. Cincotta. 2002. Fish occurrence on the Camp Dawson Collective Training Area, Kingwood, West Virginia. Proceedings of the West Virginia Academy of Natural Sciences 74:87-89 | August 2002 |
| Welsh, S. A., D. R. Smith, R. W. Laney, and R. C. Tipton. 2007. Tag-based estimates of annual fishing mortality of a mixed coastal stock of Atlantic striped bass. Transactions of the American Fisheries Society 136:34-42. | January 2007 |
| Welsh, S. A., A. W. Kahnle, B. A. Versak, and R. J. Latour. 2003. Use of tag data to compare growth rates of Atlantic coast striped bass stocks. Fisheries Management and Ecology. | October 2003 |
| Thompson, P.A., S.A. Welsh, M.P. Strager, and A.A. Rizzo. 2018. A multiscale investigation of habitat use and within-river distribution of sympatric sand darter species. Journal of Geospatial Applications in Natural Resources , Volume 2, Issue 1, Article 1. | January 2018 |
| Thompson, P.A., S.A. Welsh, A.A. Rizzo, and D.M. Smith. 2017. Effect of substrate size on sympatric sand darter benthic habitat preferences. Journal of Freshwater Ecology 32:455-465. http://dx.doi.org/10.1080/02705060.2017.1319880 | April 2017 |
| Switzer, J. F., S. A. Welsh, T. King. 2008. Microsatellite DNA primers for the candy darter, Etheostoma osburni and variegate darter, Etheostoma variatum, and cross-species amplification in other darters (Percidae). Molecular Ecology Resources 8:335-338. (doi: 10. 1111/j. 1471-8286. 2007. 01946. x) | January 2008 |
| Smith, J., S.A. Welsh, J.T. Anderson, and R. Fortney. 2015. Water quality trends in the Blackwater River watershed, West Virginia. Northeastern Naturalist 14(Special Issue 7):103-111. https://doi.org/10.1656/058.014.sp711 | October 2015 |
| Smith, J., S.A. Welsh, J.T. Anderson, and R. Fortney. 2009. Water quality trends (1980-1993) in the Blackwater River watershed, West Virginia. In Canaan Valley and its Environs: A Landscape Heritage Celebration. Canaan Valley Institute, Davis, West Virginia. http://www.canaanvi.org/canaanvi_web/events_ed.aspx?collection=past%20events&id=826 | April 2009 |
| Smith, D.M., S.A. Welsh, and P.J. Turk. 2012. Available benthic habitat type may influence predation risk in larval lampreys. Ecology of Freshwater Fishes 21:160–163. | May 2012 |
| Smith, D.M., S.A. Welsh, and P.J. Turk. 2011. Selection and preference of benthic habitat by small and large ammocoetes of the least brook lamprey (Lampetra aepyptera). Environmental Biology of Fishes 91:421-428. | August 2011 |
| Smith, D.M., S.A. Welsh, and C. HIlling. 2021. Seasonal Movement Patterns and Distribution of Walleye in a Central Appalachian Hydropower Reservoir. Pages 209-237 in J. Bruner and R. DeBruyne (eds.) Biology, Management, and Culture of Walleye, Sauger, and Yellow Perch: Status and Needs, Springer | November 2021 |
| Smith, D., Welsh, S. and Hilling, C. (n.d.). Environmental Correlates of Walleye Spawning Movements in an Appalachian Hydropower Reservoir. Journal of the Southeastern Association of Fish and Wildlife Agencies, [online] 10, pp.36–44. Available at: https://seafwa.org/sites/default/files/journal-articles/J10_05_Smith%20et%20al%2036-44.pdf | January 2023 |
| Smith, D. M., C. D. Hilling, S. A. Welsh, and D. I. Wellman, Jr. 2022. Differences in population characteristics and modeled response to harvest regulations in reestablished Appalachian Walleye populations. North American Journal of Fisheries Management 46:612–629. | June 2022 |
| Siegel, J.V., S.A. Welsh, N.D. Taylor, and Q.E. Phelps. Size Structure, Age, Growth, and Mortality of Flathead Catfish in the Robert C. Byrd Pool of the Ohio and Kanawha Rivers. Journal of the Southeastern Association of Fish and Wildlife Agencies 10:10-16. | January 2023 |
| Sheehan, K.R. and S.A. Welsh. 2021. Comparison of geographically weighted regression of benthic substrate modeling accuracy on large and small wadeable streams. Journal of Geographic Information System 13:194-209. DOI: 10.4236/jgis.2021.132011 | April 2021 |
