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A3572 Pinus edulis - Juniperus osteosperma Grassy Open Woodland Alliance

Type Concept Sentence: This open woodland and savanna alliance is characterized by diagnostic tree species Pinus edulis that forms a very open to moderately dense tree layer often with Juniperus osteosperma and an understory dominated by an open to dense layer of perennial grasses, lacking significant cover of shrubs. It occurs on dry mountain slopes, foothills, plateaus in the Colorado Plateau extending east into the west slope of the southern Rocky Mountains.


Common (Translated Scientific) Name: Two-needle Pinyon - Utah Juniper Grassy Open Woodland Alliance

Colloquial Name: Two-needle Pinyon - Utah Juniper Grassy Open Woodland

Hierarchy Level:  Alliance

Type Concept: This woodland alliance is characterized by diagnostic tree species Pinus edulis that forms a very open to moderately dense tree layer often with Juniperus osteosperma or, less frequently, Juniperus monosperma within the range of Juniperus osteosperma in northern Arizona. Juniperus osteosperma may also dominate stands as long as there is significant presence of Pinus edulis (not accidental) to characterize the stand as a pinyon-juniper stand and not the more xeric, typically lower elevation Juniperus osteosperma woodland and savanna. At higher elevations and relatively mesic sites, such as along drainages, Juniperus scopulorum may be present and sometimes dominant. Other conifers are absent or accidental with very low cover. The understory is characterized by an open to dense herbaceous layer. Scattered shrubs may be present but do not form a layer and do not exceed cover of the herbaceous layer. Perennial grasses typically dominate the herbaceous layer, although diverse forbs species are often present, but with low cover. Characteristic species include Achnatherum hymenoides, Bouteloua gracilis, Elymus elymoides, Hesperostipa comata, Hesperostipa neomexicana, Leymus salinus, Muhlenbergia pungens, Poa fendleriana, Poa secunda, Pleuraphis jamesii, and Pseudoroegneria spicata. The non-native, invasive annual grass Bromus tectorum becomes abundant in disturbed stands and may dominate the understory of highly disturbed stands. Some stands included in this alliance have been seeded with non-native perennial grasses such as Psathyrostachys juncea or Agropyron cristatum to control soil erosion or increase forage production and now have a semi-natural understory. This alliance occurs on warm, dry sites on mountain slopes, foothills, and plateaus in the Colorado Plateau extending east into the west slope of the southern Rocky Mountains. The climate of the region is semi-arid with drought not uncommon. Stands typically occur on nearly level to moderately steep, rocky slopes on hillsides and mesatops. Aspect does not seem important except in elevational extremes for a given latitude where low-elevation stands are restricted to the more mesic north slopes; canyons and high-elevation stands occur on south aspects. Sites are typically dry with shallow, rocky, calcareous and alkaline soils. Other sites include eroded "badlands," lava flows, scree slopes, and deep sands.

Diagnostic Characteristics: This woodland alliance occurs in the Colorado Plateau and extends east into the west slope of the southern Rocky Mountains within the distributional range of Juniperus osteosperma. The diagnostic tree species are Pinus edulis and Juniperus osteosperma, either of which may dominate woodland stands as long as there is significant presence of Pinus edulis (not accidental). The understory is characterized by an open to dense herbaceous layer. Characteristic species that may dominate include Achnatherum hymenoides, Bouteloua gracilis, Elymus elymoides, Hesperostipa comata, Hesperostipa neomexicana, Leymus salinus, Muhlenbergia pungens, Poa fendleriana, Poa secunda, Pleuraphis jamesii, and Pseudoroegneria spicata. The non-native, invasive annual grass Bromus tectorum and non-native perennial grasses such as Psathyrostachys juncea or Agropyron cristatum may become abundant in disturbed stands.

Rationale for Nominal Species or Physiognomic Features: No Data Available

Classification Comments: In northwestern New Mexico outside the distributional range of Juniperus osteosperma, this alliance transitions into the southern Rocky Mountain pinyon-juniper woodland alliances. The following associations in this alliance need further review and classification action and possibly rename Juniperus spp. to (Juniperus osteosperma) as they are not known outside the Juniperus osteosperma range or distribution: ~Pinus edulis - Juniperus spp. / Leymus salinus Woodland (CEGL002340)$$; ~Pinus edulis - Juniperus spp. / Poa fendleriana Woodland (CEGL000787)$$; and ~Pinus edulis - Juniperus spp. / Pseudoroegneria spicata Woodland (CEGL000788)$$.

Similar NVC Types: No Data Available
note: No Data Available

Physiognomy and Structure: Vegetation included in this alliance has an open to moderately dense tree canopy that is typically 3-10 m tall. Stands are either solely dominated by evergreen needle-leaved trees or may be codominated by scale-leaved evergreen trees. An open to moderate ground layer dominated by perennial graminoids is usually present. Perennial forbs and cacti are often scattered throughout the stands. Annual forbs and grasses may be seasonally present. Scattered shrubs may be present, but do not form a layer (<10% cover).

