Cooperative Fish and Wildlife Research Units Program: Pennsylvania
Education, Research and Technical Assistance for Managing Our Natural Resources

Pennsylvania Education Activities


Pennsylvania staff electrofishing brown and rainbow trout in Spruce Creek in central Pennsylvania.

Admission Requirements and Procedures - Prospective students are encouraged to contact Unit personnel directly regarding graduate research opportunities. Unit staff are graduate faculty members in the School of Forest Resources and the Intercollege Graduate Degree Program in Ecology. Consult the Graduate Degree Programs section of the School of Forest Resources website or the IGDP Ecology website (see links to the right) for complete details about each program and application procedures.

Financial Support - Graduate students associated with the Unit are typically supported by external funds in the form of grants or research contracts. Fully supported graduate students receive a monthly stipend, tuition waiver, and health insurance.

Title Instructor(s) Start Date End Date
Wildlife Management Diefenbach August 2012 December 2012
Structured Decision Making and Adaptive Management of Natural Resources Diefenbach, Wagner August 2012 December 2012
Quantitative Methods in Ecology Wagner January 2017 May 2017
Quantitative Methods in Ecology Wagner January 2014 May 2014
Quantitative Methods in Ecology Wagner January 2012 May 2012
Quantitative Methods in Ecology Wagner January 2011 May 2011
Quantitative Methods in Ecology Wagner January 2010 May 2010
Quantitative Methods in Ecology Wagner January 2009 May 2009
Professional Communications Carline January 2005 May 2005
Population estimation and modeling Diefenbach January 2016 May 2016
Population estimation and modeling Diefenbach January 2015 May 2015
Population Estimation and Modeling Diefenbach January 2020 May 2020
Population Estimation and Modeling Diefenbach January 2019 May 2019
Population Estimation and Modeling Diefenbach January 2018 May 2018
Population Estimation and Modeling Diefenbach August 2011 December 2011
Joint Species Distribution Models Wagner January 2021 May 2021
Hierarchical models in ecology Wagner January 2015 May 2015
Fishery Management Carline August 2000 December 2000
Fish Population Dynamics Carline January 2006 May 2006
Fish Population Dynamics Carline January 2004 May 2004
Fish Population Dynamics Carline January 2002 May 2002
Estimation of Fish and Wildlife Population Parameters Diefenbach September 2005 December 2005
Estimation of Fish and Wildlife Population Parameters Diefenbach September 2002 December 2002
Estimation of Fish and Wildlife Population Parameters Diefenbach August 2007 December 2007
Estimation of Fish and Wildlife Population Parameters Diefenbach August 2005 December 2005
Estimation of Fish and Wildlife Population Parameters Diefenbach August 2001 December 2001
Estimation of Fish and Wildlife Population Parameters Diefenbach August 2000 December 2000
Ecosystem Monitoring Diefenbach August 2020 December 2020
Communications in Natural Resources Carline January 2003 May 2003
Classic Ecology Diefenbach August 2008 December 2008
Applied Spatial Ecology Walter January 2021 April 2021
Applied Spatial Ecology Walter January 2019 April 2019
Applied Spatial Ecology Walter January 2017 April 2017
Applied Spatial Ecology Walter January 2016 April 2016
Applied Spatial Ecology Walter January 2015 May 2015
Applied Spatial Ecology Walter January 2014 April 2014
Applied Spatial Ecology Walter January 2013 April 2013
Analysis of Animal Location Data Diefenbach September 2006 December 2006
Analysis of Animal Location Data Diefenbach January 2004 May 2004
Analysis of Animal Location Data Diefenbach August 2008 December 2008
Advances in Ecology Diefenbach September 2016 September 2016
Advances in Ecology Diefenbach September 2013 October 2013
Advances in Ecology Diefenbach October 2014 October 2014
Advances in Ecology Diefenbach August 2012 December 2012
Training Provided Instructor(s) Completion Date
Virtual workshop in Applied Spatial Ecology in R - Methods for managing and analyzing ecological datasets are constantly evolving. Advances in technology have resulted in large spatial datasets for aquatic and terrestrial populations that requires data compilation and analysis for making sound resource management and conservation decisions. From GIS dataset manipulation to traditional estimators of home range, this course will provide hands-on learning to handle a variety of datasets all within the software R (R Foundation for Statistical Computing, Vienna, Austria). Walter September 2021
This was an invited workshop on Bayesian hierarchical modeling. The course was attended by faculty, students, postdocs, and state biologists. The workshop was held at the Virginia Institute of Marine Science. Wagner May 2016
This was an invited workshop on Bayesian hierarchical modeling. The course was attended by faculty, students, postdocs, and state biologists. The workshop was held at the University of Missouri. Wagner June 2015
This was an invited workshop on Bayesian hierarchical modeling. The course was attended by faculty, students, postdocs, and state biologists. The workshop was held at The Ohio State University. Wagner April 2015
Modeling survival using radio telemetry and nest monitoring data Diefenbach April 2009
Introduction to occupancy modeling Diefenbach March 2014
Introduction to R and hierarchical models was given to biologists at the PA Department of Environmental Protection. Wagner September 2016