| Sheehan, K., and S. A. Welsh. 2009. An interpolation method for stream habitat assessments. North American Journal of Fisheries Management 29:1-9. | April 2009 |
| Sheehan, K., M. Strager, and S. Welsh. 2013. Advantages of geographically weighted regression for modeling benthic substrate in two Greater Yellowstone Ecosystem Streams. Environmental Modeling and Assessment. DOI 10.1007/s10666-012-9334-2 | April 2013 |
| Sheehan, K. and S. Welsh. 2013. Accuracy of stream habitat interpolations across spatial scales. Journal of Geographic Information System 5:602-612. | October 2013 |
| Selego, S.M., C.L. Rose, G.T. Merovich, S.A Welsh, and J.T. Anderson. 2012. Community-level response of fishes and aquatic macroinvertebrates to stream restoration in a third-order tributary of the Potomac River, USA. International Journal of Ecology, DOI 10.1155/2012/753634. | April 2012 |
| Ruble, C.L., P.L. Rakes, J.R. Shute, and S.A. Welsh. 2016. Captive propagation, reproductive biology, and early life history of Etheostoma wapiti (Boulder Darter), E. vulneratum (Wounded Darter), and E. maculatum (Spotted Darter). Southeastern Naturalist 15:115-126. https://doi.org/10.1656/058.015.0109 | October 2016 |
| Ruble, C.L., P.L. Rakes, J.R. Shute, and S.A. Welsh. 2014. Captive propagation, reproductive biology, and early life history of the Diamond Darter (Crystallaria cincotta). American Midland Naturalist 172:107–118. https://doi.org/10.1674/0003-0031-172.1.107 | January 2014 |
| Rizzo, A.A., S.A. Welsh, and P.A. Thompson. 2017. A paired-laser photogrammetric method for in situ length measurement of benthic fishes. North American Journal of Fisheries Management 37:16-22. http://dx.doi.org/10.1080/02755947.2016.1235632 | January 2017 |
| Rizzo, A.A., D.J. Brown, S.A. Welsh, and P.A. Thompson. 2017. Factors influencing detection of the federally endangered Diamond Darter Crystallaria cincotta: implications for long-term monitoring strategies. American Midland Naturalist 178:123-131. https://doi.org/10.1674/0003-0031-178.1.123 | April 2017 |
| Rizzo, A.A., C.T. Rota, P.A. Thompson, D.J. Brown, and S.A. Welsh. 2018. Effects of an Extreme Flood Event on Federally Endangered Diamond Darter Abundances. American Midland Naturalist 180:108-118. | July 2018 |
| Osier, E. A., and S. A. Welsh. 2007. Habitat Use of the Spotted Darter, Etheostoma maculatum, in Elk River, West Virginia. Northeastern Naturalist 14:447-460. | September 2007 |
| Millard, M., S. Welsh, J. Fletcher, J. Mohler, A. Kahnle, K. Hattala. 2003. Mortality associated with catch and release of striped bass in the Hudson River. Fisheries Management and Ecology. | October 2003 |
| Matt, K.J., S.A. Welsh, and D.M. Smith. 2021. Spawning characteristics of Yellow Perch during periods of water level fluctuations in a hydropower reservoir. Pages 3-32 in J. Bruner and R. DeBruyne (eds.) Biology, Management, and Culture of Walleye, Sauger, and Yellow Perch: Status and Needs, Springer | November 2021 |
| Loughman, Z.L., S.A. Welsh, and R. Thoma. 2017. Cambarus (C.) appalachiensis, a new species of crayfish (Decapoda: Cambaridae) from the New River Basin of Virginia and West Virginia, USA. Zootaxa 4243:432-454. http://dx.doi.org/10.11646/zootaxa.4243.3.2 | March 2017 |
| Loughman, Z.L., A.M. Sykes, M.I. McKinney, and S.A. Welsh. 2021. Epigean crayfish of the Potomac River basin in West Virginia: Zoogeography, natural history, and conservation. Freshwater Crayfish 26:37-49. | April 2021 |
| Loughman, Z.J., and S.A. Welsh. 2010. Distribution and conservation standing of West Virginia crayfishes. Southeastern Naturalist 9(Special Issue 3):63-78. | June 2010 |
| Loughman, Z.J., T.P. Simon, and S.A. Welsh. 2011. Cambarus (Puncticambarus) smilax, a new species of crayfish (Crustacea; Decapoda: Cambaridae) from the Greenbrier River basin of West Virginia. Proceedings of the Biological Society of Washington 124:99-111. | May 2011 |
| Loughman, Z.J., T.P. Simon, and S.A. Welsh. 2010. Forward: conservation, biology, and natural history of crayfishes from the southern United States. Southeastern Naturalist 9(Special Issue 3):1-10. | June 2010 |
