Print Report
D010 Pinus monophylla - Pinus edulis - Juniperus spp. Woodland & Scrub Division
Type Concept Sentence: These pinyon pine- and juniper-dominated woodlands, scrub, and savannas generally occur just above semi-desert shrublands and grasslands or shortgrass prairies and below montane forest vegetation throughout the semi-arid Intermountain West and western Great Plains of North America.
Common (Translated Scientific) Name: Singleleaf Pinyon - Two-needle Pinyon - Juniper species Woodland & Scrub Division
Colloquial Name: Western North American Pinyon - Juniper Woodland & Scrub
Hierarchy Level: Division
Type Concept: This division is composed of woodland, savanna, and scrub characterized by an open to closed tree and shrub canopy of Pinus monophylla, Pinus edulis, Juniperus occidentalis, Juniperus osteosperma, Juniperus monosperma, or Juniperus scopulorum. Typically one, or combinations of two, of these species dominate the canopy. Stands are typically open-canopied (10-30% cover), but closed-canopy conditions with a sparse understory are not uncommon. Pinyon and juniper stands may occur as persistent woodlands, open savannas, and wooded shrublands. Persistent woodlands occur where climate substrates support pinyon pines and junipers; typically with infrequent wildfire. Open savannas occur where landform and substrates support both trees and grasses, and where more frequent surface fires were likely the historical norm. Wooded shrublands occur where climate and soils support shrublands, but where trees increase in abundance under favorable climate and limited disturbance, and decrease during droughts and following disturbance. Understory layers are variable and may be dominated by shrubs, graminoids, or be absent, especially on rocky substrates. Except for in the extreme eastern and southern portion of its range, shrub layers are frequently dominated by Artemisia tridentata, which in places can form a moderately dense shrub canopy. Other common associated shrub species include Arctostaphylos patula, Artemisia arbuscula, Artemisia bigelovii, Artemisia nova, Atriplex canescens, Cercocarpus intricatus, Cercocarpus ledifolius (in tree or shrub form), Cercocarpus montanus, Chrysothamnus viscidiflorus, Coleogyne ramosissima, Ericameria nauseosa, Fraxinus anomala, Fraxinus cuspidata, Glossopetalon spinescens, Krascheninnikovia lanata, Purshia stansburiana, Purshia tridentata, Quercus gambelii, Quercus turbinella, Quercus x pauciloba, Ribes cereum, Rhus trilobata, Tetradymia spp., Yucca baccata, and Yucca glauca. The most frequent succulents are Cylindropuntia imbricata, Opuntia phaeacantha, and Opuntia polyacantha. The herbaceous layer may be sparse to dense depending on overstory density, substrate, landscape position, and disturbance history, with the densest graminoid layer in open-tree or shrub savanna. Common graminoid associates include Bouteloua curtipendula, Bouteloua gracilis, Carex filifolia, Festuca idahoensis, Hesperostipa comata, Leymus cinereus, Leymus salinus, Pleuraphis jamesii, Pseudoroegneria spicata, Poa fendleriana, and Poa secunda. Forb species may be diverse but typically have low canopy cover values, and can include Astragalus spp., Castilleja integra, Cryptantha cinerea var. jamesii, Erigeron divergens, Eriogonum jamesii, Hymenopappus filifolius, Ipomopsis multiflora, Mentzelia spp., Penstemon spp., and Petradoria pumila.
Diagnostic Characteristics: Structurally complex, low woodland, scrub, and savanna with pinyon pine and or juniper dominance in uppermost canopy, especially Pinus monophylla, Pinus edulis, Juniperus occidentalis, Juniperus osteosperma, Juniperus monosperma, or Juniperus scopulorum. The understory is dominated by either shrub or grass species of cool-temperate affinity.