I was one of several instructors who gave talks on various aspects of environmental sampling for the PA Department of Environmental Protection pollution biologists. Carline November 2001
Applied Spatial Ecology in R - Methods for managing and analyzing ecological datasets are constantly evolving. Advances in technology have resulted in large spatial datasets for aquatic and terrestrial populations that requires data compilation and analysis for making sound resource management and conservation decisions. From GIS dataset manipulation to traditional estimators of home range, this course will provide hands-on learning to handle a variety of datasets all within the software R (R Foundation for Statistical Computing, Vienna, Austria). Walter September 2018
Applied Spatial Ecology in R - Methods for managing and analyzing ecological datasets are constantly evolving. Advances in technology have resulted in large spatial datasets for aquatic and terrestrial populations that requires data compilation and analysis for making sound resource management and conservation decisions. From GIS dataset manipulation to traditional estimators of home range, this course will provide hands-on learning to handle a variety of datasets all within the software R (R Foundation for Statistical Computing, Vienna, Austria). Walter September 2017
Applied Spatial Ecology in R - Methods for managing and analyzing ecological datasets are constantly evolving. Advances in technology have resulted in large spatial datasets for aquatic and terrestrial populations that requires data compilation and analysis for making sound resource management and conservation decisions. From GIS dataset manipulation to traditional estimators of home range, this course will provide hands-on learning to handle a variety of datasets all within the software R (R Foundation for Statistical Computing, Vienna, Austria). Walter September 2015
Applied Spatial Ecology in R - Methods for managing and analyzing ecological datasets are constantly evolving. Advances in technology have resulted in large spatial datasets for aquatic and terrestrial populations that requires data compilation and analysis for making sound resource management and conservation decisions. From GIS dataset manipulation to traditional estimators of home range, this course will provide hands-on learning to handle a variety of datasets all within the software R (R Foundation for Statistical Computing, Vienna, Austria). Walter March 2020
Applied Spatial Ecology in R - Methods for managing and analyzing ecological datasets are constantly evolving. Advances in technology have resulted in large spatial datasets for aquatic and terrestrial populations that requires data compilation and analysis for making sound resource management and conservation decisions. From GIS dataset manipulation to traditional estimators of home range, this course will provide hands-on learning to handle a variety of datasets all within the software R (R Foundation for Statistical Computing, Vienna, Austria). Walter September 2013
An overview of modeling left-censored contaminant data using hierarchical Bayesian models. Wagner September 2015
An introduction to the use of distance sampling to estimate the abundance of grassland songbirds. Diefenbach April 2004
An Introduction to R Wagner August 2008
An Introduction to Multilevel Models and their Applications in Forest, Fish, and Wildlife Management: Multilevel data structures are a common feature in forestry and fish and wildlife research. These data structures most often results in a lack of independence among observations (i.e., pseudoreplication). Ignoring multilevel data structures during analysis has implications for inference and decision making, leading to overly optimistic uncertainty estimates (e.g., underestimated standard errors of estimated parameters). Multilevel models not only account for nested data structures, but also provide a powerful approach for modeling the effects of predictors at multiple spatial scales and for partitioning variability into multiple spatial and temporal components. This course will provide an overview of multilevel models, including the concept and interpretation of random effects and variance components (day 1), and hands-on experience fitting models using the freely available programming environment R (day 2). Wagner, Hill September 2012
An Introduction to Bayesian Estimation and Inference Using JAGS This workshop provided an introduction to Bayesian estimation and inference using the program environment R and JAGS. Wagner June 2013
Title and Location Speaker Date
White-tailed deer dispersal: implications for population and disease management
Presented to: University of Georgia
Diefenbach August 2016
We're 90% certain you will want to be a Bayesian after this seminar
Presented to: Pennsylvania State University
Diefenbach, Wagner, Walter February 2012
Truths and Myths about Chronic Wasting Disease
Presented to: USDA/APHIS/WS
Walter October 2017
Truths and Myths about Chronic Wasting Disease
Presented to: Pennsylvania Forest Webinar Series
Walter June 2018
Truths and Myths about Chronic Wasting Disease
Presented to: Indiana Chapter of the Wildlife Society
Walter October 2017
The Impossible Dream: Managing White-tailed Deer to Meet Forest Management Objectives
Presented to: University of Maine
Diefenbach April 2019
Temporal, spatial, and demographic patterns of white-tailed deer mortality: implications for their ecology and management.
