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F012 Temperate Grassland & Shrubland Formation

Type Concept Sentence: Temperate Grassland, Meadow & Shrubland is dominated by perennial grasses, forbs and shrubs typical of moderately dry to moist habitats and is found in the mid-latitude regions of all continents (23° to 55°N and S), varying from large open grassland landscapes to droughty hillside meadows in forested landscapes.


Common (Translated Scientific) Name: Temperate Grassland & Shrubland Formation

Colloquial Name: Temperate Grassland & Shrubland

Hierarchy Level:  Formation

Type Concept: Temperate grasslands are dominated by mesomorphic to somewhat xeromorphic perennial grasses with a variable amount of perennial forbs. Trees are scattered to largely absent (typically <10% cover), often controlled by drought, cold or fire. Grasses and forbs grow to different heights at maturity, depending on climate, soil and other factors. Grasses may be either tussock (bunch) forming or sod forming (forming a carpet on the ground), with bunchgrasses increasingly common in droughty conditions. Both C3 and C4 grasses are found, with C3 increasingly common towards the poles. Grasses and perennial forbs are well-suited to withstand cold seasons and tolerate grazing and fire because of the position of their renewal buds near or below the ground surface.

Diagnostic Characteristics: Mesomorphic perennial grasses and shrubs are the most common growth forms in this formation, with a variable amount of perennial forbs. Trees are scattered to largely absent (typically <10% cover). Stands are found in the mid-latitude regions of all continents, typically associated with semi-arid climates or stressful sites in moist climates in the mid-latitudes (23° to 55°N and S).

Rationale for Nominal Species or Physiognomic Features: No Data Available

Classification Comments: No Data Available

Similar NVC Types: No Data Available
note: No Data Available

Physiognomy and Structure: Mesomorphic perennial grasses and shrubs are the most common growth forms in this formation, with a variable amount of perennial forbs. Trees are scattered to largely absent (typically <10% cover). Grasses and forbs grow to different heights at maturity. Furthermore, some grasses and most forbs are erect, while others are recumbent, creeping along the ground. Thus a distinct layering of plant foliage occurs in some temperate grasslands. Forbs and grasses may grow and bloom at different times during the growing season, so temperate grasslands may have recognizable color phases, or aspects, depending on what plants are blooming. This is especially evident in the east European steppes. Grasses and perennial forbs are well-suited to withstand cold seasons and tolerate grazing and fire because of the position of their renewal buds near the ground surface. The plants are generally able to regenerate from these buds after seasonal die-downs, burns, or cropping by grazing animals (Woodward 2008). Among the grasses, both sod-forming and bunchgrasses are prominent throughout, but bunchgrasses become a much more frequent component of the vegetation in the drier short-grass regions.

Floristics: No Data Available

Dynamics:  Temperate grasslands around the world have been so heavily altered by human activities (especially grazing of livestock, plowing under and conversion to agriculture, increased fire frequency, or cessation of grass fires) that little survives that is truly natural. Fire may have been the first way people managed grasslands. Annual burning eliminated tree and shrub seedlings and stimulated the growth of new grass shoots. Deliberate burning may have occurred as soon as fire itself could be managed in order to attract the wild grazing animals upon which hunting and gathering peoples, and indeed, early agriculturalists, depended for protein. Repeated firing of the forest edge may have expanded grasslands into wetter climate regions, such as the Prairie Peninsula of the U.S. and the eastern pampas of Argentina and Uruguay. Later, farmers and townsfolk stopped all wildfires, and trees invaded the grasslands (Woodward 2008).

Pastoralism in all likelihood first developed in the temperate grasslands of the Old World. Shifts in plant species abundance no doubt accompanied the spread of pastoral societies, since livestock select the most palatable plants. Weedy grasses and forbs evolved in response to the trampling and cropping of the vegetation by cattle, sheep and other hoofed grazers. Tolerant of disturbance and preferring open habitat, weeds typically germinate rapidly in full sun, have short lifespans, produce large numbers of easily dispersed seeds, and thus spread quickly to new disturbed sites. Not surprisingly, Old World weeds followed cattle, sheep, and goats when the animals were transported to other parts of the world (Woodward 2008).

Environmental Description:  Climate: Temperate grasslands occur in regions with a semi-arid to moist climate (mostly BSk in the Koeppen climate classification). Semi-arid climate regions receive 25-50 cm of precipitation a year; moist regions receive 50-90 cm. Snow may occur in some areas, including mid-continental prairie of North America and the East European steppes of Eurasia). The continental location of grasslands in both North America and Eurasia results in the wide annual range in temperatures, with winters having temperatures well below freezing and summers very hot, and the growing season extending from 5-7 months (Woodward 2008).

Continental precipitation gradients shape the distribution of regional biomes. In some instances, such as in North America, longitudinal zones dominate and replace each other in an east-west direction across the mid-continent (i.e. tallgrass prairie to shortgrass prairie). In other areas, such as Eurasia, latitudinal zonation is more prominent, with biomes changing in a north-south direction. Still other natural grasslands are a consequence of rainshadows on the lee sides of major mountain ranges or of high elevation, or of specialized droughty substrates (Woodward 2008).

Soil/substrate/hydrology: Temperate grasslands are found on soils that are highly fertile and productive. Soils are built from the combination of dense roots and the annual die-off of above ground vegetation. More living biomass exists below ground than above. The mollisol soils (USDA Soil Taxonomy, also called chernozems) are rich in humus in both the A and B horizons. The soils often contain loess material, rich in carbonates (Woodward 2008).

Geographic Range: Temperate grasslands and shrublands occur in the middle latitudes (25-55°) of the northern and southern hemispheres. The largest expanses are found in North America ("prairies") and Eurasia ("steppe), with smaller but substantial areas in South America ("pampas") and southern Africa ("veld") (Woodward 2008).

Nations: AR,AU,BR,CA,CL,CN,LS,MN,MX,NZ,RU,US,UY,ZA

States/Provinces:  No Data Available



Confidence Level: Moderate

Confidence Level Comments: No Data Available

Grank: GNR

Greasons: No Data Available


Concept Lineage: No Data Available

Predecessors: No Data Available

Obsolete Names: No Data Available

Obsolete Parents: No Data Available

Synonomy: = Temperate Grassland (Woodward 2008) [Despite the name, shrublands found within the grassland regions are included in Woodward''s concept.]

Concept Author(s): Hierarchy Revisions Working Group, Federal Geographic Data Committee (Faber-Langendoen et al. 2014) after S. Woodward (2008)

Author of Description: D. Faber-Langendoen and J. Messick

Acknowledgements: No Data Available

Version Date: 10-17-14

  • Faber-Langendoen, D., T. Keeler-Wolf, D. Meidinger, C. Josse, A. Weakley, D. Tart, G. Navarro, B. Hoagland, S. Ponomarenko, J.-P. Saucier, G. Fults, and E. Helmer. 2015c. Classification and description of world formation types. General Technical Report RMRS-GTR-000. USDA Forest Service, Rocky Mountain Research Station, Fort Collins, CO.
  • Woodward, S. 2008. Grassland biomes. Greenwood Press, Westport, CT.