Cooperative Fish and Wildlife Research Units Program: Florida
Education, Research and Technical Assistance for Managing Our Natural Resources

Florida Staff Member


Dr. Raymond Raoul Carthy

Dr. Raymond R. Carthy

Assistant Unit Leader
Phone: (352) 846 - 0545
Email: rayc@usgs.gov
Faculty Email: ngosi@ufl.edu
Faculty Website

Education

  • Ph D University of Florida 1996
  • MS Slippery Rock University 1983
  • BS Manhattan College 1980

Biography

Raymond Carthy - Assistant Unit Leader, Florida Cooperative Fish and Wildlife Research Unit

Dr. Carthy joined the Florida Unit in 1996 after receiving a Ph.D. in Zoology from the University of Florida. His previous employment included work at the Smithsonian Institution’s National Museum of Natural History and volunteer and consulting work for marine conservation NGOs. Dr. Carthy's research centers on the ecology of threatened and endangered species, and his interests involve reproductive ecology and physiology of coastal and wetland herpetofauna, with a focus on marine and freshwater turtles. Current projects include examining effects of climate change and habitat degradation on nesting and in-water marine turtle species. He is also involved in research on threatened upland species and in conservation management-oriented studies. Dr. Carthy is the Program Director of the University of Florida Uncrewed Aircraft Systems Research Program (UFUASRP), a multi-disciplinary endeavor studying the use of UAS for wildlife and habitat surveys and other natural resource conservation applications. In addition to teaching herpetology classes centered on ecology and conservation of marine chelonids, Dr. Carthy has taught courses on bioethics and critical thinking.

Areas of Expertise

Anthropogenic Impacts, Aquatic Ecology, Habitat Management, Marine/Coastal Ecology, Species Management, T&E Species Management, UAVs, Wetland Ecology, Wildlife Management