Rationale for Nominal Species or Physiognomic Features: No Data Available
Classification Comments: This division occurs over a very broad biogeographic range with many diagnostic species. The southern end of the range of this division occurs over a very broad transition zone with 1.B.1.Nd ~Madrean-Balconian Forest & Woodland Division (D060)$$ which is distinguished by the presence of other Madrean tree species, such as Juniperus coahuilensis, Juniperus deppeana, Juniperus pinchotii, Pinus cembroides, Pinus discolor, or evergreen oaks such as Quercus emoryi, Quercus grisea, or Quercus mohriana along with Madrean grasses, forbs, and characteristically warm-temperate subshrubs such as Nolina spp. and Dasylirion spp. Classifying open juniper savanna stands from grasslands with scattered trees is challenging, especially where juniper trees are invading grasslands. The invasive juniper savanna stands are typically characterized by younger, shorter (<3 m tall), pointed-crown trees of lower density.
Similar NVC Types: No Data Available
note: No Data Available
Physiognomy and Structure: This broadly defined evergreen division is composed of a woodland, savanna (or open woodland), and scrub structure. Stands are typically short (2-10 m tall), with an open to closed (10-60% cover), evergreen needle-leaved or scale-leaved or broad-leaved, sclerophyllous tree and/or shrub canopy. The understory is variable with lush grass cover and occasionally scattered shrubs in the savanna stands to a sparse to dense short-shrub layer and/or herbaceous layer in woodland stands. This division encompasses savanna that has widely spaced, short (2-10 m tall), mature (>150-year-old) trees with a moderately dense to dense herbaceous layer dominated by perennial graminoids. On extremely xeric sites, diagnostic juniper and pinyon trees species may only attain 2 m in height and have a more shrub form. However, Juniperus occidentalis-dominated stands have two different tree canopy structures: (1) an old-growth woodland with large, fairly well-spaced trees with rounded crowns, and (2) relatively young, often dense junipers trees with pointed crowns. Cover of understory species sharply declines when tree canopy cover exceeds 40% (Young et al. 1982). Many of the tree savannas have a sparse shrub layer present. Herbaceous layers are variable depending on the density of woody canopy, substrate, landscape position, and disturbance history. Perennial graminoids typically dominate most herbaceous layers with most species individually contributing low cover.
Floristics: This division includes woodland, savanna (or open woodland), and scrub characterized by an open to closed tree and shrub canopy of Pinus monophylla and/or Pinus edulis, Juniperus occidentalis, Juniperus osteosperma, Juniperus monosperma, or Juniperus scopulorum. Typically one, or combinations of two, of these species dominate the canopy. In the Mohave Desert mountains, Yucca brevifolia may be an associate of Pinus monophylla. Shrub species include Arctostaphylos patula, Artemisia arbuscula, Artemisia bigelovii, Artemisia nova, Artemisia tridentata, Atriplex canescens, Cercocarpus intricatus, Cercocarpus ledifolius (in tree or shrub form), Cercocarpus montanus, Chrysothamnus viscidiflorus, Coleogyne ramosissima, Ericameria nauseosa, Fraxinus anomala, Fraxinus cuspidata, Glossopetalon spinescens, Krascheninnikovia lanata, Purshia stansburiana, Purshia tridentata, Quercus gambelii, Quercus turbinella, Quercus x pauciloba, Ribes cereum, Rhus trilobata, Tetradymia spp., Yucca baccata, Yucca brevifolia, and Yucca glauca. Shrub species more characteristic of 1.B.1.Nc ~Californian Forest & Woodland Division (D007)$$, such as Juniperus californica, Quercus chrysolepis, and/or Quercus john-tuckeri, may occur near the southwestern limits of the range. The most frequent succulents are Cylindropuntia imbricata, Opuntia phaeacantha, and Opuntia polyacantha. The herbaceous layer may be sparse to dense depending on overstory density, substrate, landscape position, and disturbance history, with the densest graminoid layer in open-tree or shrub savanna. Common graminoid associates include Bouteloua curtipendula, Bouteloua eriopoda, Bouteloua gracilis, Carex filifolia, Hesperostipa comata, Hesperostipa neomexicana, Festuca idahoensis, Leymus cinereus (= Elymus cinereus), Leymus salinus, Muhlenbergia pauciflora, Pleuraphis jamesii, Poa fendleriana, Poa secunda, Pseudoroegneria spicata, and the non-native invasive annual Bromus tectorum. Forb species may be diverse but typically have low canopy cover values, and can include Astragalus spp., Castilleja integra, Cryptantha cinerea var. jamesii (= Cryptantha jamesii), Eriogonum jamesii, Erigeron divergens, Hymenopappus filifolius, Ipomopsis multiflora, Mentzelia spp., Penstemon spp., and Petradoria pumila.