Floristics: This alliance is characterized by a very open to moderately dense tree canopy typically 3-12 m tall. The diagnostic tree species is Pinus edulis often with Juniperus osteosperma or, less frequently, Juniperus monosperma within the range of Juniperus osteosperma in northern Arizona. Juniperus osteosperma may also dominate stands as long as there is significant presence (>5% cover) of Pinus edulis to characterize the stand as a pinyon-juniper stand and not the more xeric, typically lower elevation Juniperus osteosperma woodland and savanna. At higher elevations and relatively mesic sites, such as along drainages, Juniperus scopulorum may be present and sometimes dominant. Other conifers are absent or accidental with very low cover. The understory is characterized by an open to dense herbaceous layer. Scattered shrubs may be present, such as Artemisia spp., Cercocarpus montanus, Ephedra viridis, Ericameria nauseosa, Fraxinus anomala, Gutierrezia sarothrae, Opuntia polyacantha, and Rhus aromatica, but do not form a layer and do not exceed cover of the herbaceous layer. Perennial grasses typically dominate the herbaceous layer, although diverse forbs species are often present, but with low cover. Characteristic species include Achnatherum hymenoides, Bouteloua gracilis, Elymus elymoides, Hesperostipa comata, Hesperostipa neomexicana, Leymus salinus, Muhlenbergia pungens, Poa fendleriana, Poa secunda, Pleuraphis jamesii, and Pseudoroegneria spicata. Commonly present forbs include species of Arenaria, Artemisia, Eriogonum, Heterotheca, Hymenoxys, Mirabilis, Oxytropis, Penstemon, Phlox, Senecio, Stenotus, and Zinnia. Annual grasses and forbs are seasonally present. The non-native, invasive annual grass Bromus tectorum becomes abundant in disturbed stands and may dominate the understory of highly disturbed stands. Some stands included in this alliance have been seeded with non-native perennial grasses such as Psathyrostachys juncea or Agropyron cristatum to control soil erosion or increase forage production and now have a semi-natural understory.

Dynamics:  Pinus edulis is extremely drought-tolerant and slow-growing (Little 1987, Powell 1988b, Muldavin et al. 1998c). It is also non-sprouting and may be killed by fire (Wright et al. 1979). The effect of fire on a stand is largely dependent on the tree height and density, fine-fuel load on the ground, weather conditions, and season (Dwyer and Pieper 1967, Wright et al. 1979). Trees are more vulnerable in open stands where fires frequently occur in the spring, when the relative humidity is low, wind speeds are over 10-20 mph, and there are adequate fine fuels to carry fire (Wright et al. 1979). Under other conditions, burns tend to be spotty with low tree mortality. Large trees are generally not killed unless fine fuels, such as tumbleweeds, have accumulated beneath the tree to provide ladder fuels for the fire to reach the crown (Jameson 1962). Closed-canopy stands rarely burn because they typically do not have enough understory or wind to carry a fire (Wright et al. 1979).

Although Pinus edulis is drought-tolerant, prolonged droughts will weaken trees and promote mortality by secondary agents. Periodic die-offs of pinyon pine caused by insects, such as the pinyon Ips beetle (Ips confusus), or fungal agents, such as blackstain root-rot (Leptographium wageneri), tend to be correlated with droughts (Anhold 2005). These mortality events may be localized or widespread but can result in 50 to 90% mortality of Pinus edulis (Harrington and Cobb 1988).

Climatic and other factors have resulted in denser and expanded pinyon-juniper stands throughout the Colorado Plateau and Great Basin. Denser stands are more susceptible to attack by insects and disease (Anhold 2005). In addition, altered fire regimes, cutting trees for fencing or firewood, and improper grazing by livestock have significant impacts on the quality of sites. Grazing by livestock can modify the fire regime by removing the fine fuels that carry fire. Fire, livestock grazing, and trampling by recreationalists and vehicles disturb cryptogamic soil crusts that help maintain soil structure, reduce soil erosion, provide habitat for plants and preserve biological diversity (Ladyman and Muldavin 1996). More study is needed to understand and manage these woodlands ecologically.