| Loughman, Z.J., S.A. Welsh, and T.P. Simon. 2012. Occupancy rates of primary burrowing crayfish in natural and disturbed large river bottomlands. Journal of Crustacean Biology 32:557–564. | April 2012 |
| Loughman, Z.J., S.A. Welsh, N.M. Sadecky, Z.W. Dillard, and R.K. Scott. 2017. Evaluation of physiochemical and physical habitat associations for Cambarus callainus (Big Sandy Crayfish), an imperiled crayfish endemic to the Central Appalachians. Aquatic Conservation: Marine and Freshwater Ecosystems 2017:1-9. DOI: 10.1002/aqc.2746 | January 2017 |
| Loughman, Z.J., S.A. Welsh, N.M. Sadecky, Z.W. Dillard, and R.K. Scott. 2016. Environmental covariates associated with Cambarus veteranus Faxon, 1914 (Decapoda: Cambaridae), an imperiled Appalachian crayfish endemic to West Virginia, USA. Journal of Crustacean Biology 36:642-648. https://doi.org/10.1163/1937240X-00002456 | December 2016 |
| Loughman, Z.J., S.A. Welsh, J.W. Fetzner, and R.F. Thoma. 2015. Conservation of imperiled crayfish species - Cambarus veteranus (Decapoda: Reptantia: Cambaridae). Journal of Crustacean Biology 35:850-860. https://doi.org/10.1163/1937240X-00002383 | October 2015 |
| Loughman, Z.J., R. Fagundo, S.A. Welsh, E. Lau, and R.F. Thoma. 2013. Cambarus (C.) hatfieldi, a new species of crayfish (Decapoda:Cambaridae) from the Tug Fork River Basin of Kentucky, Virginia and West Virginia, USA. Zootaxa. | July 2013 |
| Loughman, Z.J., D. Foltz, and S.A. Welsh. 2013. Baited lines, an active nondestructive collection method for burrowing crayfish. Southeastern Naturalist 12:809-815. | October 2013 |
| Loughman, Z., D. Foltz, N. Garrison, and S. Welsh. 2013. Cambarus theepiensis sp. n., a new species of crayfish (Crustacea, Decapoda, Cambaridae) from the coalfields region of Eastern Kentucky and Southwestern West Virginia, USA. Zootaxa 3641:63-73. | April 2013 |
| Loughman, Z. J., Simon, T. P., and S. A. Welsh. 2009. West Virginia crayfishes (Decapoda:Cambaridae): Observations on distribution, natural history, and conservation. Northeastern Naturalist 16:225-238. | January 2009 |
| Loughman, Z, S. Welsh, and R. Thoma. Cambarus fetzneri sp. nov., a new species of burrowing crayfish (Decapoda: Cambaridae) from the Allegheny Mountains of Virginia and West Virginia. Zootaxa 4651 (1): 038–050. https://doi.org/10.11646/zootaxa.4651.1.2 | August 2019 |
| Kinziger, A. P., R. M. Wood, and S. A. Welsh. 2001. Systematics of Etheostoma tippecanoe and Etheostoma denoncourti (Perciformes: Percidae). Copeia 2001:235-239. | October 2001 |
| Hilling, C.D., N.D. Taylor, S.A. Welsh, and D.M. Smith. Population Characteristics of Yellow Perch in a Central Appalachia Hydropower Reservoir. Journal of Fish and Wildlife Management 9(2):486–495. doi:10.3996/012018-JFWM-007 | Abstract | December 2018 |
| Hilling, C. D., S. A. Welsh, and D. M. Smith. 2016. Age, growth and fall diet of channel catfish in Cheat Lake, West Virginia. Journal of Fish and Wildlife Management 7:304-314. https://doi.org/10.3996/092015-JFWM-091 | April 2016 |
| Hedrick, L.B., S.A. Welsh, and J.T. Anderson. 2007. Effects of highway construction on sediment and benthic macroinvertebrates in two tributaries of the Lost River, West Virginia. Journal of Freshwater Ecology 22:561-569. | June 2007 |
| Hedrick, L.B., S.A. Welsh, J.T. Anderson, L. Lin, Y. Chen, and X. Wei. 2010. Response of benthic macroinvertebrate communities to highway construction in an Appalachian watershed. Hydrobiologia 641:115-131. | March 2010 |
| Hedrick, L., S. Welsh, and J. Anderson. 2009. Influences of high flow events on a stream channel altered by construction of a highway bridge – a case study. Northeastern Naturalist 16:375-394. | December 2009 |
| Hedrick, L., J. Anderson, S. Welsh, and L-S. Lin. 2013. Sedimentation in mountain streams: a review of methods of measurement. Natural Resources 4:92-104. | January 2013 |
| Hedrick, L. B., S. A. Welsh, and J. T. Anderson. 2007. Evaluation of a new sediment sampling device. Proceedings of the Annual Conference of the Southeastern Association of Fish and Wildlife Agencies 61:35-39. | December 2007 |