Dynamics: Key dynamic processes are drought, fire, herbivory, and insect/disease outbreaks. Characteristic Pinus spp. and Juniperus spp. are relatively short (generally <15 m tall), shade-intolerant, drought-tolerant, slow-growing, long-lived trees (especially Juniperus osteosperma can reach 650 years old). Pinus spp. are non-sprouting and may be killed by fire. The effect of a fire on these stands is largely dependent on the tree height and density, fine fuel load on the ground, weather conditions and season. Large trees generally survive unless the fire gets into the crown due to heavy fuel loads in the understory. Fire acts to open stands, increase diversity and productivity in understory species, and create a mosaic of stands of different sizes and ages across the landscape while maintaining the boundary between woodlands and adjacent shrublands or grasslands.
As modeled by Landfire, this division is generally characterized by a spectrum of fire regimes, including frequent non-lethal fires, mixed-severity mosaic fires (mean FRI of 50-200 years), and very infrequent replacement fires (mean FRI of 200-500 years). Surface fire was likely the most frequent in savannas. Frequently, fire spreads from adjacent vegetation. Severe climatic events occurring during the growing season, such as frosts and drought, are thought to limit the distribution and density of pinyon and/or juniper stands to relatively narrow altitudinal belts on mountainsides and foothills. Weather-related stress thins trees in more closed stands. Insects/disease has a similar effect, but with a greater frequency in closed stands (mean return interval of 100 years) than open ones (mean return interval of 1000 years). Competition from grasses and older trees in late open stands is also included as a disturbance that maintains open woody canopies.
As modeled by Landfire, this division is generally characterized by a spectrum of fire regimes, including frequent non-lethal fires, mixed-severity mosaic fires (mean FRI of 50-200 years), and very infrequent replacement fires (mean FRI of 200-500 years). Surface fire was likely the most frequent in savannas. Frequently, fire spreads from adjacent vegetation. Severe climatic events occurring during the growing season, such as frosts and drought, are thought to limit the distribution and density of pinyon and/or juniper stands to relatively narrow altitudinal belts on mountainsides and foothills. Weather-related stress thins trees in more closed stands. Insects/disease has a similar effect, but with a greater frequency in closed stands (mean return interval of 100 years) than open ones (mean return interval of 1000 years). Competition from grasses and older trees in late open stands is also included as a disturbance that maintains open woody canopies.
Environmental Description: Stands of this division in the Colorado Plateau, Great Basin, and Mohave Desert mountains occur between 1500 and 2600 m elevation on warm, dry sites of lower mountain slopes, hills, mesas, plateaus, ridges, and more recently on basins and flats where trees are expanding into semi-desert grasslands and steppe. Substrates are variable, but are generally shallow, cobbly, gravelly, or sandy loams to clay loam or clay. Parent materials are variable. Juniper stands in the Columbia Plateau range from under 200 m elevation along the Columbia River in central Washington to over 1500 m. In central Oregon, they occur on all aspects and slope positions. Stands of this division also occur in dry mountains and foothills in southern Colorado south into central New Mexico, extending east into the plains on breaks in the southwestern Great Plains. They are found in dry sites in lower slopes of mountains, plateaus and foothills and on limestone and shale breaks in the plains. In this portion of the range, stands are found at elevations from 1370 to 2900 m. Climate is cool-temperate. Severe weather events occurring during the growing season, such as frosts and drought, are thought to limit the distribution of pinyon-juniper woodlands to relatively narrow altitudinal belts on a given mountainside, and particularly influence the proportion of pinyon trees relative to juniper.
