Print Report
A3733 Cercocarpus montanus - Quercus x pauciloba Shrubland Alliance
Type Concept Sentence: This alliance is characterized by shrublands dominated or codominated by Cercocarpus montanus and Quercus x pauciloba occurring in the southern Rocky Mountains south to the northern Chihuahuan Desert and east to the Southern Shortgrass Prairie with outlying occurrences in adjacent ecoregions.
Common (Translated Scientific) Name: Alderleaf Mountain-mahogany - Wavyleaf Oak Shrubland Alliance
Colloquial Name: Alderleaf Mountain-mahogany - Wavyleaf Oak Shrubland
Hierarchy Level: Alliance
Type Concept: The vegetation of this alliance is characterized by shrublands most commonly dominated by Cercocarpus montanus, but in the southern portion of its range is codominated by Quercus x pauciloba. Structure varies from dense to having little to no canopy overlap with lush herbaceous layers. Shrub associates may include Ceanothus greggii, Dasylirion leiophyllum, Quercus grisea (shrub form), and Quercus turbinella. The understory is most commonly graminoid-dominated by Bouteloua curtipendula, Hesperostipa neomexicana, Muhlenbergia montana, or Petrophytum caespitosum. Additional species may include Achnatherum scribneri, Bouteloua gracilis, Carex geyeri, Elymus albicans, Festuca arizonica, and the perennial forbs Helianthus pumilus, Heterotheca villosa, Potentilla fissa, and Scutellaria brittonii. Introduced annual grass Bromus tectorum is often present. This alliance primarily occurs in the southern Rocky Mountains, northern Chihuahuan Desert and east to the Southern Shortgrass Prairie, but has outlying occurrences in adjacent ecoregions. It occurs on montane slopes at 915 to 2440 m (3000-8000 feet) elevation. Stands are found on various aspects, but typically the higher elevation and northern sites are on warmer, southern exposures, and lower elevation and southern stands are restricted to more mesic north slopes.
Diagnostic Characteristics: Broad-leaved shrublands dominated or codominated by the hybrid scrub oak Quercus x pauciloba and Cercocarpus montanus with grasses predominant in the understory. Understory associated species tend to reflect southern distributions.
Rationale for Nominal Species or Physiognomic Features: No Data Available
Classification Comments: This alliance includes Cercocarpus montanus associations previously included in one larger Rocky Mountain alliance, but has been created separately to reflect the more southern floristic elements of this new alliance. Stands in this alliance appear to be separated from stands in ~Cercocarpus montanus Madrean Montane Chaparral Alliance (A3791)$$ by the density of the shrub layer alone. Stands in this shrub-herbaceous alliance have only been described from southwestern New Mexico but may occur throughout the range of Cercocarpus montanus. A Cercocarpus montanus / Stipa neomexicana shrubland has been described from the Colorado Front Range foothills by the Colorado Natural Heritage Program but has not yet been included in the National Vegetation Classification (S. Kettler pers. comm.).
Similar NVC Types: No Data Available
note: No Data Available
Physiognomy and Structure: Vegetation included in this alliance has a moderately sparse to moderately dense cover of broad-leaved deciduous shrubs 2 m tall. The sparse to moderately dense herbaceous layer is usually less than 1 m tall and dominated by graminoids. Some stands have herbaceous layers dominated by perennial forbs, but generally forb cover is sparse. Annuals are seasonally present.