Taxon Groups Studied

Amphibians, Marine Mammals, Reptiles, Sea Turtles

Research Publications Publication Date
Wooding, J. B., and R. R. Carthy. 2000. Interactions of humans and manatees in Crystal River, Florida. Proceedings of the Southeastern Association of Fish and Wildlife Agencies, 52:260-264. September 2000
Williams,N.C., Bjorndal, K. A., Lamont, M.M., Carthy, R.R. 2014. Winter diets of immature green turtles (Chelonia mydas) on a northern feeding ground: integrating stomach contents and stable isotope analyses. Estuaries and Coasts. 37:986-994. | Download July 2014
Wilkinson, B., H. A. Lassiter, A. Abd-Elrahman, R. R. Carthy, P. Ifju, E. Broadbent, and N. Grimes. 2019. Geometric targets for UAS Lidar. Remote Sensing 11(24), 3019; https://doi.org/10.3390/rs11243019 December 2019
Tiwari, M., R. R. Carthy, A. Silveria, and K. A. Bjorndal. 2003. A comparison of nest architecture in loggerhead sea turtles (Caretta caretta) nesting in Florida and Brazil. Herpetological Review, Vol. 34(2):138-139. August 2003
Swindall, J.E., H.K. Ober, M.M. Lamont and R.R. Carthy. 2019. Informing sea turtle outreach efforts to maximize effectiveness. Wildlife Society Bulletin 1–11; 2019; DOI: 10.1002/wsb.1004 | Download September 2019
Silver-Georges, I., S. A. Ceriani,M. Ware, M. Lamb, M. Lamont, J. Becker, R.R. Carthy, C, Matechik, J. Mitchell, R. Pruner, M. Reynolds, B. Smith, C. Snyder, and M. M. P. B. Fuentes. 2021. Using systems thinking to inform management of imperiled species: a case study with sea turtles. Biological Conservation 260: 109201 | Abstract | Publisher Website June 2021
Shamblin, B.M., M.G. Dodd, D.A. Bagley, L.M. Ehrhart, A.D. Tucker, C. Johnson, R.R. Carthy, R.A. Scarpino, E. McMichael, D.S. Addison, K.L. Williams, M.G. Frick, S. Ouellette, A.B. Meylan, M.H. Godfrey, S.R. Murphy, C.J. Nairn. Genetic structure of the southeastern United States loggerhead turtle nesting aggregation: evidence of additional structure within the peninsular Florida recovery unit. 2010. Marine Biology (2011) 158:571-587 December 2010
Shamblin, B.M., A.B. Bolten, K.A. Bjorndal, P.H. Dutton, J.T. Nielsen, F.A. Abreu-Grobois, K.J. Reich. B.E. Witherington, D.A. Bagley, L.M. Ehrhart, A.D. Tucker, D.S. Addison, A. Arenas, C. Johnson, R.R. Carthy, M.M. Lamont, M.G. Dodd, M.S. Gaines, E. LaCasella, and C.J. Nairn. 2012. Expanded mitochondrial control region sequences increase resolution of stock structure among North Atlantic loggerhead turtle rookeries. Marine Ecology Progress Series,. 469:145-160. | Abstract July 2012
Senko, J., V. Koch, W.M. Megill, R.R. Carthy, R.P. Templeton, W.J. Nichols. 2010. Fine scale daily movements and habitat use of East Pacific green turtles at a shallow coastal lagoon in Baja California Sur, Mexico. Journal of Experimental Marine Biology and Ecology. 391(2010): 92-100 July 2010
Salmon, M., R. Carthy, C. M.F. Lohmann, K. J. Lohmann, and J. Wyneken. 2012. Collecting a sample of loggerhead sea turtle hatchlings before a natural emergence does not reduce nest productivity. Endangered Species Research. 16: 296-299. doi: 10.3354/esr00409 IP-034556 | Download March 2012
Rees, A. F., L. Avens, K. Ballorain, E. Bevan, A. C. Broderick, R. R. Carthy, M. J. Marjolijn, J. A. Christianen, G. Cuclos, M. R. Heithaus, D. W. Johnston, J . C. Mangel, F. Paladino, K. Pendoley, R. D. Reina, N. J. Robinson, R. Ryan, S. T. Sykora-Bodie, D. Tilley, M. R. Varela, E. R. Whitman, P. A. Whittock, T. Wibbels, and B. J. Godley. 2018.The potential of unmanned aerial systems for sea turtle research and conservation: a review and future directions. Endangered Species Research. 35: 81-100. February 2018
Prosdocimi, L., N. Teryda, G. Navarro, and R. R. Carthy. 2020. Use of remote sensing tools to predict focal areas for sea turtle conservation in the Southwestern Atlantic. Aquatic Conservation: Marine and Freshwater Ecosystems, 2020:1–11. DOI: 10.1002/aqc.3478 | Abstract December 2020
Phillips, A.C.N., J. Coutou, S. Rajh, N. Stewart, A. Watson, A. Jehu, H.Asmath, F. Dziva, C. Unakal R.B. Holder, and R.R. Carthy. 2017. Temporospatial dynamic and public health significance of bacterial flora identified on a major leatherback turtle (Dermochelys coriacea) nesting beach in the Southern Caribbean. Marine Ecology. 15pp IP- 068594. https://doi.org/10.1111/maec.12412 | Abstract | Publisher Website April 2017