Geographic Range: This division generally occurs just above semi-desert shrublands and grasslands or shortgrass prairies and below montane forest vegetation throughout the semi-arid Intermountain West and western Great Plains of North America.
Nations: US
States/Provinces: AZ, CA, CO, ID, MT, NM, NV, OK, OR, TX, UT, WA, WY
Plot Analysis Summary:
http://vegbank.org/natureserve/ELEMENT_GLOBAL.2.860343
Confidence Level: High
Confidence Level Comments: No Data Available
Grank: GNR
Greasons: No Data Available
Type | Name | Database Code | Classification Code |
---|---|---|---|
Class | 1 Forest & Woodland Class | C01 | 1 |
Subclass | 1.B Temperate & Boreal Forest & Woodland Subclass | S15 | 1.B |
Formation | 1.B.2 Cool Temperate Forest & Woodland Formation | F008 | 1.B.2 |
Division | 1.B.2.Nc Western North American Pinyon - Juniper Woodland & Scrub Division | D010 | 1.B.2.Nc |
Macrogroup | 1.B.2.Nc.1 Singleleaf Pinyon - Utah Juniper - Western Juniper Intermountain Woodland Macrogroup | M896 | 1.B.2.Nc.1 |
Macrogroup | 1.B.2.Nc.2 Two-needle Pinyon - One-seed Juniper Southern Rocky Mountain Woodland Macrogroup | M897 | 1.B.2.Nc.2 |
Concept Lineage: No Data Available
Predecessors: No Data Available
Obsolete Names: No Data Available
Obsolete Parents: No Data Available
Synonomy: No Data Available
- Barbour, M. G., and J. Major, editors. 1988. Terrestrial vegetation of California: New expanded edition. California Native Plant Society, Special Publication 9, Sacramento. 1030 pp.
- Belnap, J., J. Kaltenecker, R. Rosentreter, J. Williams, S. Leonard, and D. Eldridge. 2001. Biological soil crusts: Ecology and management. Technical Report 1730-2. USDI Bureau of Land Management. 110 pp.
- Blackburn, W. H., and P. T. Tueller. 1970. Pinyon and juniper invasion in black sagebrush communities in east-central Nevada. Ecology 51:841-848.
- Bradley, A. F., N. V. Noste, and W. C. Fischer. 1992a. Fire ecology of forests and woodlands in Utah. General Technical Report INT-287. USDA Forest Service, Intermountain Research Station, Ogden, UT. 128 pp.
- Brown, D. E., C. H. Lowe, and C. P. Pase. 1979. A digitized classification system for the biotic communities of North America with community (series) and association examples for the Southwest. Journal of the Arizona-Nevada Academy of Science 14:1-16.
- Brown, D. E., editor. 1982a. Biotic communities of the American Southwest-United States and Mexico. Desert Plants Special Issue 4(1-4):1-342.
- Dick-Peddie, W. A. 1993. New Mexico vegetation: Past, present, and future. University of New Mexico Press, Albuquerque. 244 pp.
- Dwyer, D. D., and R. D. Pieper. 1967. Fire effects on blue gramma-pinyon-juniper rangeland in New Mexico. Journal of Range Management 20:359-362.
- Eyre, F. H., editor. 1980. Forest cover types of the United States and Canada. Society of American Foresters, Washington, DC. 148 pp.
- Faber-Langendoen, D., J. Drake, S. Gawler, M. Hall, C. Josse, G. Kittel, S. Menard, C. Nordman, M. Pyne, M. Reid, L. Sneddon, K. Schulz, J. Teague, M. Russo, K. Snow, and P. Comer, editors. 2010-2019a. Divisions, Macrogroups and Groups for the Revised U.S. National Vegetation Classification. NatureServe, Arlington, VA. plus appendices. [in preparation]
- Floyd, M. L., D. D. Hanna, and G. Salamacha. 2001. Post-fire treatment of noxious weeds in Mesa Verde National Park, Colorado. Pages 147-157 in: C. Van Riper, III, et al., editors. Proceedings of the 5th Biennial Conference of Research on the Colorado Plateau, U.S. Geological Survey / FRESC Report Series USGSFRESC/COPL/2001/24. [http://www.rmrs.nau.edu/lab/people/csieg/sieg_pubs/Exotic_and%20_noxious_plants.pdf].
