Print Report
A3732 Amelanchier utahensis - Cercocarpus montanus - Cercocarpus intricatus Shrubland Alliance
Type Concept Sentence: This alliance is characterized by shrublands dominated by Amelanchier utahensis, Cercocarpus montanus or Cercocarpus intricatus in the southern Rocky Mountains, Wyoming Basins, Colorado Plateau and extending west to the Great Basin.
Common (Translated Scientific) Name: Utah Serviceberry - Alderleaf Mountain-mahogany - Littleleaf Mountain-mahogany Shrubland Alliance
Colloquial Name: Utah Serviceberry - Mountain-mahogany Shrubland
Hierarchy Level: Alliance
Type Concept: This alliance is known from the southern Rocky Mountains, Wyoming Basins, Colorado Plateau and extending west to the Great Basin. Stands are characterized by mixed tall-shrub canopies dominated by Amelanchier utahensis, Cercocarpus montanus, or Cercocarpus intricatus. Species that may codominate in some stands include Artemisia tridentata, Fendlera rupicola, Glossopetalon spinescens, and Rhus trilobata. Total canopy cover ranges from sparse to moderately dense. Occasional trees, such as Juniperus spp., Quercus spp., Pinus edulis, Pinus ponderosa, or Pseudotsuga menziesii, may be scattered in some stands. The herbaceous layer is variable and may have sparse to moderate cover. The most typical dominants are graminoids and may include Carex geyeri, Elymus lanceolatus ssp. lanceolatus, Hesperostipa comata, and Pseudoroegneria spicata. Elevations range from 1800-2700 m. Sites are variable but are generally xeric and rocky with moderate to very steep slopes or on ridges. Stands are found on various aspects, but typically the higher elevation and northern sites are on warmer, southern exposures, and lower elevation and southern stands are restricted to more mesic north slopes.
Diagnostic Characteristics: Mixed montane shrublands dominated Amelanchier utahensis, Cercocarpus montanus, or occasionally Cercocarpus intricatus. Shrub cover may be sparse to moderate. These shrublands may be either codominated by other shrub species or have sparse to dense understories of graminoids. Tree may be present, but are scattered and have low cover.
Rationale for Nominal Species or Physiognomic Features: No Data Available
Classification Comments: Inclusion of ~Cercocarpus intricatus Montane Shrubland (CEGL002587)$$ is tentative as it occupies the same geographic range and montane environments of other associations in this alliance and has similar floristics. ~Cercocarpus intricatus - Glossopetalon spinescens Shrubland (CEGL005426)$$ is included in that it is montane with some floristic overlap, but is a geographic outlier, therefore placement here is low in confidence. Other Cercocarpus intricatus associations are more associated with slickrock environments and range more western and southern.
Similar NVC Types: No Data Available
note: No Data Available
Physiognomy and Structure: This alliance is characterized by a moderate to dense canopy of cold-deciduous shrubs up to 4 m in height. There is often another stratum (sometimes two) of low cold-deciduous shrubs from 0.5-2 m tall. The ground layer is usually a species-rich assemblage of cespitose graminoids and erect forbs.
Floristics: These shrublands are typically sparse to dense and tall with multiple vegetation layers. The upper shrub layer is typically dominated by Amelanchier utahensis, Cercocarpus montanus, or Cercocarpus intricatus. Species that may codominate in some stands include Artemisia tridentata, Fendlera rupicola, Glossopetalon spinescens, and Rhus trilobata. Other shrub associates may include Chrysothamnus viscidiflorus, Ephedra viridis, Eriogonum corymbosum, Holodiscus dumosus, Mahonia repens, Purshia tridentata, Ribes cereum, Rosa woodsii, Symphoricarpos oreophilus, and Tetradymia canescens. Occasional trees, such as Juniperus spp., Quercus spp., Pinus edulis, Pinus ponderosa or Pseudotsuga menziesii, may be scattered in some stands. The herbaceous layer is variable and may have sparse to moderate cover and is most often dominated by graminoids. Dominant graminoids may include Carex geyeri, Elymus lanceolatus ssp. lanceolatus, Hesperostipa comata, and Pseudoroegneria spicata. Other associated graminoids may include Achnatherum hymenoides, Koeleria macrantha, Leymus salinus, Poa fendleriana, and Poa fendleriana. Forbs do not contribute significant cover, but may be diverse in some stands. Associates may include Artemisia ludoviciana, Balsamorhiza sagittata, Collinsia parviflora, Hedeoma hispida, Lathyrus pauciflorus, and Symphyotrichum oblongifolium (= Aster oblongifolius).