| Hedrick, L. B., S. A. Welsh, and J. D. Hedrick. 2005. A New Sampler Design for Measuring Sedimentation in Streams. North American Journal of Fisheries Management 25:238-244. | January 2005 |
| Hammond, S.D. and S.A. Welsh. 2009. Seasonal movements of large yellow American eels downstream of a hydroelectric dam, Shenandoah River, West Virginia. Pages 309-323 in J.M. Casselman and D.K. Cairns, editors. Eels at the edge: science, status, and conservation concerns. American Fisheries Society Symposium 58, Bethesda, Maryland. | August 2009 |
| Gibson, I., A. Welsh, S. Welsh, and D. Cincotta. 2018. Genetic swamping and species collapse: tracking introgression between the native Candy Darter and introduced Variegate Darter. Conservation Genetics 20:287–298. doi.org/10.1007/s10592-018-1131-2 | Abstract | December 2018 |
| Foltz, D.A., N. Sadecky, G.. Meyers, J. Fetzner, S. Welsh, G. Stocker, M. Glon, and R. Thoma. 2018. Cambarus loughmani, a New Species of Crayfish (Decapoda: Cambaridae) Endemic to the Pre-glacial Teays River Valley in West Virginia, USA. Journal of Natural History 52:45-46, 2875-2897, DOI:10.1080/00222933.2018.1557271 | December 2018 |
| Eyler, S.M., S.A. Welsh, D.R. Smith and M.M. Rockey. 2016. Downstream passage and impact of turbine shutdowns on survival of silver American eels at five hydroelectric dams on the Shenandoah River. Transactions of the American Fisheries Society 145:964-976. http://dx.doi.org/10.1080/00028487.2016.1176954 | October 2016 |
| Cincotta, D.A., and S.A. Welsh. 2010. Discovery of Ammocrypta clara (western sand darter) in the upper Ohio River of West Virginia. American Midland Naturalist 163:318-325. | April 2010 |
| Cincotta, D.A., D.P. Wegmen, T.E. Oldham, S.A. Welsh, and L.B. Hedrick. 2015. Fishes of the Blackwater River drainage, Tucker County, West Virginia. Northeastern Naturalist 14(Special Issue 7):297-313. https://doi.org/10.1656/058.014.sp725 | October 2015 |
| Cincotta, D.A., D.P. Wegmen, T.E. Oldham, S.A. Welsh, and L.B. Hedrick. 2009. Fishes of the Blackwater River drainage, Tucker County, West Virginia. In Canaan Valley and its Environs: A Landscape Heritage Celebration. Canaan Valley Institute, Davis, West Virginia. http://www.canaanvi.org/canaanvi_web/events_ed.aspx?collection=past%20events&id=826 | April 2009 |
| Cincotta, D.A. and S.A. Welsh. 2024. An Update of the Ichthyofauna of West Virginia with Notes on Historic Sportfish Stockings. Northeastern Naturalist 31:(monograph 24):1-48. | July 2024 |
| Chen, Y., X. Wei, L. Lin, R. Viadera, L.B. Hedrick, S.A. Welsh, J.T. Anderson. 2009. Effects of highway construction on stream water quality and macroinvertebrate condition in a Mid-Atlantic highlands watershed, USA. Journal of Environmental Quality 38:1672-1682. | April 2009 |
| Braham, M.A., S.A. Welsh, D.M. Smith. 2024. An experimental study of benthic habitat selection in yellow-phase American eels (Anguilla rostrata). Environmental Biology of Fishes 107:513-522. | June 2024 |
| Argentina, J.E., P.L. Angermeier, E.M. Hallerman, and S.A. Welsh. 2018. Spatial extent of analysis influences observed patterns of population genetic structure in a widespread darter species (Percidae). Freshwater Biology 63: 1185–1198. | April 2018 |
| Aldinger, J.L. and S.A. Welsh. 2017. Diel periodicity and chronology of upstream migration in yellow-phase American eels (Anguilla rostrata). Environmental Biology of Fishes 100:839-838. DOI 10.1007/s10641-017-0614-1 | April 2017 |
| Presentations | Presentation Date |
|---|---|
| Whitaker, J. A. Welsh, D. Hondorp, J. Boase, S. Welsh, and G. Merovich. 2015. Characterization of the Migratory Phenotype in Lake Sturgeon. Presented at the 71st Annual Northeast Fish and Wildlife Meeting, New Port, Rhode Island. | April 2015 |
| Welsh, S.A., K. Zipfel, A. Peters, C. Hoffman, and C. Layne. 2024. An Assessment of Movement and Dam Passage of Paddlefish in the Ohio River, West Virginia. Meeting of the West Virginia Academy of Science, Glenville, WV, 6 April 2024. | April 2024 |