- Floyd, M. L., W. H. Romme, D. D. Hanna, M. Winterowd, D. Hanna, and J. Spence. 2008. Fire history of pinon-juniper woodlands on Navajo Point, Glen Canyon National Recreation Area. Natural Areas Journal 28:26-36.
- Franklin, J. F., and C. T. Dyrness. 1973. Natural vegetation of Oregon and Washington. General Technical Report PNW-8. USDA Forest Service, Pacific Northwest Forest and Range Experiment Station, Portland, OR. 417 pp.
- Gottfried, G. J., T. W. Swetnam, C. D. Allen, J. L. Betancourt, and A. L. Chung-MacCoubrey. 1995. Pinyon-juniper woodlands. Pages 95-132 in: D. M. Finch and J. A. Tainter, editors. Ecology, diversity, and sustainability of the middle Rio Grande Basin. General Technical Report RM. 268. USDA Forest Service, Rocky Mountain Forest and Range Experiment Station, Fort Collins, CO.
- Gruell, G. E., L. E. Eddleman, and R. Jaindl. 1994. Fire history of the pinyon-juniper woodlands of Great Basin National Park. NPS/PNROSU/NRTR-94/01. National Park Service, Pacific Northwest Region, Seattle, WA. 27 pp.
- Holland, V. L., and D. J. Keil. 1995. California vegetation. Kendall/Hunt Publishing Company, Dubuque, IA. 516 pp.
- Knight, D. H. 1994. Mountains and plains: Ecology of Wyoming landscapes. Yale University Press, New Haven, MA. 338 pp.
- LANDFIRE [Landfire National Vegetation Dynamics Database]. 2007a. Landfire National Vegetation Dynamics Models. Landfire Project, USDA Forest Service, U.S. Department of Interior. (January - last update) [http://www.LANDFIRE.gov/index.php] (accessed 8 February 2007).
- Ladyman, J. A. R., and E. Muldavin. 1996. Terrestrial cryptograms of pinyon-juniper woodlands in the southwestern United States: A review. General Technical Report RM-GTR-280. USDA Forest Service, Rocky Mountain Forest and Range Experiment Station, Fort Collins, CO. 33 pp.
- Larson, M., and W. H. Moir. 1987. Forest and woodland habitat types of northern New Mexico and northern Arizona. Edition 2. USDA Forest Service, Southwestern Region, Albuquerque, NM.
- Little, E. L. 1987. Pinyon trees (Pinus edulis) remeasured after 47 years. Pages 65-68 in: Proceedings - pinyon-juniper conference. General Technical Report INT-215. USDA Forest Service, Intermountain Research Station, Ogden, UT.
- Margolis, E. Q. 2014. Fire regime shift linked to increased forest density in a pinyon-juniper savanna landscape. International Journal of Wildland Fire 23:234-245.
- Meeuwig, R. O., and R. L. Bassett. 1983. Pinyon-juniper. Pages 84-86 in: R. M. Burns, compiler. Silvicultural systems for the major forest types of the United States. Agriculture Handbook No. 445. USDA Forest Service, Washington, DC.
- Muldavin, E., A. Kennedy, C. Jackson, P. Neville, T. Neville, K. Schulz, and M. Reid. 2011b. Vegetation classification and map: Bandelier National Monument. Natural Resource Technical Report NPS/SCPN/NRTR--2011/438. National Park Service, Fort Collins, CO.
- Muldavin, E., C. Baisan, T. Swetnam, L. DeLay, and K. Morino. 2003b. Woodland fire history studies in the Oscura and northern San Andres mountains, White Sands Missile Range, New Mexico. Natural Heritage New Mexico Publication No. 03-GTR-256, WSMR document No. 92F018, Albuquerque, NM.