Floristics: The sparse to moderately dense shrub layer is 1-2 m tall and is dominated by the cold-deciduous, broad-leaved shrub Cercocarpus montanus or the hybrid oak Quercus x pauciloba. Occasional trees, such as Juniperus spp., Pinus edulis, Pinus ponderosa, Pseudotsuga menziesii, or Quercus spp., may be scattered in some stands. Shrub cover is generally denser on northern aspects and increases with elevation. Lower elevation stands grade into shrub savannas with decreasing elevation. The herbaceous layer is sparse to moderately dense depending on the density of the shrub layer and soil moisture. It is dominated by perennial graminoids or forbs with annuals present seasonally. Associated species vary with location. In the western Great Plains, Artemisia frigida and Rhus trilobata are common woody species. The herbaceous layer is sparse to moderately dense and is dominated by the warm-season, perennial midgrass Bouteloua curtipendula. Achnatherum hymenoides (= Oryzopsis hymenoides), Aristida purpurea, Elymus elymoides, Hesperostipa comata (= Stipa comata), and Piptatheropsis micrantha (= Oryzopsis micrantha) may also be present. Common forbs are Artemisia ludoviciana, Hedeoma hispida, and Symphyotrichum oblongifolium (= Aster oblongifolius). Stands in the Rocky Mountain foothills are similar, but with the graminoid layer usually dominated by the cool-season, perennial mid grasses Bouteloua gracilis, Hesperostipa comata, or Hesperostipa neomexicana (= Stipa neomexicana). In addition to Artemisia frigida and Rhus trilobata, associated shrubs and dwarf-shrubs may include Purshia tridentata, Quercus gambelii, and Ribes cereum. Other associated species, such as Allium textile, Artemisia ludoviciana, Astragalus parryi, Bouteloua gracilis, Eriogonum umbellatum, Helianthus pumilus, and Poa spp., are common in the herbaceous layer. In the Chihuahuan Desert montane scrublands, the dominant species may be a different subspecies, Cercocarpus montanus var. paucidentatus. Associated shrubs and dwarf-shrubs include Madrean elements such as Agave parryi, Ceanothus greggii, Dasylirion leiophyllum, Dasylirion wheeleri, Ericameria laricifolia, Garrya flavescens, Garrya wrightii, Petrophytum caespitosum, Quercus grisea (shrub form), Quercus turbinella, Yucca baccata, and shrubby species of Opuntia, sometimes forming a mixed cold-deciduous/evergreen stand. Occasional trees such as Quercus arizonica, Quercus gambelii, and other Quercus spp. and conifers such as Juniperus deppeana may be present. The herbaceous layer is sparse on the very rocky sites to moderately dense, and usually is dominated by perennial grasses, including Aristida ternipes, Bothriochloa barbinodis, Bouteloua curtipendula, Bouteloua gracilis, Eragrostis intermedia, Koeleria macrantha, Lycurus phleoides, Muhlenbergia emersleyi, Muhlenbergia montana, and Muhlenbergia pauciflora. Forbs may be diverse and include Artemisia ludoviciana, Eriogonum jamesii, Hedeoma plicata, Heliomeris longifolia, Heliomeris multiflora, Mirabilis multiflora, and Viguiera dentata.
Dynamics: Cercocarpus montanus is a dominant understory species in several woodland and forests dominated by Juniperus deppeana, Juniperus monosperma, Juniperus osteosperma, Juniperus scopulorum, Pinus edulis, Pinus ponderosa, Pseudotsuga menziesii, Quercus gambelii, Quercus grisea, and Quercus x pauciloba. Cercocarpus montanus stands often occur in the more xeric habitat below these woodland and forest stands. In xeric habitats studied by Greenwood and Brotherson (1978), Cercocarpus montanus sites had significantly more rock than the Pinus edulis and Juniperus osteosperma sites. Brotherson et al. (1984) suggested that stands are moisture-limited because stands on southern aspects were always higher in elevation than stands on northern aspects. However, Cercocarpus montanus did not typically occur with mesic shrubs such as Symphoricarpos oreophilus and Amelanchier alnifolia. There is often a broad Cercocarpus montanus shrub/herbaceous ecotone between these shrublands and grasslands. Ecological factors that control shrub densities such as fire and drought need more investigation. Unlike other species of Cercocarpus, Cercocarpus montanus is a fire-resistant species because it can resprout from the base after a fire has killed the top (Cronquist et al. 1997). In the southern portion of its range, Cercocarpus montanus functions as an evergreen shrub by retaining leaves during mild winters and losing them during cold winters (Dick-Peddie 1993). Cercocarpus montanus is preferred winter range browse for deer (Roughton 1966, 1972, Hoffman and Alexander 1987). Stands can also produce significant forage that can be utilized by grazing livestock provided the slopes are not too steep (Hoffman and Alexander 1987). More information is needed on the effects of livestock grazing and wildlife browsing on the structure and function of these shrublands.
Environmental Description: Shrublands included in this alliance are found on prairie breaks and on slopes of foothills, mountains and canyons. Elevations range from 915 to 2440 m (3000-8000 feet). Climate is semi-arid. Summers are hot and winters are typically cold, with freezing temperatures and snow common. Mean annual precipitation ranges from 24-55 cm with precipitation occurring bimodally during the winter and late summer with a droughty period in late spring/early summer. The late-summer rain often occurs as high-intensity thunderstorms. Sites are variable but are generally xeric and rocky with moderate to very steep slopes or on ridges. Stands are found on various aspects, but typically the higher elevation and northern sites are on warmer, southern exposures, and lower elevation and southern stands are restricted to more mesic north slopes. Substrates are typically thin, well-drained, poorly developed, lithic soils with abundant rock outcrops. Soil textures range from sandy loam to clay. Soils range from slightly acidic to slightly alkaline depending on parent material, which includes alluvium, colluvium and residuum from igneous, metamorphic or sedimentary rocks such as granite, gneiss, limestone, quartz monzonite, rhyolite, sandstone, schist and shale.