Presented to: University of Delaware
Diefenbach February 2010
Spatial, Temporal, and Demographic variation in male turkey harvest rates in New York, Ohio, and Pennsylvania.
Presented to: Pennsylvania State University
Diefenbach February 2010
Putting together pieces of the puzzle: contaminants, pathogens, and smallmouth bass population dynamics in the Susquehanna River Basin
Presented to: The Pennsylvania State University
Wagner October 2019
Predator-Prey Dynamics of Hunters and White-tailed Deer: Implications for Deer Population Management
Presented to: Pennsylvania State University
Diefenbach October 2004
Online Database Infrastructures in Animal Ecology: A Novel way of Data Management and Sharing of Animal derived Sensor Data
Presented to: National Wildlife Research Center
Walter June 2013
Linking lakes and landscapes: applying principles of landscape limnology to the management of inland lakes
Presented to: Pennsylvania State University
Wagner December 2009
Landscape Genetics to Inform Management for Chronic Wasting Disease
Presented to: USGS, USDA, AFWA
Walter July 2021
Hierarchical Models
Presented to: Michigan State University
Wagner June 2011
Genetics of cervids and susceptibility to chronic wasting disease
Presented to: Newcastle University
Walter July 2016
Genetics of Cervids as it relates to susceptibility to Chronic Wasting Disease
Presented to: Eastern Tennessee State University
Walter October 2019
Fish Habitat and Climate Change: Implications for the Desert Southwest, Midwestern Smallmouth Bass, and Eastern Brook Trout
Presented to: NCCWSC Climate Change Science and Management Webinar Series
Wagner, Deweber April 2013
Effects of highway construction on sediment loading in Spring Creek
Presented to: Penn State University
Carline November 2001
Chronic wasting disease in white-tailed deer in the east: Coping With Disease (CWD)
Presented to: State University of New York - Environmental Science and Forestry
Walter July 2015
CWD genetics in the wild and model results outline potential for spread within Pennsylvania
Presented to: Pennsylvania Department of Agriculture
Walter June 2017
Buruli ulcer disease: insights into a 110 year mystery using landscape-based models
Presented to: Pennsylvania State University
Wagner May 2008
Brook Trout ecology and management: land use, climate and transboundary population monitoring
Presented to: University of Georgia
Wagner November 2013
Assessing response of trout to habitat manipulation
Presented to: Lycoming College
Carline December 2001
A conceptual framework for understanding multiscaled interactions between terrestrial and aquatic ecosystems
Presented to: University of Maryland Center for Environmental Science
Wagner April 2013
Title Presenter Date
Wildlife Disease Walter October 2013
Wildlife Disease Walter October 2012
Understanding dependency in Ecological data Walter September 2014
Understanding dependency in Ecological data Walter November 2015
Trout Population Dynamics Carline March 2003
Stream habitat management Carline March 2004
Stream ecology Carline October 2005
Stream ecology Carline November 2005
Spatial epidemiology of unique diseases: prions, parasites, and bacteria Walter November 2014
Spatial Epidemiology of Unique Diseases: Prions, parasites,and bacteria Walter April 2012
Sampling fish and wildlife populations Diefenbach October 2001
Sampling aquatic environments Carline November 2002
Salmonid spawning habitat Carline November 2000
Risk Assessment and Disease Management Walter September 2018
Risk Assessment and Disease Management Walter September 2017
Risk Assessment and Disease Management Walter September 2016
Risk Assessment and Disease Management Walter December 2015
Risk Assessment and Disease Management Walter December 2014
Riparian restoration Carline April 2005
Riparian management in the Spruce Creek watershed Carline March 2004
Reintroductions of wildlife Diefenbach November 2014
Population restoration Diefenbach March 2018
