Cooperative Fish and Wildlife Research Units Program: Wyoming
Education, Research and Technical Assistance for Managing Our Natural Resources

Wyoming Staff Member

Arthur Middleton

Arthur Middleton

Phone: (307) 766 - 6415
Personal website


Arthur Middleton is a Ph.D. candidate in the Program in Ecology at the University of Wyoming, based in the Cooperative Fish and Wildlife Research Unit within the Department of Zoology and Physiology.

Research Interest

Prior to starting his research, Arthur received a master’s degree in environmental management from Yale University and a bachelor’s degree in English and government from Bowdoin College. Before coming to Wyoming, most of Arthur’s experience was with birds of prey - as a falconer, he trained numerous raptor species for public education and for hunting, and as a field assistant, he contributed to studies of the swallow-tailed kite in South Carolina and the harpy eagle in Panama. He was also responsible for the field component of a bald eagle restoration project in New York. Since coming to Wyoming in 2007, Arthur has coordinated the Absaroka Elk Ecology Project in collaboration with the Wyoming Game and Fish Department and the U.S. Fish and Wildlife Service, studying elk migration and elk-wolf interactions in the Absaroka Mountains. The primary focus of his dissertation research is evaluating the relative influence of top-down (predation risk) versus bottom-up (habitat quality) forces on the nutritional condition and reproduction of elk. Arthur is also initiating new work with Emiliano Donadio and others on the ecology of a high Andean food web in Argentina, where pumas and condors interact strongly with native camelids. Arthur has received a Thomas J. Watson Fellowship, a Doris Duke Conservation Fellowship, and a Resident Fellowship at the Buffalo Bill Historical Center in Cody, Wyoming. He anticipates finishing his degree in spring 2012.

Research Publications Publication Date
Sawyer, H., M.J. Kauffman, A.D. Middleton, T.A. Morrison, R.M. Nielson, and T.B. Wyckoff. 2013. A framework for understanding semi-permeable barrier effects on migratory ungulates. Journal of Applied Ecology, 50 (1). DOI: 10.1111/1365-2664.12013 | Abstract | Download | Publisher Website 2013-02-28
Pauli, J.N., J.P. Whiteman, M. Riley, and A.D. Middleton. 2010. Defining noninvasive for sampling of vertebrates. Conservation Biology 24:349-352. | Download 2010-02-28
Middleton, A.D., M.J. Kauffman, D.E. McWhirter, J.G. Cook, R.C. Cook, A.A. Nelson, M.D. Jimenez, and R.W. Klaver. 2013. Animal migration amid shifting patterns of phenology and predation: lessons from a Yellowstone elk herd. Ecology 94:1245-1256. 2013-12-31
Martínez del Rio, C. and A.D. Middleton. 2010. Laws for ecology? (book review). Ecology 91:1244-1245. | Download 2010-04-30
Cross, P.C., E.K. Cole, A.P. Dobson, W.H. Edwards, K.L. Hamlin, G. Luikart, A.D. Middleton, B.M. Scurlock, and P.J. White. 2010. Probable causes of increasing elk brucellosis in the Greater Yellowstone Ecosystem. Ecological Applications 20: 278-288. | Download 2010-01-31
Christianson, D., R.W. Klaver, A. Middleton, and M. Kauffman. 2013. Confounded winter and spring phenoclimatology on large herbivore ranges. Landscape Ecology 28:427–437. 2013-01-31
Technical Publications Publication Date
Absaroka Elk Ecology Project - 2010 Annual Report | Download 2010-12-31
Absaroka Elk Ecology Project - 2008 Annual Report | Download 2008-06-30
Absaroka Elk Ecology - 2009 Annual Report | Download 2009-12-31