- Neely, B., P. Comer, C. Moritz, M. Lammerts, R. Rondeau, C. Prague, G. Bell, H. Copeland, J. Humke, S. Spakeman, T. Schulz, D. Theobald, and L. Valutis. 2001. Southern Rocky Mountains: An ecoregional assessment and conservation blueprint. Prepared by The Nature Conservancy with support from the U.S. Forest Service, Rocky Mountain Region, Colorado Division of Wildlife, and Bureau of Land Management.
- Romme, W. H., C. D. Allen, J. D. Balley, W. L. Baker, B. T. Bestelmeyer, P. M. Brown, K. S. Eisenhart, M. L. Floyd, D. W. Huffman, B. F. Jacobs, R. F. Miller, E. H. Muldavin, T. W. Swetnam, R. J. Tausch, and P. J. Weisberg. 2009. Historical and modern disturbance regimes, stand structures, and landscape dynamics in pinon-juniper vegetation of the western United States. Rangeland Ecology & Management 62:203-222.
- Sawyer, J. O., T. Keeler-Wolf, and J. Evens. 2009. A manual of California vegetation. Second edition. California Native Plant Society, Sacramento CA. 1300 pp.
- Shiflet, T. N., editor. 1994. Rangeland cover types of the United States. Society for Range Management. Denver, CO. 152 pp.
- Stuever, M. C., and J. S. Hayden. 1997b. Plant associations of Arizona and New Mexico. Volume 2: Woodlands. USDA Forest Service, Southwestern Region, Habitat Typing Guides. 196 pp.
- Thorne, R. F., A. A. Schoenherr, C. D. Clements, and J. A. Young. 2007. Transmontane coniferous vegetation. Pages 574-586 in: M. G. Barbour, T. Keeler-Wolf, and A. A. Schoenherr, editors. Terrestrial vegetation of California. Third edition. University of California Press, Berkeley.
- Tuhy, J., P. Comer, D. Dorfman, M. Lammert, B. Neely, L. Whitham, S. Silbert, G. Bell, J. Humke, B. Baker, and B. Cholvin. 2002. An ecoregional assessment of the Colorado Plateau. The Nature Conservancy, Moab Project Office. 112 pp. plus maps and appendices.
- West, N. E. 1999a. Juniper-pinon savannas and woodlands of western North America. Pages 288-308 in: R. C. Anderson, editor. Savannas, barrens, and rock outcrop plant communities of North America. Cambridge University Press, New York.
- West, N. E. 1999b. Distribution, composition, and classification of current juniper-pinyon woodlands and savannas across western North America. Pages 20-23 in: S. B. Monsen and R. Stevens, editors. Proceedings: Ecology and management of pinyon-juniper communities within the Interior West. Proceedings RMRS-P-9. USDA Forest Service, Rocky Mountain Research Station, Ogden, UT.
- West, N. E., R. J. Tausch, and P. T. Tueller. 1998. A management-oriented classification of pinyon-juniper woodlands of the Great Basin. General Technical Report RMRS-GTR-12. USDA Forest Service, Rocky Mountain Research Station, Ogden, UT. 42 pp.
- West, N. E., and J. A. Young. 2000. Intermountain valleys and lower mountain slopes. Pages 255-284 in: M. G. Barbour and W. D. Billings, editors. North American Terrestrial Vegetation, second edition. Cambridge University Press, Cambridge.
- Wright, H. A., L. F. Neuenschwander, and C. M. Britton. 1979. The role and use of fire in sagebrush-grass and pinyon-juniper plant communities: A state of the art review. General Technical Report INT-58. USDA Forest Service, Intermountain Forest and Range Experiment Station, Ogden, UT.
- Young, J. A., R. A. Evans, J. D. Budy, and A. Torell. 1982. Cost of controlling maturing western juniper trees. Journal of Range Management 35(4):437-442.