Geographic Range: This alliance is known from the southern Rocky Mountains south to the northern Chihuahuan Desert and east to the Southern Shortgrass Prairie of Colorado, New Mexico and Texas.
Nations: US
States/Provinces: CO, NE, NM, SD, TX, WY
Plot Analysis Summary:
http://vegbank.org/natureserve/ELEMENT_GLOBAL.2.899785
Confidence Level: Low
Confidence Level Comments: No Data Available
Grank: GNR
Greasons: No Data Available
Concept Lineage: A.896, A.921.
Predecessors: No Data Available
Obsolete Names: No Data Available
Obsolete Parents: No Data Available
Synonomy: ? Cercocarpus montanus - Rhus aromatica / Bouteloua curtipendula Plant Association (Johnston 1987) [Rhus aromatica is a synonym for R. tridentata]
? Cercocarpus montanus / Bouteloua curtipendula Habitat Type (Hoffman and Alexander 1987)
? Cercocarpus montanus (Mountain Mahogany) Series (Muldavin et al. 1998b)
? Mountain Mahogany-Mixed Shrub Series (Dick-Peddie 1993) [included within the Montane Scrubland Vegetation Type]
? Mountain mahogany Series (133.34) (Brown 1982a) [included within Interior Chaparral]
? Cercocarpus montanus / Bouteloua curtipendula Habitat Type (Hoffman and Alexander 1987)
? Cercocarpus montanus (Mountain Mahogany) Series (Muldavin et al. 1998b)
? Mountain Mahogany-Mixed Shrub Series (Dick-Peddie 1993) [included within the Montane Scrubland Vegetation Type]
? Mountain mahogany Series (133.34) (Brown 1982a) [included within Interior Chaparral]
- Brotherson, J. D., D. L. Anderson, and L. A. Szyska. 1984. Habitat relations of Cercocarpus montanus (true mountain mahogany) in central Utah. Journal of Range Management 37:321-324.
- Brown, D. E., editor. 1982a. Biotic communities of the American Southwest-United States and Mexico. Desert Plants Special Issue 4(1-4):1-342.
- Cronquist, A., N. H. Holmgren, and P. K. Holmgren. 1997. Intermountain flora: Vascular plants of the Intermountain West, USA. Volume 3, Part A, subclass Rosidae (except Fabeles). New York Botanical Garden, Bronx, NY. 446 pp.
- Diamond, D. D. 1993. Classification of the plant communities of Texas (series level). Unpublished document. Texas Natural Heritage Program, Austin. 25 pp.
- Dick-Peddie, W. A. 1993. New Mexico vegetation: Past, present, and future. University of New Mexico Press, Albuquerque. 244 pp.
- Faber-Langendoen, D., J. Drake, M. Hall, G. Kittel, S. Menard, C. Nordman, M. Pyne, M. Reid, M. Russo, K. Schulz, L. Sneddon, K. Snow, and J. Teague. 2013-2019b. Screening alliances for induction into the U.S. National Vegetation Classification: Part 1 - Alliance concept review. NatureServe, Arlington, VA.
- Greenwood, L. R., and J. D. Brotherson. 1978. Ecological relationships between pinyon-juniper and true mountain mahogany stands in the Uintah Basin, Utah. Journal of Range Management 31(3):164-167.
- Hoffman, G. R., and R. R. Alexander. 1987. Forest vegetation of the Black Hills National Forest of South Dakota and Wyoming: A habitat type classification. Research Paper RM-276. USDA Forest Service, Rocky Mountain Forest and Range Experiment Station, Fort Collins, CO. 48 pp.
- Johnston, B. C. 1987. Plant associations of Region Two: Potential plant communities of Wyoming, South Dakota, Nebraska, Colorado, and Kansas. R2-ECOL-87-2. USDA Forest Service, Rocky Mountain Region. Lakewood, CO. 429 pp.
- Kettler, S. Personal communication. Colorado Natural Heritage Program, Colorado State University, Fort Collins.
- Muldavin, E., G. Harper, P. Neville, and Y. Chauvin. 1998b. The vegetation of White Sands Missile Range, New Mexico. Volume II. Vegetation map. Final report for Cooperative Agreement No. 14-16-00-91-233 White Sands Missile Range, U.S. Fish & Wildlife Service. The Nature Conservancy and the University of New Mexico.
- Roughton, R. D. 1966. Age structure of browse populations. Unpublished thesis, Colorado State University, Fort Collins. 154 pp.
- Roughton, R. D. 1972. Shrub age structures on a mule deer winter range in Colorado. Ecology 53(4):615-625.