Cooperative Fish and Wildlife Research Units Program: Pennsylvania
Education, Research and Technical Assistance for Managing Our Natural Resources

Pennsylvania Education Activities


Pennsylvania staff electrofishing brown and rainbow trout in Spruce Creek in central Pennsylvania.

Admission Requirements and Procedures - Prospective students are encouraged to contact Unit personnel directly regarding graduate research opportunities. Unit staff are graduate faculty members in the School of Forest Resources and the Intercollege Graduate Degree Program in Ecology. Consult the Graduate Degree Programs section of the School of Forest Resources website or the IGDP Ecology website (see links to the right) for complete details about each program and application procedures.

Financial Support - Graduate students associated with the Unit are typically supported by external funds in the form of grants or research contracts. Fully supported graduate students receive a monthly stipend, tuition waiver, and health insurance.


Title Instructor(s) Start Date End Date
Wildlife Management Diefenbach 2012-08-22 2012-12-14
Structured Decision Making and Adaptive Management of Natural Resources Diefenbach, Wagner 2012-08-22 2012-12-14
Quantitative Methods in Ecology Wagner 2017-01-10 2017-05-05
Quantitative Methods in Ecology Wagner 2014-01-10 2014-05-05
Quantitative Methods in Ecology Wagner 2012-01-10 2012-05-05
Quantitative Methods in Ecology Wagner 2011-01-10 2011-05-05
Quantitative Methods in Ecology Wagner 2010-01-10 2010-05-05
Quantitative Methods in Ecology Wagner 2009-01-10 2009-05-05
Professional Communications Carline 2005-01-11 2005-05-06
Population estimation and modeling Diefenbach 2016-01-12 2016-05-07
Population estimation and modeling Diefenbach 2015-01-12 2015-05-06
Population Estimation and Modeling Diefenbach 2020-01-12 2020-05-08
Population Estimation and Modeling Diefenbach 2019-01-08 2019-05-05
Population Estimation and Modeling Diefenbach 2018-01-15 2018-05-05
Population Estimation and Modeling Diefenbach 2011-08-25 2011-12-05
Hierarchical models in ecology Wagner 2015-01-14 2015-05-04
Fishery Management Carline 2000-08-24 2000-12-12
Fish Population Dynamics Carline 2006-01-10 2006-05-10
Fish Population Dynamics Carline 2004-01-13 2004-05-07
Fish Population Dynamics Carline 2002-01-07 2002-05-03
Estimation of Fish and Wildlife Population Parameters Diefenbach 2007-08-27 2007-12-14
Estimation of Fish and Wildlife Population Parameters Diefenbach 2007-08-27 2007-12-10
Estimation of Fish and Wildlife Population Parameters Diefenbach 2005-09-02 2005-12-12
Estimation of Fish and Wildlife Population Parameters Diefenbach 2005-08-31 2005-12-09
Estimation of Fish and Wildlife Population Parameters Diefenbach 2002-09-01 2002-12-10
Estimation of Fish and Wildlife Population Parameters Diefenbach 2001-08-21 2001-12-07
Estimation of Fish and Wildlife Population Parameters Diefenbach 2000-08-21 2000-12-07
Ecosystem Monitoring Diefenbach 2020-08-22 2020-12-15
Communications in Natural Resources Carline 2003-01-07 2003-05-17
Classic Ecology Diefenbach 2008-08-28 2008-12-12
Applied Spatial Ecology Walter 2019-01-29 2019-04-26
Applied Spatial Ecology Walter 2017-01-09 2017-04-28
Applied Spatial Ecology Walter 2016-01-09 2016-04-28
Applied Spatial Ecology Walter 2015-01-05 2015-05-01
Applied Spatial Ecology Walter 2014-01-07 2014-04-26
Applied Spatial Ecology Walter 2013-01-07 2013-04-26
Analysis of Animal Location Data Diefenbach 2008-08-28 2008-12-12
Analysis of Animal Location Data Diefenbach 2006-09-06 2006-12-15
Analysis of Animal Location Data Diefenbach 2004-01-05 2004-05-07
Advances in Ecology Diefenbach 2016-09-13 2016-09-22
Advances in Ecology Diefenbach 2014-10-01 2014-10-15
Advances in Ecology Diefenbach 2013-09-24 2013-10-04
Advances in Ecology Diefenbach 2012-08-22 2012-12-14
Training Provided Instructor(s) Completion Date
This was an invited workshop on Bayesian hierarchical modeling. The course was attended by faculty, students, postdocs, and state biologists. The workshop was held at the Virginia Institute of Marine Science. Wagner 2016-05-26
This was an invited workshop on Bayesian hierarchical modeling. The course was attended by faculty, students, postdocs, and state biologists. The workshop was held at the University of Missouri. Wagner 2015-06-18
This was an invited workshop on Bayesian hierarchical modeling. The course was attended by faculty, students, postdocs, and state biologists. The workshop was held at The Ohio State University. Wagner 2015-04-24
Modeling survival using radio telemetry and nest monitoring data Diefenbach 2009-04-23
Introduction to occupancy modeling Diefenbach 2014-03-13
Introduction to R and hierarchical models was given to biologists at the PA Department of Environmental Protection. Wagner 2016-09-22
I was one of several instructors who gave talks on various aspects of environmental sampling for the PA Department of Environmental Protection pollution biologists. Carline 2001-11-13