Environmental Description:  This woodland alliance occurs on warm, dry sites on canyon and mountain slopes, mesatops, foothills, and plateaus in the Colorado Plateau extending east into the west slope of the southern Rocky Mountains. Climate is semi-arid and droughts are not uncommon. Summers are generally hot, and winters are cold with occasional snows and extended periods of freezing temperatures. The seasonality of precipitation varies from east to west with summer rain more common in the southern and eastern portion of the alliance''s range and winter precipitation more common in the western portion of the range. Mean annual precipitation ranges from 30-46 cm. Elevations normally range from 1500-2465 m (4921-8087 feet). Stands occur on the lower and middle slopes of ridges, on benches, terraces and dunes. These sites range from moderately to steeply sloping (47% to more than 100% slopes), although a minority of stands occur on gentle slopes with gradients not exceeding 10%. Aspect does not seem important except in elevational extremes for a given latitude where low-elevation stands are restricted to the more mesic north slopes; canyons and high-elevation stands occur on south aspects. Soils are shallow to moderately deep, well-drained, often calcareous and alkaline with textures ranging from sand to clay depending on the underlying geology. Parent materials vary from shale, limestone and sandstone to metamorphic, and granitic rocks, eolian deposits, and basalt alluvium or colluvium overlying Moenkopi siltstone. Rocks and bare ground are often common and occupy most of the unvegetated surface, up to 90% in some stands and averaging 60%.

Geographic Range: The core distribution of this woodland and savanna alliance is the Colorado Plateau extending east into the west slope of the southern Rocky Mountains.

Nations: US

States/Provinces:  AZ, CO, NM, UT, WY?




Confidence Level: Moderate

Confidence Level Comments: No Data Available

Grank: GNR

Greasons: No Data Available

Type Name Database Code Classification Code
Class 1 Forest & Woodland Class C01 1
Subclass 1.B Temperate & Boreal Forest & Woodland Subclass S15 1.B
Formation 1.B.2 Cool Temperate Forest & Woodland Formation F008 1.B.2
Division 1.B.2.Nc Western North American Pinyon - Juniper Woodland & Scrub Division D010 1.B.2.Nc
Macrogroup 1.B.2.Nc.1 Singleleaf Pinyon - Utah Juniper - Western Juniper Intermountain Woodland Macrogroup M896 1.B.2.Nc.1
Group 1.B.2.Nc.1.a Two-needle Pinyon - Utah Juniper Woodland Group G900 1.B.2.Nc.1.a
Alliance A3572 Two-needle Pinyon - Utah Juniper Grassy Open Woodland Alliance A3572 1.B.2.Nc.1.a
Association CEGL000778 Two-needle Pinyon - (Utah Juniper) / Blue Grama Woodland CEGL000778 1.B.2.Nc.1.a
Association CEGL000787 Two-needle Pinyon - Juniper species / Muttongrass Woodland CEGL000787 1.B.2.Nc.1.a
Association CEGL000788 Two-needle Pinyon - Juniper species / Bluebunch Wheatgrass Woodland CEGL000788 1.B.2.Nc.1.a
Association CEGL002340 Two-needle Pinyon - Juniper species / Saline Wildrye Woodland CEGL002340 1.B.2.Nc.1.a
Association CEGL002364 Two-needle Pinyon - Utah Juniper / Indian Ricegrass Woodland CEGL002364 1.B.2.Nc.1.a
Association CEGL002367 Two-needle Pinyon - Utah Juniper / Cheatgrass Ruderal Woodland CEGL002367 1.B.2.Nc.1.a
Association CEGL002368 Two-needle Pinyon - Utah Juniper / Russian Wildrye Ruderal Woodland CEGL002368 1.B.2.Nc.1.a
Association CEGL002371 Two-needle Pinyon - Utah Juniper / New Mexico Feathergrass Woodland CEGL002371 1.B.2.Nc.1.a
Association CEGL002373 Two-needle Pinyon - Utah Juniper / Sandhill Muhly Woodland CEGL002373 1.B.2.Nc.1.a
Association CEGL002379 Two-needle Pinyon - Utah Juniper / James'' Galleta Woodland CEGL002379 1.B.2.Nc.1.a
Association CEGL002819 Two-needle Pinyon - Utah Juniper / Bluebunch Wheatgrass - Cushion Plant Woodland CEGL002819 1.B.2.Nc.1.a
Association CEGL005652 Two-needle Pinyon - Utah Juniper / Needle-and-Thread Open Woodland CEGL005652 1.B.2.Nc.1.a

Concept Lineage: This alliance is composed of 10 herbaceous understory associations from Old Alliance II.A.4.N.a. Pinus edulis - (Juniperus spp.) Woodland Alliance (A.516).

Predecessors: No Data Available

Obsolete Names: No Data Available

Obsolete Parents: No Data Available

Synonomy: > Pinus edulis Series (Francis 1986)
> Colorado Pinyon-Utah Juniper Series (Dick-Peddie 1993)

Concept Author(s): K.A. Schulz, in Faber-Langendoen et al. (2013)

Author of Description: K.A. Schulz

Acknowledgements: We have incorporated significant descriptive information previously compiled by J. Coles.

Version Date: 03-14-14

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