| Welsh, S.A., D.R. Jerry, and D.W. Burrows. A New Species of Freshwater Eel-tailed Catfish of the Genus Tandanus (Teleostei: Plotosidae) from the Wet Tropics Region of Eastern Australia. Annual Joint Meeting of Ichthyologists and Herpetologists, Chattanooga, TN, 1 Aug 2014. | August 2014 |
| Welsh, S.A., D. Jerry, D. Burrows, and M. Rourke. 2017. A New Species of Freshwater Eel-tailed Catfish of the Genus Tandanus (Teleostei: Plotosidae) from Coastal Rivers of Mid-northern New South Wales, Australia. Joint Meeting of Ichthyologists and Herpetologists | July 2017 |
| Welsh, S.A. and Z.J. Loughman. A Nonnative Crayfish (Faxonius virilis) Use of an Eel Ladder, Potomac River Drainage, USA . Meeting of the International Association of Astacologists, Pittsburgh, PA, 10 July 2018. | July 2018 |
| Welsh, S.A. and Z.J. Loughman. 2016. Use of an eel pass by virile crayfish on the lower Shenandoah River. Annual Meeting of the West Virginia Academy of Science, Huntington, WV, 9 April 2016 | April 2016 |
| Welsh, S.A. and Z.J. Loughman. 2015. Upstream dispersal of an invasive crayfish aided by a fish passage facility. Presented at the Southern Division American Fisheries Society meeting, Savannah, GA | January 2015 |
| Welsh, S.A. and S.E. Welsh. 2017. Smoke screen foraging: an observation of an unusual foraging behavior of the northern pike, Esox lucius. Annual Meeting of the West Virginia Academy of Science | April 2017 |
| Welsh, S.A. and J.L. Aldinger. 2016. Patterns and periodicity of upstream passage of American Eels at an eel ladder. Annual Meeting of the Southern Division American Fisheries Society, Wheeling, WV, 20 February 2016 | February 2016 |
| Welsh, S.A. and J.L. Aldinger. 2015. Upstream migrant American Eels: A 12-year time series from an eel pass on the lower Shenandoah River. Presented at the 145th Annual Meeting of the American Fisheries Society, 19 August 2015, Portland, OR. | August 2015 |
| Welsh, S.A. and J.L. Aldinger. 2015. Periodicity and timing of upstream migration of yellow-phase American Eels, Potomac River drainage, USA. The 95th Annual Meeting of the American Society of Ichthyologists and Herpetologists. University of Nevada, Reno, Nevada, 19 July 2015. | July 2015 |
| Welsh, S.A. and J. L. Aldinger. 2014. A Photographic Counter for Monitoring American Eels at an Eel Ladder. Annual meeting of the Southern Division of the American Fisheries Society, Charleston, SC, 23 January 2014. | January 2014 |
| Welsh, S., and D. Wichterman. 2013. Angler and creel surveys of the walleye fishery at Summersville Reservoir, West Virginia. Joint meeting of the Ohio and WV Chapters of the American Fisheries Society, 20 February 2013, Huntington, West Virginia. | February 2013 |
| Welsh, S., S. Hammond, H. Liller, J. Zimmerman, M. Braham, and J. Aldinger. 2014. An 11-year series of passage counts of upstream migrant American Eels at the Millville Dam eel ladder, lower Shenandoah River, USA. 144th annual meeting of the American Fisheries Society, Quebec City, Canada | August 2014 |
| Welsh, S., D. Smith, and N. Taylor. 2013. Nocturnal microhabitat use of the diamond darter. Annual meeting of the Southern Division American Fisheries Society, 9 February 2013, Nashville, Tennessee. | February 2013 |
| Welsh, S. and J. Aldinger. A 12-year time series of American eel passage at an eel ladder, lower Shenandoah River. Presented at the joint meeting of VA and WV Chapters of AFS, Blacksburg, VA. | February 2015 |
| Welsh, S. K. Matt, and D. Smith. Spawning Characteristics of Yellow Perch During Periods of Water Level Fluctuations in a Hydropower Reservoir. Annual Meeting of the Southeastern Association of Fish and Wildlife Agencies, Charleston, West Virginia, 23-26 October 2022. | October 2022 |
| Thompson, P., S. Welsh, A. Rizzo. 2016. Substrate selection of sympatric sand darters from the Elk River, WV. Annual Meeting of the West Virginia Academy of Science, Huntington, WV, 9 April 2016 | April 2016 |