Parris, L., M. M. Lamont, and R. R. Carthy. 2002. Increased incidence of red imported fire ant (Hymenoptera: formicidae) presence in loggerhead sea turtle (testudines: cheloniidae) nests and observations of hatchling mortality. Florida Entomologist, Vol. 85(3):514-517. September 2002
Nomani, S. Z., R. R. Carthy and M. K. Oli. Comparison of methods for estimating abundance of gopher tortoises. Applied Herpetology. 5:13-31 January 2008
Nomani, S. Z., M. K. Oli, and R.R. Carthy. 2012. Line transects by design: the influence of study design, spatial distribution and density of objects on estimates of abundance. The Open Ecology Journal. 5: 25-44. | Abstract October 2012
McMichael, E., R.R. Carthy, J.A. Seminoff and R.A. Scarpino. Kemp's ridley turtles in St. Joseph Bay, Florida, USA: Results from an in-water study in the northern Gulf of Mexico. NOAA Technical Memorandum NMFS-SEFSC-630 May 2012
McMichael, E., J. Seminoff, R. Carthy. Kemp's ridley sea turtles in St. Joseph Bay, Florida, USA: results from an in-water study in the northeastern Gulf of Mexico. 2009. Journal of Natural History. 42 (43-44; 2293-2307). December 2009
McMichael, E., J. A. Seminoff and R. R. Carthy. Growth rates of wild green turtles, Chelonia mydas, at a temperate foraging habitat in the northern Gulf of Mexico: assessing short-term affects of cold stunning on growth. Journal of Natural History. December 2008
Manes, C., R. R. Carthy, and V. Hull. 2023. A coupled human and natural systems framework to characterize emerging infectious diseases- the case of fibropapillomatosis in marine turtles. Animals 2023, 13(9), 1441; https://doi.org/10.3390/ani13091441 | Abstract April 2023
Manes, C., R. R. Carthy, R. M. Herren, D. J. Duffy, J. A. Farrell, A. Page-Karjian, C. Skibicki, F. Dunlap, and I. Capua. 2023. Green turtle Fibropapillomatosis: tumor morphology and growth rate in a rehabilitation setting. Vet. Sci. 2023, 10(7), 421; https://doi.org/10.3390/vetsci10070421 | Abstract July 2023
Maglothin, M. L., M. M. Lamont, and R. R. Carthy. 2002. Nest Chamber anomalies in loggerhead sea turtle (Caretta caretta) nests from Northwest Florida, USA. Herpetological Review., Vol. 33(1):48-50. August 2002
Layton, J.E., T. Wibbels, T. Tucker, J. Wyneken, L. Ehrhart, R. Carthy, E.R. Martin, M. Bressette, C.Johnson, S. Fournier, and J. Schmid. 2009. Developing a Comprehensive Long-term Database on Nesting Beach Temperatures of the Loggerhead Sea Turtle in the Southeastern U.S: Applications and Implications for Global Climate Change. Integrative and Comparative Biology. 49:E227-E227 | Publisher Website October 2009
Lamont, M.M., Fujisaki, I., Carthy, R.R. 2014. Estimates of vital rates for a declining loggerhead turtle (Caretta caretta) subpopulation: implications for management. Marine Biology. 161:2659–2668 DOI 10.1007/s00227-014-2537-0 | Abstract September 2014
Lamont, M. M., and R. R. Carthy. 2007. Response of nesting sea turtles to barrier island dynamics. Chelonian Conservation and Biology 6(2): 206-212 January 2007
Lamont, M. M. D. Johnson, and R. R. Carthy. 2020. The incubation environment of nests deposited by a genetically distinct group of loggerhead sea turtles in Northwest Florida. Global Ecology and Conservation 23: https://doi.org/10.1016/j.gecco.2020.e01070 | Abstract | Publisher Website September 2020
Hirama, S., B. Witherington, K. Kneifl, A. Sylvia, M. Wideroff, and R. R. Carthy. 2021. Environmental factors predicting the orientation of sea turtle hatchlings on a naturally lighted beach: A baseline for light-management goals. Journal of Experimental Marine Biology and Ecology 541 (2021) 151568. | Abstract May 2021
Hirama, S. B. Witherington, S. Hirsch, A. Sylvia, and R. R. Carthy. 2023. Accuracy and precision of hatchling sea-finding orientation as a function of dune proximity in hatchlings of two species of sea turtles. Marine and Freshwater Research (IF 1.8) , DOI:10.1071/mf23052. | Abstract June 2023
Hill, M. K., M. C. Monroe, T. T. Ankersen, R. R. Carthy, and T. A. Kay. 2019. Conservation easements and coastal armoring: protecting sea turtle nesting habitat through property ownership., Ocean and Coastal Management, https://doi.org/10.1016/j.ocecoaman.2019.104944 September 2019
Hill, M. K., M. C. Monroe, T. T. Ankersen, R. R. Carthy, T. A. Kay. 2019. Coastal Armoring and Sea Turtles: Beachfront Homeowners’ Opinions and Intent. Coastal Management, DOI: 10.1080/08920753.2019.1669102 September 2019