Applied Spatial Ecology in R - Methods for managing and analyzing ecological datasets are constantly evolving. Advances in technology have resulted in large spatial datasets for aquatic and terrestrial populations that requires data compilation and analysis for making sound resource management and conservation decisions. From GIS dataset manipulation to traditional estimators of home range, this course will provide hands-on learning to handle a variety of datasets all within the software R (R Foundation for Statistical Computing, Vienna, Austria). Walter 2020-03-11
Applied Spatial Ecology in R - Methods for managing and analyzing ecological datasets are constantly evolving. Advances in technology have resulted in large spatial datasets for aquatic and terrestrial populations that requires data compilation and analysis for making sound resource management and conservation decisions. From GIS dataset manipulation to traditional estimators of home range, this course will provide hands-on learning to handle a variety of datasets all within the software R (R Foundation for Statistical Computing, Vienna, Austria). Walter 2018-09-18
Applied Spatial Ecology in R - Methods for managing and analyzing ecological datasets are constantly evolving. Advances in technology have resulted in large spatial datasets for aquatic and terrestrial populations that requires data compilation and analysis for making sound resource management and conservation decisions. From GIS dataset manipulation to traditional estimators of home range, this course will provide hands-on learning to handle a variety of datasets all within the software R (R Foundation for Statistical Computing, Vienna, Austria). Walter 2017-09-20
Applied Spatial Ecology in R - Methods for managing and analyzing ecological datasets are constantly evolving. Advances in technology have resulted in large spatial datasets for aquatic and terrestrial populations that requires data compilation and analysis for making sound resource management and conservation decisions. From GIS dataset manipulation to traditional estimators of home range, this course will provide hands-on learning to handle a variety of datasets all within the software R (R Foundation for Statistical Computing, Vienna, Austria). Walter 2015-09-30
Applied Spatial Ecology in R - Methods for managing and analyzing ecological datasets are constantly evolving. Advances in technology have resulted in large spatial datasets for aquatic and terrestrial populations that requires data compilation and analysis for making sound resource management and conservation decisions. From GIS dataset manipulation to traditional estimators of home range, this course will provide hands-on learning to handle a variety of datasets all within the software R (R Foundation for Statistical Computing, Vienna, Austria). Walter 2013-09-25
An overview of modeling left-censored contaminant data using hierarchical Bayesian models. Wagner 2015-09-29
An introduction to the use of distance sampling to estimate the abundance of grassland songbirds. Diefenbach 2004-04-05
An Introduction to R Wagner 2008-08-15
An Introduction to Multilevel Models and their Applications in Forest, Fish, and Wildlife Management: Multilevel data structures are a common feature in forestry and fish and wildlife research. These data structures most often results in a lack of independence among observations (i.e., pseudoreplication). Ignoring multilevel data structures during analysis has implications for inference and decision making, leading to overly optimistic uncertainty estimates (e.g., underestimated standard errors of estimated parameters). Multilevel models not only account for nested data structures, but also provide a powerful approach for modeling the effects of predictors at multiple spatial scales and for partitioning variability into multiple spatial and temporal components. This course will provide an overview of multilevel models, including the concept and interpretation of random effects and variance components (day 1), and hands-on experience fitting models using the freely available programming environment R (day 2). Wagner, Hill 2012-09-28
An Introduction to Bayesian Estimation and Inference Using JAGS This workshop provided an introduction to Bayesian estimation and inference using the program environment R and JAGS. Wagner 2013-06-18
Title and Location Speaker Date
White-tailed deer dispersal: implications for population and disease management
Presented to: University of Georgia
Diefenbach 2016-08-18
We're 90% certain you will want to be a Bayesian after this seminar
Presented to: Pennsylvania State University
Diefenbach, Wagner, Walter 2012-02-17
Truths and Myths about Chronic Wasting Disease
Presented to: USDA/APHIS/WS
Walter 2017-10-17
Truths and Myths about Chronic Wasting Disease
Presented to: Pennsylvania Forest Webinar Series
Walter 2018-06-12
Truths and Myths about Chronic Wasting Disease
Presented to: Indiana Chapter of the Wildlife Society
Walter 2017-10-04
The Impossible Dream: Managing White-tailed Deer to Meet Forest Management Objectives
Presented to: University of Maine
Diefenbach 2019-04-17
Temporal, spatial, and demographic patterns of white-tailed deer mortality: implications for their ecology and management.