| Thompson, P., S. Welsh, A. Rizzo. 2016. A tale of two darters: Substrate selection and preference of sympatric sand darters from the Elk River, WV. Annual Meeting of the Southern Division of the American Fisheries Society, Wheeling, WV, 20 February 2016 | February 2016 |
| Taylor, N., S. Welsh, and D. Smith. 2013. Assessment of age, diet, and growth of yellow perch In Cheat Lake, West Virginia. Annual meeting of the Southern Division American Fisheries Society, 10 February 2013, Nashville, Tennessee. | February 2013 |
| Taylor, N., S. Welsh, and D. Smith. 2013. Age structure, growth, and summer diet of yellow perch (Perca flavescens) in a West Virginia Reservoir. Joint meeting of the Ohio and WV Chapters of the American Fisheries Society, 20 February 2013, Huntington, West Virginia. | February 2013 |
| Smith, D.M., S.A. Welsh, and C. Hilling. 2020. Seasonal Movement and Distribution of Walleye in a WV hydropower reservoir. American Fisheries Society, Virtual Annual Meeting | September 2020 |
| Smith, D.M., S.A. Welsh, N.D. Taylor, and C.D. Hilling. 2016. Environmental and Biological Correlates of Seasonal Walleye Movements in a Hydropower Reservoir. Annual Meeting of the Southern Division of the American Fisheries Society, Wheeling, WV, 20 February 2016 | February 2016 |
| Smith, D.M., S.A. Welsh, N.D. Taylor, and C.D. Hilling. 2015. Population Dynamics and Seasonal Movement Patterns of Walleye in a Hydropower Reservoir. Presented at the Southern Division American Fisheries Society meeting, Savannah, GA. | January 2015 |
| Smith, D.M., S.A. Welsh, N.D. Taylor, and C.D. Hilling. 2015. Population Dynamics and Seasonal Movement Patterns of Walleye in a Hydropower Reservoir. Presented at the Joint meeting of VA and WV AFS Chapters, Blacksburg, VA | February 2015 |
| Smith, D., S. Welsh, and N. Taylor. 2013. Investigating Seasonal Movements and Spawning Locations of Walleye in Cheat Lake, WV using Acoustic Telemetry. Annual meeting of the Southern Division American Fisheries Society, 10 February 2013, Nashville, Tennessee. | February 2013 |
| Smith, D., S. Welsh, and N. Taylor. 2013. Determining Seasonal Movements and Spawning Habitat Use of Walleye in Cheat Lake, WV using Acoustic Telemetry. Joint meeting of the Ohio and WV Chapters of the American Fisheries Society, 20 February 2013, Huntington, West Virginia. | February 2013 |
| Smith, D., S. Welsh, N. Taylor, and C. Hilling. 2014. Use of Acoustic Telemetry to Determine Seasonal Movements of Walleye in a West Virginia Hydropower Reservoir. Annual meeting of the Southern Division of the American Fisheries Society, Charleston, SC, 23 January 2014. | January 2014 |
| Siegel, J.V., S.A. Welsh, N.D. Taylor, and B.A Murry. Population Structure, Age, and Growth of Flathead and Blue Catfish within the Robert C. Byrd Pool of the Ohio River. West Virginia Chapter of the American Fisheries Society, virtual meeting. | April 2021 |
| Siegel, J., S. Welsh, N. Taylor, and Q. Phelps. Size Structure, Age, Growth, and Mortality of Flathead Catfish in the Robert C. Byrd Pool of the Ohio and Kanawha Rivers. Annual Meeting of the Southeastern Association of Fish and Wildlife Agencies, Charleston, West Virginia, 23-26 October 2022. | October 2022 |
| Rizzo, A., and S. Welsh. 2016. Photogrammetric analysis of fish lengths. Annual Meeting of the Southern Division of the American Fisheries Society, Wheeling, WV, 20 February 2016 | February 2016 |
| Rizzo, A., S. Welsh, P. Thompson. 2016. Photogrammetric analysis of fish lengths. Annual Meeting of the West Virginia Academy of Science, Huntington, WV, 9 April 2016 | April 2016 |
| Matt, K.J., S.A. Welsh, and D.M. Smith. 2020. Spawning characteristics of Yellow Perch during periods of water level fluctuations in a hydropower reservoir. American Fisheries Society, Virtual Annual Meeting. | September 2020 |
| Loughman, Z.L. and S.A. Welsh. 2016. Physical Habitat and Water Quality Correlates for Cambarus veteranus in the Upper Guyandotte Basin of West Virginia. Annual Meeting of the Southern Division American Fisheries Society, Wheeling, WV, 20 February 2016 | February 2016 |