Hart, K.M., M. Lamont, I. Fujisaki, A.D. Tucker, R.R. Carthy. Common coastal foraging areas in the Gulf of Mexico used by nesting loggerheads from 3 different subpopulations: Opportunities for marine conservation. Published.Biological Conservation. (145) 1:185-194. 2012. January 2012
Gross, T. S., D. A. Crain, K. A. Bjorndal, A. B. Bolten, and R. R. Carthy. 1995. Identification of sex hatchling loggerhead turtles (Caretta caretta) by analysis of steriod concentrations in chorioallantoic/amniotic fluid. Gen. Comp. Endocrinol. 99:204-210. March 1995
Gregory, C. J., R. R. Carthy, and L. G. Pearlstine. 2006. Application of Species at Risk at Camp Blanding Training Site, FLARNG(Florida). Southeastern Naturalist. 5(3):473-498. January 2006
Gonzalez Perez, A., B. Wilkinson. A. Abd-Elrahman, R. R. Carthy, and D.J. Johnson. 2022. Deep and machine learning image classification of coastal wetlands using unpiloted aircraft system multispectral images and lidar datasets. Remote Sensing, 14(16), 3937; https://doi.org/10.3390/rs14163937 | Abstract August 2022
Goforth, K. M. and R.R. Carthy. 2022. Tidally-driven gas exchange in beaches: implications for sea turtle nest success. Journal of Coastal Research. 38 (3): 523–537, https://doi.org/10.2112/JCOASTRES-D-21-00082.1 | Abstract February 2022
Garland, K.A. and R.R. Carthy. 2010. Changing taste preferences, market demands and traditions in Pearl Lagoon, Nicaragua: a community reliant on Chelonia mydas for income and nutrition. Conservation and Society 8(1):55-72. April 2010
Gallaway, Benny J. ,William J. Gazey, Charles W. Caillouet, Jr., Pamela T. Plotkin, Donna J. Shaver, F. Alberto Abreu Grobois, Anthony M. Amos, Patrick M. Burchfield, Ray R. Carthy, Marco A. Castro Martínez, John G. Cole, Andrew T. Coleman, Melissa Cook, Steven DiMarco, Sheryan P. Epperly, Allen M. Foley, Masami Fujiwara, Daniel Gomez Gamez, Gary L. Graham, Wade L. Griffin, Selina S. Heppell, Francisco Illescas Martínez, Jackie Isaacs, Margaret M. Lamont, Rebecca L. Lewison, Kenneth J. Lohmann, James M. Nance, Jonathan Pitchford, Nathan F. Putman, Scott W. Raborn, Jeffery K. Rester, Jack J. Rudloe, Laura Sarti Martínez, Mark Schexnayder, Jeffrey R. Schmid, Christopher Slay, Mandy Tumlin, Thane Wibbels and Blanca M. Zapata Najera. 2017. Development of a Kemp’s Ridley Sea Turtle Stock Assessment Model. Gulf of Mexico Science. | Publisher Website February 2017
Fujisaki, I., M. Lamont and R.R. Carthy. 2018.Temporal shift of loggerhead nest sites in a highly eroding beach in a dynamic barrier island. Ocean and Coastal Management. 155 (2018) 24–29. https://doi.org/10.1016/j.ocecoaman.2017.12.032 April 2018
Evans, D. R., R. A. Valverde, C. Ordóñez , and R. R. Carthy. 2021. Identification of the Gulf of Mexico as an important high-use habitat for leatherback turtles from Central America. Ecosphere 12(8):e03722. 10.1002/ecs2.3722 May 2021
Evans, D. R., L. Pemberton, and R. R. Carthy. 2024. Wide ranging dispersal of hawksbill sea turtles (Eretmochelys imbricata) from nesting beaches on the island of Nevis. Marine Biology, 171:171; https://doi.org/10.1007/s00227-024-04491-6 | Abstract August 2024
Evans, D. R. , R. R. Carthy, and S. A. Ceriani. 2019. Migration routes, foraging behavior, and individual fidelity of loggerhead sea turtles (Caretta caretta) satellite tracked from a globally important rookery. Marine Biology, 166:134 https://doi.org/10.1007/s00227-019-3583-4. | Publisher Website September 2019
Diaz, A. L., A. E. Ortega, H. Tingle, A. Pulido, O, Cordero, M. Nelson, N. Cocoves, J. Shin. R. R. Carthy, B. E. Wilkinson, and P. G. Ifju. 2022, The Bathy-drone: An autonomous Uncrewed drone-tethered sonar system. Drones 6(10), 294; https://doi.org/10.3390/drones6100294 | Abstract October 2022
Carthy, R.R. The leatherbacks of Grande Riviere Trinidad and Tobago. 2012. Florida Environmental Outreach. (3), 1- 10-13. IP-034506 December 2012
Carthy, R. R., A. M. Foley, and Y. Matsuzawa. 2002. The incubation environment of nests of the loggerhead turtle (Caretta caretta) and effects of hatching success and the characteristics of hatchlings. Book Chapter, for "The Biology of the Loggerhead Sea Turtle". Smithsonian Institution Press, August 2003. August 2003
Carthy, R. R. 2016. The leatherback turtle: Biology and Conservation. Herpetological Review, 47(4):703-705. December 2016
Carthy, R. R. 2006. An owner's manual for the Chelonioidea. Conservation Biology. 20(1)259-260. February 2006