Presented to: University of Delaware
Diefenbach 2010-02-24
Spatial, Temporal, and Demographic variation in male turkey harvest rates in New York, Ohio, and Pennsylvania.
Presented to: Pennsylvania State University
Diefenbach 2010-02-12
Putting together pieces of the puzzle: contaminants, pathogens, and smallmouth bass population dynamics in the Susquehanna River Basin
Presented to: The Pennsylvania State University
Wagner 2019-10-15
Predator-Prey Dynamics of Hunters and White-tailed Deer: Implications for Deer Population Management
Presented to: Pennsylvania State University
Diefenbach 2004-10-05
Online Database Infrastructures in Animal Ecology: A Novel way of Data Management and Sharing of Animal derived Sensor Data
Presented to: National Wildlife Research Center
Walter 2013-06-12
Linking lakes and landscapes: applying principles of landscape limnology to the management of inland lakes
Presented to: Pennsylvania State University
Wagner 2009-12-14
Hierarchical Models
Presented to: Michigan State University
Wagner 2011-06-27
Genetics of cervids and susceptibility to chronic wasting disease
Presented to: Newcastle University
Walter 2016-07-26
Genetics of Cervids as it relates to susceptibility to Chronic Wasting Disease
Presented to: Eastern Tennessee State University
Walter 2019-10-23
Fish Habitat and Climate Change: Implications for the Desert Southwest, Midwestern Smallmouth Bass, and Eastern Brook Trout
Presented to: NCCWSC Climate Change Science and Management Webinar Series
Wagner, Deweber 2013-04-11
Effects of highway construction on sediment loading in Spring Creek
Presented to: Penn State University
Carline 2001-11-19
Chronic wasting disease in white-tailed deer in the east: Coping With Disease (CWD)
Presented to: State University of New York - Environmental Science and Forestry
Walter 2015-07-09
CWD genetics in the wild and model results outline potential for spread within Pennsylvania
Presented to: Pennsylvania Department of Agriculture
Walter 2017-06-20
Buruli ulcer disease: insights into a 110 year mystery using landscape-based models
Presented to: Pennsylvania State University
Wagner 2008-05-09
Brook Trout ecology and management: land use, climate and transboundary population monitoring
Presented to: University of Georgia
Wagner 2013-11-08
Assessing response of trout to habitat manipulation
Presented to: Lycoming College
Carline 2001-12-13
A conceptual framework for understanding multiscaled interactions between terrestrial and aquatic ecosystems
Presented to: University of Maryland Center for Environmental Science
Wagner 2013-04-04
Title Presenter Date
Wildlife Disease Walter 2013-10-24
Wildlife Disease Walter 2012-10-30
Understanding dependency in Ecological data Walter 2015-11-12
Understanding dependency in Ecological data Walter 2014-09-18
Trout Population Dynamics Carline 2003-03-04
Stream habitat management Carline 2004-03-29
Stream ecology Carline 2005-11-10
Stream ecology Carline 2005-10-07
Spatial epidemiology of unique diseases: prions, parasites, and bacteria Walter 2014-11-05
Spatial Epidemiology of Unique Diseases: Prions, parasites,and bacteria Walter 2012-04-16
Sampling fish and wildlife populations Diefenbach 2001-10-04
Sampling aquatic environments Carline 2002-11-02
Salmonid spawning habitat Carline 2000-11-14
Risk Assessment and Disease Management Walter 2018-09-13
Risk Assessment and Disease Management Walter 2017-09-09
Risk Assessment and Disease Management Walter 2016-09-13
Risk Assessment and Disease Management Walter 2015-12-01
Risk Assessment and Disease Management Walter 2014-12-09
Riparian restoration Carline 2005-04-17
Riparian management in the Spruce Creek watershed Carline 2004-03-23
Reintroductions of wildlife Diefenbach 2014-11-07
Population restoration Diefenbach 2018-03-15