| Loughman, Z, N. Sadecky, and S. Welsh. 2015. Determination of Crayfish Occupancy Rates Across Mined Watersheds in Eastern Kentucky. Presented at the Southern Division American Fisheries Society meeting, Savannah, GA | January 2015 |
| Hilling, C.D., D.M. Smith, and S.A. Welsh. 2020. Percids Kicking Acid: Biology of Recovered Walleye and Yellow Perch Populations in a Historically Acidified Hydropower Reservoir. American Fisheries Society, Virtual Annual Meeting. | September 2020 |
| Hilling, C, S. Welsh, and D. Smith. 2016. Population characteristics of Channel Catfish in Cheat Lake, West Virginia. Annual Meeting of the Southern Division of the American Fisheries Society, Wheeling, WV, 20 February 2016 | February 2016 |
| Hilling, C, S. Welsh, and D. Smith. 2015. Evaluation of age, growth and condition of Channel Catfish in Cheat Lake, West Virginia. Presented at the Joint Meeting of VA and WV AFS Chapters, Blacksburg, VA | February 2015 |
| Harrell, J., S. Welsh, D. Smith, C. Arantes, and D. Wellman. Smallmouth bass population characteristics and an evaluation of management responses in two West Virginia watersheds. Annual Meeting of the Southeastern Association of Fish and Wildlife Agencies, Charleston, West Virginia, 23-26 October 2022. | October 2022 |
| Eyler, S., S. Welsh, and D. Smith. 2015. Seasonal turbine shutdowns used to protect downstream migrating silver American Eels (Anguilla rostrata) at five hydroelectric dams on the Shenandoah River. Presented at the 71st Annual Northeast Fish and Wildlife Meeting, New Port, Rhode Island. | April 2015 |
| Eyler, S., S. Welsh, D. Smith, and M. Rockey. 2015. Passage method, turbine mortality, and migratory delay of silver American Eels (Anguilla rostrata) at five hydroelectric dams on the Shenandoah River. Presented at the Connecticut River Atlantic Salmon Commission (CRASC) Research Forum, MADFW Westborough Field Office, Westborough, MA | March 2015 |
| Eyler, S., S. Welsh, D. Smith, and M. Rockey. 2015. Environmental variables associated with American Eel (Anguilla rostrata) downstream migrations in the Shenandoah River. Presented at the American Fisheries Society Meeting, Portland, Oregon. | August 2015 |
| Aldinger, J. and S. Welsh. Nocturnal periodicity of upstream migration of yellow-phase American Eels at an eel ladder on the Shenandoah River. Presented at the Southern Division American Fisheries Society meeting, Savannah, GA | January 2015 |
| Technical Publications | Publication Date |
|---|---|
| Welsh, S.A., Skjeveland, J.E., M.F. Mangold. 2000. Progress report of investigations and research on Atlantic and shortnose sturgeon in Maryland waters of the Chesapeake Bay (1996-1998). U.S. Fish and Wildlife Service, Maryland Fisheries Resource Office, Annapolis, MD, 15 June 1999. | June 1999 |
| Welsh, S.A., D.A. Cincotta, and J.F. Switzer. 2006. Fishes of Bluestone National Scenic River. National Park Service, Natural Resources Technical Report NPS/NER/NRTR--2006/049. | April 2006 |
| Welsh, S.A. 2004. Overestimation of tag-based fishing mortality rates by monotonic trend models: examples from simulated and real data. Report submitted to the ASMFC Striped Bass Tagging Committee, 1 October 2004. | October 2004 |
| Welsh, S.A. 2001. Annual tag-encounter and annual exploitation rates of Atlantic striped bass estimated from tag recoveries (1987 - 2000). Report to the ASMFC Striped Bass Tagging Committee, Striped Bass Stock Assessment Committee, and Striped Bass Technical Committee. 45 pp. | June 2001 |
| Welsh, S.A. 2000. Summary of data from the York River (1991-1999) of the USFWS Cooperative Tagging Program. Report to the ASMFC Striped Bass Tagging Committee, 25 August 2000. | August 2000 |
| Welsh, S.A. 2000. Summary of data from the North Carolina Winter Trawl Survey (1988 -1999) of the USFWS Cooperative Tagging Program. Report to the ASMFC Striped Bass Tagging Committee, 15 August 2000. | August 2000 |