Carthy, R. R. 1992. Scanning electron microscopy (SEM) of loggerhead (Caretta caretta) eggshell structure. In: M. Salmon and J. Wyneken (compilers), Proceedings of the 11th Annual Workshop on Sea Turtle Biology and Conservation. NOAA Technical Memorandum NMFS-SEFSC-302:143-144. January 1992
Bjorndal, K.A., Bolten, A.B., Chaloupka, M., Saba, V.S., Bellini, C., Marcovaldi, M.A.G., Santos, A.J.B., Bortolon, L.F.W., Meylan, A.B., Meylan, P.A., Gray, J., Hardy, R., Brost, B., Bresette, M., Gorham, J.C., Connett, S., Crouchley, B.V.S., Dawson, M., Hayes, D., Diez, C.E., van Dam, R.P., Willis, S., Nava, M., Hart, K.M., Cherkiss, M.S., Crowder, A.G., Pollock, C., Hillis-Starr, Z., Muñoz Tenería, F.A., Herrera-Pavón, R., Labrada-Martagón, V., Lorences, A., Negrete-Philippe, A., Lamont, M.M., Foley, A.M., Bailey, R., Carthy, R.R., Scarpino, R., McMichael, E., Provancha, J.A., Brooks, A., Jardim, A., López-Mendilaharsu, M., González-Paredes, D., Estrades, A., Fallabrino, A., Martínez-Souza, G., Vélez-Rubio, G.M., Boulon Jr., R.H., Collazo, J.A., Wershoven, R., Guzmán Hernández, V., Stringell, T.B., Sanghera, A., Richardson, P.B., Broderick, A.C., Phillips, Q., Calosso, M., Claydon, J.A.B., Metz, T.L., Gordon, A.L., Landry Jr., A.M., Shaver, D.J., Blumenthal, J., Collyer, L., Godley, B.J., McGowan, A., Witt, M.J., Campbell, C.L., Lagueux, C.J., Bethel, T.L., and Kenyon, L., 2017, Ecological regime shift drives declining growth rates of sea turtles throughout the West Atlantic. 2017. Global Change Biology, 23(11):4556-4568, http://dx.doi.org/10.1111/gcb.13712. | Abstract | Publisher Website June 2017
Bishop, N., Polisar, J., Eliazar, P.J., Carthy, R.R., & Bjorndal, K.A. 2022. Diet of Dermatemys mawii, an aquatic turtle that relies heavily on terrestrial vegetation. Chelonian Conservation and Biology. 21 (1): 37–45. https://doi.org/10.2744/CCB-1467.1 | Abstract March 2022
Bishop, N. D., Hudson, R., Marlin, J., Pop, T., Rainwater, T., Boylan, S., Atkinson, B., and Carthy, R. R. 2021. Using growth rates to estimate the minimum age and size at sexual maturity in a captive population of the critically endangered Central American river turtle Dermatemys mawii. Journal of Zoo and Aquarium Research, 9(3), 150–156. https://doi.org/10.19227/jzar.v9i3.432 | Abstract July 2021
Askren, V., R. Carthy, C. Chase, G. Comstock, E. Franz and T. Oriola. 2002. More Citation Vigilance. The Scientist, Vol. 16(17):14. August 2002
Project Completion Date
Pre-assessment Plan to Determine Potential Exposure and Injuries of Nesting and Hatching Loggerhead Sea Turtles July 2011
Incubation temperatures of loggerhead turtle (caretta caretta) nests On NW Florida Beaches March 2012
Gopher Tortoise (gopherus polyphermus)Population Survey For St. Marks NWR-Line Transect Distance Sampling August 2011
Observing Flight Initiation Distances of Florida Chelonia mydas in the Crystal Bay Area October 2022
St. Joseph Peninsula beach restoration project. October 2008
Assessment of beach compaction and associated effects on loggerhead sea turtles nesting on natural and nourished beaches in northwest Florida. September 2008
Assessing the effects of coastline alteration on sea turtle nesting and faunal assemblages at Cape San Blas, Florida. March 2009
Living Green: Living with sea turtles. April 2010
Assessment of coastal habitats and listed species. March 2010
Spectral and response assessment of wildlife-friendly lighting. April 2010
Effects of climate change on barrier island habitat and nesting sea turtles. May 2012
Environmental correlates of fibropapilloma in green turtles. December 2025
Assessing effects of anthropogenic and climate-induced change on health of coastal ecosystems. December 2026
Effects of Coastal Dynamics & Climate on Loggerhead Turtle N... May 2018
Evaluation of the impact of artificial lighting on sea turtle hatchling orientation. December 2020
Nutritional ecology and husbandry of Dermatemys mawii August 2020
Green turtle spatial distribution, abundance and habitat models in the Northeastern Gulf of Mexico. August 2021
Integration, Validation and fusion of Small Unmanned Aircraft systems August 2019
Composition, distribution and ecology of the Nature Coast sea turtle assemblage August 2020
D. Duke Conservation Scholars Program Partnership Through UF September 2023
Aguilar, Trenton
PhD
trenton.aguilar@ufl.edu