Dynamics: Yake and Brotherson (1979) noted that regeneration of Amelanchier utahensis was most favorable on moderately deep, fine-textured soils of northerly-aspected sites. They noted that seedling herbivory by native ungulates was greater at such sites. Annual grasses were noted to have a negative effect on seedling densities. Amelanchier utahensis hybridizes with Amelanchier alnifolia, and distinctions between the two species may become blurred in areas where their ranges overlap. Cercocarpus montanus is a dominant understory species in several woodland and forests dominated by Juniperus deppeana, Juniperus monosperma, Juniperus osteosperma, Juniperus scopulorum, Pinus edulis, Pinus ponderosa, Pseudotsuga menziesii, Quercus gambelii, Quercus grisea, and Quercus x pauciloba. Cercocarpus montanus stands often occur in the more xeric habitat below these woodland and forest stands. In xeric habitats studied by Greenwood and Brotherson (1978), Cercocarpus montanus sites had significantly more rock than the Pinus edulis and Juniperus osteosperma sites. Brotherson et al. (1984) suggested that stands are moisture-limited because stands on southern aspects were always higher in elevation than stands on northern aspects. They also found that the Cercocarpus montanus stands were more likely to occur on northern slopes than on southern in central Utah. However, Cercocarpus montanus did not typically occur with mesic shrubs such as Symphoricarpos oreophilus and Amelanchier alnifolia. There is often a broad Cercocarpus montanus shrub/herbaceous ecotone between these shrublands and grasslands. Ecological factors that control shrub densities such as fire and drought need more investigation. Unlike other species of Cercocarpus, Cercocarpus montanus is a fire-resistant species because it can resprout from the base after a fire has killed the top (Cronquist et al. 1997). In the southern portion of its range, Cercocarpus montanus functions as an evergreen shrub by retaining leaves during mild winters and losing them during cold winters (Dick-Peddie 1993). Cercocarpus montanus is preferred winter range browse for deer (Roughton 1966, 1972, Hoffman and Alexander 1987). Stands can also produce significant forage that can be utilized by grazing livestock provided the slopes are not too steep (Hoffman and Alexander 1987). More information is needed on the effects of livestock grazing and wildlife browsing on the structure and function of these shrublands.
Environmental Description: Plant associations in this alliance occur at middle elevations (1850-2700 m) of mountains. Climate is semi-arid. Summers are hot and winters are typically cold, with freezing temperatures and snow common, however, stands on the southwestern U.S. may have mild winter temperatures. Precipitation ranges from 30-50 cm annually with a large proportion falling as winter snow. Sites are variable but are generally xeric and rocky with moderate to very steep slopes or on ridges. Stands are found on various aspects, but typically the higher elevation and northern sites are on warmer, southern exposures, and lower elevation and southern stands are restricted to more mesic north slopes. Environments include sheltered rocky canyons, mesa slopes, slickrock, moderate to very steep slopes and ridges. Soils are variable, from shallow and skeletal near rock outcroppings, to moderately deep with abundant organic matter. Soil textures range from sandy loam to clay. Soils range from slightly acidic to slightly alkaline depending on parent material, which includes alluvium, colluvium and residuum from igneous, metamorphic or sedimentary rocks such as granite, gneiss, limestone, quartz monzonite, rhyolite, sandstone, schist and shale.