Management of Game Birds Diefenbach 2010-04-22
Management of Game Birds Diefenbach 2009-04-14
Management of Game Birds Diefenbach 2008-04-11
Mammalian dispersal Diefenbach 2018-04-22
Mammalian dispersal Diefenbach 2015-03-06
Lake ecology Carline 2006-04-19
Impacts of climate change on eastern U.S. fisheries Wagner 2015-11-17
History of deer management in Pennsylvania. Diefenbach 2005-03-16
Fundamentals of Riparian Management Carline 2005-04-27
Field ecology of Big Spring Creek Carline 2002-01-23
Elements of Electrofishing Carline 2003-09-11
Effects of acidic deposition on stream biota Carline 2002-12-04
Dispersal in White-tailed Deer Diefenbach 2017-02-22
Dietary uptake of PCBs by rainbow trout Carline 2003-03-26
Deer Management in Pennsylvania Diefenbach 2006-03-13
Advances in Ecology, Statistical Modelling Wagner 2012-10-04
Advances in Ecology, Statistical Modelling Wagner 2011-10-04
Advances in Ecology, Statistical Modelling Wagner 2010-10-13
Advances in Ecology, Statistical Modelling Wagner 2009-10-13
Activity patterns of female black bears Diefenbach 2005-11-29
Student (Degree) Advisor Start Year End Year
Wilson, Andy (PhD) Diefenbach 2005 2009
Williamson, Neihl (MS) Wagner 2008 2010
White, Shannon (PhD) Diefenbach 2017 2019
Wentworth, Kevin (PhD) Diefenbach 2002 2005
Weiderholt, Ruscena (PhD) Diefenbach 2007 2011
Weber, Andrew (MS) Diefenbach 2010 2014
Weather, Thomas (PhD) Walter 2016 2019
Voorhees, Christine (MS) Diefenbach 2000 2002
Tolley, Keeley (MS) Diefenbach 2000 2002
Stokke, Grant (MS) Diefenbach 2005 2014
Stainbrook, David (MS) Wagner 2008 2011
Smith, Lori (MS) Walter 2012 2014
Slawson, Deborah (PhD) Carline 2000 2004
Sklebo, Jared (MS) Diefenbach 2014 2017
Sidelick, Jennifer (MS) Carline 2003 2005
Schall, Megan (PhD) Walter 2013 2017
Rummel, Shawn (PhD) Diefenbach 2007 2011
Rodemaker, Vitor (PhD) Diefenbach 2007 2007
Rocco, Gian (PhD) Carline 2000 Present
Regmi, Arun (PhD) Diefenbach 2020 Present
Reed, Brendan (MS) Wagner 2012 2014
Price, Keith (PhD) Wagner 2008 2011
Petprakob, Krittika (PhD) Wagner 2009 Present
Pedersen, Christian (PhD) Diefenbach 2004 2008
O'Driscoll, Michael (PhD) Carline 2001 2004
Norton, Andrew (MS) Wagner 2008 2011
Munoz, David (MS) Diefenbach 2014 2015
Mollenhauer, Robert (MS) Diefenbach 2009 2012
Miller, Tricia (MS) Diefenbach 2005 2007
Means, Jennifer (MS) Carline 2003 Present
McNeil, Leonard (MS) Carline 2003 2005
May, Melissa (MS) Wagner 2009 2011
Mason, Doris (MS) Wagner 2008 2009
Long, Eric (PhD) Carline 2001 2005
Lees, Bryan (MS) Carline 2003 2005
Kerby, Jeff (PhD) Wagner 2010 2014
Julian, James (PhD) Diefenbach 2005 2009
Jones, David (MS) Carline 2005 Present
Ikis, Didem (PhD) Wagner 2010 2017
Hill, Jason (PhD) Wagner 2009 2012
Grassie, Cairsty (PhD) Wagner 2009 2013
Govenar, Breea (PhD) Diefenbach 2003 2005
Goodrich, Laurie (PhD) Diefenbach 2003 2010
Gingery, Tess (MS) Walter 2015 2018
Garneau, Danielle (PhD) Diefenbach 2002 2005
Ganoe, Laken (MS) Diefenbach 2018 2019
Fronk, Nate (MS) Walter 2012 2013
Forman, Nicholas (MS) Diefenbach 2012 2015
Fisher, Douglas (MS) Carline 2002 Present
Fergus, Emi (PhD) Wagner 2012 2016
Espindola, Walter (PhD) Walter 2020 Present
Deweber, Jefferson (PhD) Walter 2012 2014
Davis, Courtney (PhD) Wagner 2015 Present
Davis, Courtney (MS) Wagner 2013 2015
Crawford, Charles (MS) Diefenbach 2012 2015
Compton, Justin (PhD) Diefenbach 2005 2007
Chen, Xin (PhD) Diefenbach 2017 Present
Byrne, Loren (PhD) Diefenbach 2002 2006
Buda, Anthony (PhD) Carline 2002 Present
Brandell, Ellen (PhD) Walter 2016 Present
Becker, Douglas (MS) Diefenbach 2003 2005
Barton, Ethan (MS) Diefenbach 2011 2014
Barry, Patrick (MS) Diefenbach 2003 2004
Anders, Angela (PhD) Diefenbach 2003 2007
Amburgey , Staci (PhD) Wagner 2015 Present
Alexander, Richard (PhD) Wagner 2010 2014