Broxton, Jazmyn
Master's
jbroxton@ufl.edu

Harlow, Jaiere
Master's
jharlow1@ufl.edu

Herren, Richard
PhD
rherren@ufl.edu

Manes, Constanza
PhD
c.manes@ufl.edu

Smith, Rachel
PhD
jrachel.smith@ufl.edu

Teryda, Natalia
PhD
nteryda@ufl.edu

Presentations Presentation Date
Zhang, X.M. N. Kemal, B. Smith, N. Bishop and R. Carthy. 2015. Establishing baseline monitoring data for sea turtle nesting beaches with varying anthropogenic usage in St. Johns County, Florida. 100th Meeting, Ecological Society of America, Baltimore, Maryland, 9-14 August 2015. August 2015
Zeitoune, S., R. McKee, R. McCleery, C. Romagosa, and R. R. Carthy. 2023. Prey naiveté or pre-adaptation in cotton rats facing invasive pythons. Poster presentation at The Wildlife Society meeting, Louisville, Kentucky. November 2023
Vilchez, M., L Nicholson, L. Braun de Torrez , R. Carthy, and C. Romagosa. 2021.The implications of Acoustic Software Match Ratios for bat species identification and composition. Poster presentation, ESA, Long Beach, California. August 2021
Trujillo, N., M. McCampbell, C. Gulick, C. Romagosa, and R.R. Carthy. 2022. Assessment of cotton rat <i>(Sigmodon hispidus</i>) home range sizes in the Greater Everglades Ecosystem. Poster presentation at 29th Annual Meeting of TWS, Spokane, WA. November 2022
Teryda, N. S., L. Prosdocimi, E. A. Pinheiro dos Santos, G. Stahelin, M. Lopez-Mendilaharzu, and R. R. Carthy. 2025. SURFACE DETECTION PROBABILITY OF LEATHERBACK TURTLES AT THE RIO DE LA PLATA, SOUTH AMERICA. Oral presentation at 43rrd Annual International Sea Turtle Symposium, Accra, Ghana. March 2025
Teryda, N. S., L. Prosdocimi, G. M. Velez-Rubio, and R. R. Carthy. 2020. Unmanned aircraft system surveys target leatherback conservation in the Rio de la Plata Estuary, AR/UR. Poster presentation, 40th International Symposium on Sea Turtle Biology and Conservation, Cartagena, Colombia, March 14-20, 2020. March 2022
Teryda, N. S., G. M. Velez-Rubio, L. Prosdocimi, and R. R. Carthy. 2023. Uncrewed Aerial Systems as tools for green turtle population assessment in coastal marine protected areas in Uruguay. Poster presentation at International Sea Turtle Symposium, Cartagena, Colombia. March 2023
Summo, S., L. Nicholson, C. Romagosa, and R.R. Carthy.2022. Watering holes - An exploration of the relationship between proximity to different types of open water bodies and bat activity in south Florida. Poster presentation at 29th Annual Meeting of TWS, Spokane, WA. November 2022
Stratton, L. D., E. M. Trotman, C. N. Keiser, C. M. Romagosa, and R. R. Carthy. 2023. Food provisioning and juvenile survival in the fishing spider, <i>Dolomedes triton. </i>Poster presentation at The Wildlife Society meeting, Louisville, Kentucky. November 2023
Smith, R., E. Hardin, and R. R. Carthy. 2025. rust, Traps, and Turtles: Leveraging Stakeholder and Fisher Perspectives to Enhance Sea Turtle Bycatch Reporting and Mitigation in Florida’s Trap Fisheries. Oral presentation at 43rd Annual International Sea Turtle Symposium, Accra, Ghana. March 2025
Smith, R., E. Hardin, C. J. Bohannon, S. Schaf, and R. R. Carthy. 2025.Unraveling Sea Turtle Bycatch in Florida's Trap Fisheries: An Interdisciplinary Threat Assessment. Oral presentation. South East Regional Sea Network Meeting, Hilton Head, SC. March 2025
Smith, R. S., S. Schaf, and R. R. Carthy. 2023. Entanglement of marine turtles in Florida’s commercial and recreational trap fisheries: 30+ years of data reveal a potentially ubiquitous and persistent threat. Oral presentation at International Sea Turtle Symposium, Cartagena, Colombia. March 2023
Silver-Gorges, I., J. Becker, R. R. Carthy, S. A. Ceriani, M. Lamb, M. M. Lamont, C. Matechik, J. Mitchell, M. Reynolds, B. Smith, C. Snyder, M. Ware, M. M. P. B. Fuentes. 2020. Maximizing Loggerhead Turtle (Caretta caretta) Hatchling Production in Light of Disturbances in Northern Gulf of Mexico. Gulf of Mexico Oil Spill & Ecosystem Science Conference (GoMOSES). February 3-6, 2020, Tampa, FL. February 2022
Siddiqui, S., N. Bishop and R.Carthy. 2016. Nutrient Influxes from Caretta caretta (loggerhead sea turtle) nesting. Annual Meeting, Ecological Society of America, Fort Lauderdale, Florida, 7-12 August, 2016. August 2016
Shigetomo Hirama, Morgan Wideroff. Raymond R. Carthy, and Blair Witherington . 2018. Measuring the impact of light pollution on hatchling orientation. Presentation at the 38th International Symposium on Sea Turtle Biology and Conservation, Kobe, Japan. Feb. 18-24, 2018 February 2018
Sherr, E., W.W. Boone, C. M. Romagosa, R.R. Carthy and N.D. Bishop. 2017. Interactions between ctenophores (Mnemiopsis leidyi) and microplastic beads within ecologically relevant parameters. Ecological Society of America, Portland, Oregon. August 2017
Serano, J., T.S. Coleman, N. Bishop, T. Osborne, and R. R. Carthy. 2023. Measuring tidally-driven gas exchange on sea turtle nesting beaches. Poster presentation at Li-COR Connect 2023, Atlanta, GA. November 2023
Selden, J., N. Bishop, B. Smith and R. Carthy. 2015. Natural inventory and habitat classification of the beach-to-estuary ecotones and establishment of a long-term monitoring site for barrier island habitats and species in St. Johns County, Florida. 100th Meeting, Ecological Society of America, Baltimore, Maryland, 9-14 August 2015. August 2015
Roy, F., L. Elmquist, J. Broxton, L. Hussain, P. Xiong, R. R. Carthy, C. Romagosa, and R. Fletcher. 2024. Potential boat interactions between recreational boating and snail kite (<i>Rostramus sociabilis plumbeus</i>) nesting. Poster presentation at The Wildlife Society meeting, Baltimore Maryland. October 2024