Geographic Range: This alliance is known from the southern Rocky Mountains, Arizona-New Mexico mountains, north to the Wyoming Basins, east to escarpments of the Great Plains and west to the Colorado Plateau and parts of the Great Basin.
Nations: US
States/Provinces: AZ, CA, CO, NE, NM, NV, SD, TX, UT, WY
Plot Analysis Summary:
http://vegbank.org/natureserve/ELEMENT_GLOBAL.2.899784
Confidence Level: Low
Confidence Level Comments: No Data Available
Grank: GNR
Greasons: No Data Available
Concept Lineage: A.2620, A.896, A.2659, A.916.
Predecessors: No Data Available
Obsolete Names: No Data Available
Obsolete Parents: No Data Available
Synonomy: > Cercocarpus intricatus (Small leaf mountain mahogany scrub) Provisional Alliance (Sawyer et al. 2009) [76.300.00]
> Cercocarpus intricatus Shrubland Alliance (Evens et al. 2014)
> Cercocarpus intricatus Shrubland Alliance (Evens et al. 2014)
- Brotherson, J. D., and S. J. Barnes. 1984. Habitat relationships of Glaux maritima in central Utah. Great Basin Naturalist 44(2):299-309.
- Cronquist, A., N. H. Holmgren, and P. K. Holmgren. 1997. Intermountain flora: Vascular plants of the Intermountain West, USA. Volume 3, Part A, subclass Rosidae (except Fabeles). New York Botanical Garden, Bronx, NY. 446 pp.
- Dick-Peddie, W. A. 1993. New Mexico vegetation: Past, present, and future. University of New Mexico Press, Albuquerque. 244 pp.
- Evens, J. M., K. Sikes, D. Hastings, and J. Ratchford. 2014. Vegetation alliance descriptions for Lake Mead National Recreation Area, Death Valley National Park and Mojave National Preserve. Unpublished report submitted to USDI National Park Service, Mojave Desert Network Inventory and Monitoring Program. California Native Plant Society, Sacramento, CA.
- Faber-Langendoen, D., J. Drake, M. Hall, G. Kittel, S. Menard, C. Nordman, M. Pyne, M. Reid, M. Russo, K. Schulz, L. Sneddon, K. Snow, and J. Teague. 2013-2019b. Screening alliances for induction into the U.S. National Vegetation Classification: Part 1 - Alliance concept review. NatureServe, Arlington, VA.
- Greenwood, L. R., and J. D. Brotherson. 1978. Ecological relationships between pinyon-juniper and true mountain mahogany stands in the Uintah Basin, Utah. Journal of Range Management 31(3):164-167.
- Hoffman, G. R., and R. R. Alexander. 1987. Forest vegetation of the Black Hills National Forest of South Dakota and Wyoming: A habitat type classification. Research Paper RM-276. USDA Forest Service, Rocky Mountain Forest and Range Experiment Station, Fort Collins, CO. 48 pp.
- Roughton, R. D. 1966. Age structure of browse populations. Unpublished thesis, Colorado State University, Fort Collins. 154 pp.
- Roughton, R. D. 1972. Shrub age structures on a mule deer winter range in Colorado. Ecology 53(4):615-625.
- Sawyer, J. O., T. Keeler-Wolf, and J. Evens. 2009. A manual of California vegetation. Second edition. California Native Plant Society, Sacramento CA. 1300 pp.
- Thomas, K. A., T. Keeler-Wolf, J. Franklin, and P. Stine. 2004. Mojave Desert Ecosystem Program: Central Mojave vegetation mapping database. U.S. Geological Survey, Western Regional Science Center. 251 pp.
- Yake, S., and J. D. Brotherson. 1979. Differentiation of serviceberry habitats in the Wasatch Mountains of Utah. Journal of Range Management 32(4):379-386.