| Welsh, S.A. 2000. Length-frequency data of tagged cohorts from the Cooperative Striped Bass Tagging. Program Report to the ASMFC Striped Bass Tagging Committee, Striped Bass Stock Assessment Committee, and Striped Bass Technical Committee, 2 April 2000. | April 2000 |
| Welsh, S.A. 2000. Annual tag-encounter rates and annual exploitation rates of Atlantic striped bass estimated from tag-recoveries: spring tagging programs (1987-1999). Report to the ASMFC Striped Bass Tagging Committee, Striped Bass Stock Assessment Committee, and Striped Bass Technical Committee, 25 August 2000. | August 2000 |
| Welsh, S.A. 2000. Annual tag-encounter rates and annual exploitation rates of Atlantic striped bass estimated from tag-recoveries. Report to the ASMFC Striped Bass Tagging Committee, Striped Bass Stock Assessment Committee, and Striped Bass Technical Committee, 2 April 2000. | April 2000 |
| Welsh, S., R. Harris, B. Versak, D. Kahn, T. Baum,V. Vecchio, K. Hattala, and G. Nelson. 2003. ASMFC tag working group: summary of USFWS cooperative tagging program results. Submitted to the Technical and Stock Assessment Committees, Atlantic States Marine Fisheries Commission. | September 2003 |
| Welsh, S., R. Harris, B. Versak, D. Kahn, T. Baum,V. Vecchio, K. Hattala, and G. Nelson. 2002. ASMFC tag working group: summary of USFWS cooperative tagging program results. Submitted to the Technical and Stock Assessment Committees, Atlantic States Marine Fisheries Commission. | June 2002 |
| Skjeveland, J.E., S.A. Welsh, M.F. Mangold, S.M. Eyler, S.W. Nachbar. 2000. A report of investigations and research on Atlantic and shortnose sturgeon in Maryland waters of the Chesapeake Bay. U.S. Fish and Wildlife Service, Maryland Fisheries Resource Office, Annapolis, MD, 18 October 2000. 44 pages. | October 2000 |
| Millard, M.J., S.A. Welsh, J. Skjeveland, J. Fletcher, J. Mohler, M. Hendrix, A. Kahnle, and K. Hattala. 2000. Mortality associated with catch and release of American shad and striped bass in the Hudson River. Final report: 1999 project year. Submitted to NY State Department of Environmental Conservation, New Paltz, NY 12561. | December 2000 |
| Eyler, S.M., J.E. Skjeveland, M.F. Mangold, and S.A. Welsh. 2000. Distribution of sturgeons in candidate open water dredged material placement sites in the Potomac River (1998-2000). Final Report. U.S. Fish and Wildlife Service, Maryland Fisheries Resource Office, Annapolis, MD, 21 November 2000. 26 pages. | November 2000 |
| Annual tag-encounter and annual exploitation rates of Atlantic striped bass estimated from tag recoveries (1987-2001). Report to the ASMFC Striped Bass Stock Assessment Commitee, and Striped Bass Technical Committee. | August 2002 |
| Role and Organization | Individual | Start Date | End Date |
|---|---|---|---|
| Southeastern Fishes Council - Member | Welsh | June 2001 | Present |
| Southeastern Association of Fish and Wildlife Agencies - Member, Program Committee | Welsh | January 2001 | December 2002 |
| Southeastern Association of Fish and Wildlife Agencies - Chair, Best Student Paper Award Committee | Welsh | September 2007 | December 2007 |
| International Association of Astacology - Member | Welsh | January 2011 | Present |
| American Society of Ichthyologists and Herpetologists - Student paper judge | Welsh | April 1993 | June 1993 |
| American Society of Ichthyologists and Herpetologists - Member, Best Student Paper Award committee | Welsh | July 2001 | July 2001 |
| American Society of Ichthyologists and Herpetologists - Member | Welsh | August 1993 | Present |
| American Fisheries Society - President-elect, West Virginia Chapter | Welsh | April 2012 | April 2013 |
| American Fisheries Society - President, West Virginia Chapter | Welsh | April 2013 | April 2015 |
| American Fisheries Society - Member, West Virginia Chapter | Welsh | May 2000 | Present |
| American Fisheries Society - Member | Welsh | August 1993 | Present |
| American Fisheries Society - Chair, Student Affairs Committee, Southern Division | Welsh | August 2002 | May 2008 |
| American Fisheries Society - Chair, Annual Meeting, West Virginia Chapter | Welsh | August 2002 | August 2003 |