Rodriguez-Riverol, J. J., E. M. Trotman, C. N. Keiser, C. M. Romagosa, and R. R. Carthy. 2023. Parasite infection and antipredator behavior in freshwater snails. Poster presentation at The Wildlife Society meeting, Louisville, Kentucky. November 2023
Quintilani, M., N. bishop, B.J. Smith,C. Romagosa and R. Carthy. 2016. Effects of man-made oyster reefs on sediment composition in estuaries. Annual Meeting, Ecological Society of America, Fort Lauderdale, Florida, 7-12 August, 2016. August 2016
Prosdocimi, L., N. S. Teryda, G. M. Velez-Rubio, and R. R. Carthy. 2020. Unmanned Aerial Vehicles: Assessing use for surveys of sea turtle populations in feeding areas. Poster presentation, 40th International Symposium on Sea Turtle Biology and Conservation, Cartagena, Colombia, March 14-20, 2020. March 2022
Pedrozo, C., J. Brisbane, B.J. Smith, C. Romagosa, N. Bishop and R. Carthy. 2016. Predicting patterns of native snake occurrence along Main Park Road in Everglades National Park, Florida. Annual Meeting, Ecological Society of America, Fort Lauderdale, Florida, 7-12 August, 2016. August 2016
Martucci, E., L. Elmquist, L. Hussain, P. Xiong, J. Broxton, R. R. Carthy, C. Romagosa, and R. Fletcher. 2024. Mapping boat activity: Comparing boat counts between peak recreational events and snail kite surveys. Poster presentation at The Wildlife Society meeting, Baltimore Maryland. October 2024
Manes, C., D. Pinton, R. Carthy, A. Canestrelli, and I. Capua. 2023. Occurrence of Fibropapillomatosis in Green Turtles (<i>Chelonia mydas</i>) in Relation to Environmental Changes in Coastal Ecosystems in Texas and Florida: A Retrospective Study. Oral presentation at International Sea Turtle Symposium, Cartagena, Colombia.<b><br><b> </b></b> March 2023
Manes, C. R. Herren, D. Godfrey, E. Cooper, and R. R. Carthy. 2025. Fibropapillomatosis elusive virus ChHV5 reveals novel multifactorial relationships in the Northeastern Gulf of Mexico. Oral presentation. South East Regional Sea Network Meeting, Hilton Head, SC. March 2025
Lopez Romero, A., R. Buron, R. Carthy , and C. Romagosa. 2021. Using eBird to determine the effects of urbanization on breeding Great Horned Owls (<i>Bubo virginianus</i>). Poster presentation, ESA, Long Beach, California. August 2021
Innocent, H. M. Celestin, N.D. Bishop, C. M. Romagosa, R.R. Carthy, W.W. Boone. 2017. Shrub encroachment in a south Florida grassland: implications for mammalian inhabitants. Ecological Society of America, Portland, Oregon. August 2017
Herren, R., D. R. Evans, and R. R. Carthy. 2025. Juvenile Green Turtle Overwintering Behavior in the Northeastern Gulf of Mexico. Oral presentation. South East Regional Sea Network Meeting, Hilton Head, SC. February 2025
Herren, R., D. Evans, D. Godfrey, and R. R. Carthy. 2023. Green turtle overwintering behavior in the northeastern Gulf of Mexico. Oral presentation at International Sea Turtle Symposium, Cartagena, Colombia. March 2022
Gonzalez, D., P. Xiong, L. Hussain, J. Broxton, R. R. Carthy, and C. Romagosa. 2024. Assessing non-target species interference in camera trapping: Implications for tegu trapping. Poster presentation at The Wildlife Society meeting, Baltimore Maryland. October 2024
Goforth, K.M. and R.R. Carthy. 2015. Effects of tidal pumping on gas exchange in sea turtle nesting beaches. Florida Undergraduate Research Conference, Daytona Beach, FL., 27-28 February, 2015. February 2015
Encinosa, M., L. Nicholson, C. Romagosa, and R.R. Carthy. 2022. Investigating the impact of roadway type and proximity on bat activity within a restoration context. Poster presentation at 29th Annual Meeting of TWS, Spokane, WA. November 2022
Ely, M, N. Bishop, B.J. Smith, C. Romagosa and R. Carthy. 2016. Monitoring changes in sea turtle nesting beaches in St. Johns County, Florida. Annual Meeting, Ecological Society of America, Fort Lauderdale, Florida, 7-12 August, 2016. August 2016
DeJesus, C., W.W. Boone, R.R. Carthy, C.M. Romagosa, and N.D. Bishop. 2017. The vulture restaurant is OPEN: Comparative feeding ethology of black and turkey vultures. Ecological Society of America, Portland, Oregon. August 2017
Cuevas- Hernandez, J R. Buron, C. Romagosa, and R. Carthy. 2021.<b> </b>Using eBird to assess the impact of urban land development on Least Tern (<i>Sternula antillarum) </i>breeding occupancy in Florida. Poster presentation, ESA, Long Beach, California. August 2021
Cronin, A., A. Wynn, B. Smith, N. Bishop and R. Carthy. 2015.Potential for ghost crab predation on sea turtle nests on driving and non-driving beaches in St. Johns County, Florida.100th Meeting, Ecological Society of America, Baltimore, Maryland, 9-14 August 2015. August 2015
Celestin, M. H. Innocent, N.D. Bishop, C.M. Romagosa, R.R.Carthy, and W.W. Boone. 2017. Investigating the influence of environmental covariates on marsh rabbit occupancy in southwest Florida. Ecological Society of America, Portland, Oregon. August 2017
Carthy, R.R., M.A. Burgess, P.G. Ifju, B.E. Wilkinson, and H.F. Percival. 2014. Development and application of small unmanned aircraft systems (sUAS) for natural resources: logistics, legalities, and lessons learned. 68th Annual Conference of the Southeastern Association of Fish and Wildlife Agencies. Destin, Florida. October 2014
Carthy, R.R., M. Burgess, P. Ifju, B. Wilkinson, S. Smith, and T. Whitley. 2016. UAs Technology for Sea Turtle Research and Conservation.Workshop Presentation at the South East Regional Sea Turtle Network Meeting. Mobile, AL, February 10-12,2016. February 2016