Management of Game Birds Diefenbach April 2010
Management of Game Birds Diefenbach April 2009
Management of Game Birds Diefenbach April 2008
Mammalian dispersal Diefenbach March 2015
Mammalian dispersal Diefenbach April 2018
Lake ecology Carline April 2006
Impacts of climate change on eastern U.S. fisheries Wagner November 2015
History of deer management in Pennsylvania. Diefenbach March 2005
Fundamentals of Riparian Management Carline April 2005
Field ecology of Big Spring Creek Carline January 2002
Elements of Electrofishing Carline September 2003
Effects of acidic deposition on stream biota Carline December 2002
Dispersal in White-tailed Deer Diefenbach February 2017
Dietary uptake of PCBs by rainbow trout Carline March 2003
Deer Management in Pennsylvania Diefenbach March 2006
Advances in Ecology, Statistical Modelling Wagner October 2012
Advances in Ecology, Statistical Modelling Wagner October 2011
Advances in Ecology, Statistical Modelling Wagner October 2010
Advances in Ecology, Statistical Modelling Wagner October 2009
Activity patterns of female black bears Diefenbach November 2005
Student (Degree) Advisor Start Year End Year
Wilson, Andy (PhD) Diefenbach 2005 2009
Williamson, Neihl (MS) Wagner 2008 2010
Wikle, Nathan (PhD) Wagner 2017 2021
White, Shannon (PhD) Diefenbach 2017 2019
Wentworth, Kevin (PhD) Diefenbach 2002 2005
Weiderholt, Ruscena (PhD) Diefenbach 2007 2011
Weber, Andrew (MS) Diefenbach 2010 2014
Weather, Thomas (PhD) Walter 2016 2019
Voorhees, Christine (MS) Diefenbach 2000 2002
Tolley, Keeley (MS) Diefenbach 2000 2002
Stokke, Grant (MS) Diefenbach 2005 2014
Stenger, Logan (MS) Wagner 2021 Present
Stark, Sydney (MS) Wagner 2021 Present
Stainbrook, David (MS) Wagner 2008 2011
Smith, Lori (MS) Walter 2012 2014
Slawson, Deborah (PhD) Carline 2000 2004
Sklebo, Jared (MS) Diefenbach 2014 2017
Sidelick, Jennifer (MS) Carline 2003 2005
Schall, Megan (PhD) Walter 2013 2017
Rummel, Shawn (PhD) Diefenbach 2007 2011
Rodemaker, Vitor (PhD) Diefenbach 2007 2007
Rocco, Gian (PhD) Carline 2000 Present
Regmi, Arun (PhD) Diefenbach 2020 Present
Reed, Brendan (MS) Wagner 2012 2014
Price, Keith (PhD) Wagner 2008 2011
Petprakob, Krittika (PhD) Wagner 2009 2015
Pedersen, Christian (PhD) Diefenbach 2004 2008
O'Driscoll, Michael (PhD) Carline 2001 2004
Norton, Andrew (MS) Wagner 2008 2011
Munoz, David (MS) Diefenbach 2014 2015
Mollenhauer, Robert (MS) Diefenbach 2009 2012
Miller, Tricia (MS) Diefenbach 2005 2007
Means, Jennifer (MS) Carline 2003 Present
McNeil, Leonard (MS) Carline 2003 2005
May, Melissa (MS) Wagner 2009 2011
Mason, Doris (MS) Wagner 2008 2009
Long, Eric (PhD) Carline 2001 2005
Lees, Bryan (MS) Carline 2003 2005
Kerby, Jeff (PhD) Wagner 2010 2014
Julian, James (PhD) Diefenbach 2005 2009
Jones, David (MS) Carline 2005 Present
Ikis, Didem (PhD) Wagner 2010 2017
Hill, Jason (PhD) Wagner 2009 2012
Guerin, Dan (PhD) Diefenbach 2021 Present
Grassie, Cairsty (PhD) Wagner 2009 2013
Govenar, Breea (PhD) Diefenbach 2003 2005
Goodrich, Laurie (PhD) Diefenbach 2003 2010
Gingery, Tess (MS) Walter 2015 2018
Garneau, Danielle (PhD) Diefenbach 2002 2005
Ganoe, Laken (MS) Diefenbach 2018 2019
Fronk, Nate (MS) Walter 2012 2013
Forman, Nicholas (MS) Diefenbach 2012 2015
Fisher, Douglas (MS) Carline 2002 Present
Fergus, Emi (PhD) Wagner 2012 2016
Espindola, Walter (MS) Walter 2020 2021
Deweber, Jefferson (PhD) Walter 2012 2014
Davis, Courtney (PhD) Wagner 2015 2019
Davis, Courtney (MS) Wagner 2013 2015
Crawford, Charles (MS) Diefenbach 2012 2015
Corondi, Avery (MS) Diefenbach 2021 Present
Compton, Justin (PhD) Diefenbach 2005 2007
Chen, Xin (PhD) Diefenbach 2017 2020
Byrne, Loren (PhD) Diefenbach 2002 2006
Buda, Anthony (PhD) Carline 2002 Present
Brandell, Ellen (PhD) Walter 2016 2021
Becker, Douglas (MS) Diefenbach 2003 2005
Barton, Ethan (MS) Diefenbach 2011 2014
Barry, Patrick (MS) Diefenbach 2003 2004
Anders, Angela (PhD) Diefenbach 2003 2007
Amburgey , Staci (PhD) Wagner 2015 2019
Alexander, Richard (PhD) Wagner 2010 2014