Carthy, R. R., T, Wibbels, and A. Rees. 2019. 3rd Workshop on the use of UAVs in sea turtle conservation and research. Workshop presentation, 39th International Sea Turtle Symposium, Charleston, SC.. February 2019
Carthy, R. R, P. J. Ifju, B. E. Wilkinson, A. Ortega, and M. Burgess. 2022. Overview of the University of Florida Uncrewed Aircraft Systems Research Program (UFUASRP): Two decades of drones for natural resource applications. Oral presentation at The Wildlife Society Drone Symposium, Spokane, WA. November 2022
Carthy, R. R, M. Mota, and N Dejardin. 2019. Effects of engineered beaches on sea turtle incubation and hatching. Workshop presentation, 39th International Sea Turtle Symposium, Charleson, SC. March 2019
Burgess, M. A., Zweig, C. L., Newman, S., Cook, M. I., Rodgers, H. L., Carthy, R. R., Wilkinson, B. E., Whitley, T. J., Ward, T. S., DiRodio, J. G., Frederick, P. C., Ifju, P. G., Smith, S. E., Percival, H. F. (2015, April). Applications of high-resolution aerial imagery and a small unmanned aircraft system in everglades science. Presented at the 2015 Greater Everglades Ecosystem Restoration (GEER) conference, Coral Springs, Fl. April 2015
Broxton, J, L Nicholson, L. Braun de Torrez, H. Ober, C. Romagosa, and R. Carthy. 2021.The implications of Acoustic Software Match Ratios for bat species identification and composition. Poster presentation, ESA, Long Beach, California. August 2021
Brisbane, J., C. Pedrozo, B.J. Smith, C. Romagosa, N. Bishop and R. Carthy. 2016. Environmental factors influencing the detection of frog species in three habitat types of Everglades National Park. Annual Meeting, Ecological Society of America, Fort Lauderdale, Florida, 7-12 August, 2016. August 2016
Bishop, N., T. Pop, J. Marlin, R. HUdson, S. Boylan, J. Polisar and R.R. Carthy. 2016. Utilizing growth rates to determine the onset of sexual maturity of Dermatemys mawii at the HCRC. 2nd Hicatee Conservation Forum and Workshop, Belize, February 25-26, 2016. February 2016
Bishop, N. D., and R. R. Carthy. 2019. A Novel Method for Assessing Color Differences in Dichotomous Groupings: An Example with Sexual Dichromatism in the Central American River Turtle, Dermatemys mawii. Poster presentation, 16th Annual Symposium on the Conservation and Biology of Tortoises and Freshwater Turtles,Fort Worth, TX. August 2018
Bishop, J. Polisar, P. Eliazar, K. Bjorndal and R.R. Carthy. 2016. Wild diet of Dermatemys mawii assessed by stomach content analysis. 2nd Hicatee Conservation Forum and Workshop, Belize, February 25-26, 2016. February 2016
Araya, P., P. Xiong., M. Harmon, A. Figueroa, L. Hussain, J. Broxton, R. R. Carthy, and C. Romagosa. 2024. Investigating the seed dispersal mechanisms of plant seeds in tegu (<i>Salvator merianae</i>), diet samples in Everglades National Park. Poster presentation at The Wildlife Society meeting, Baltimore Maryland. October 2024
Alvarez, O., C. K. Gulick, R. R. Carthy, and C. Romagosa. 2023. Alabama’s inshore islands have significantly shrunk over the last two decades. Poster presentation at The Wildlife Society meeting. Louisville, Kentucky. November 2023
Raymond R. Carthy, J. Roger Brothers, Vanessa Bézy,Kenneth J. Lohmann, Margaret M. Lamont. 2018. Ontogeny of orientation responses in developing sea turtle embryos. Presentation at the 38th International Symposium on Sea Turtle Biology and Conservation, Kobe, Japan. Feb. 18-24, 2018 February 2018
Daniel R. Evans, Raymond R. Carthy, and Cristina Ordoñez. 2018. Which Way Do You Go: Satellite Telemetry Reveals Regionally Different Foraging Behavior Of Leatherback Turtles. Presentation at the 38th International Symposium on Sea Turtle Biology and Conservation, Kobe, Japan. Feb. 18-24, 2018 February 2018
Technical Publications Publication Date
Rees, A. F. and R. R. Carthy. 2018. The Second Annual Workshop on the Use of UAVs in Sea Turtle Research and Conservation at the 38th International Sea Turtle Symposium: 19 Feb 2018. Marine Turtle Newsletter. April 2018
Nomani, S.Z., R.R. Carthy and M.K. Oli. 2006. Analysis of Abundance Estimation Methods for Gopher Tortoises (Gopherus polyphemus) July 2006
Langan, E.M., R.R. Carthy, S.Z. Nomani, S. Johnson, and M.K. Oli. 2006. Guidelines and standard techniques for estimating gopher tortoise abundance. August 2006
Gregory, C.J., B. Bird, and R.R. Carthy. 2003. The Grand Bay-Banks Lake Ecosystem Site Conservation Plan. Report to The Nature Conservancy, Florida Cooperative Fish and Wildlife Research Unit, U.S. Geological Survey, Department of Wildlife Ecology and Conservation, and the University of Florida. May 2003
Carthy, R., M.K. Oli, J.B. Wooding, J.E. Berish, and W.D. Meyer. 2005. Analysis of gopher tortoise population estimation techniques. Report to USACOE-CERL. October 2005
Role and Organization Individual         Start Date        End Date       
The Wildlife Society - Member Carthy January 1997 Present
Southeastern Association of Fish and Wildlife Agencies - Minorities in Natural Resources Committee Member Carthy August 1996 December 2006
Southeastern Association of Fish and Wildlife Agencies - Member Carthy May 2002 Present
Society for the Study of Amphibians and Reptiles - Member Carthy May 2001 Present
Other - President-Elect Carthy May 2022 Present
Other - President 2013 Carthy February 2012 May 2013
Other - Coordinator and Chair for paper sessions at 2005-2007 Anual Meetings Carthy 2005 2007
Other - Chair, Nominations Committee Carthy January 2015 Present
Other - Board of Directors Member Carthy May 2012 May 2017
American Society of Ichthyologists and Herpetologists - Member Carthy May 2001 Present