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A3615 Abies lasiocarpa - Picea engelmannii Southern Rocky Mountain Moist Forest Alliance
Type Concept Sentence: These subalpine forests and woodlands of the Rocky Mountains with southern distributions are dominated by Abies lasiocarpa and/or Picea engelmannii.
Common (Translated Scientific) Name: Subalpine Fir - Engelmann Spruce Southern Rocky Mountain Moist Forest Alliance
Colloquial Name: Southern Rocky Mountain Subalpine Fir - Engelmann Spruce Moist Forest
Hierarchy Level: Alliance
Type Concept: These conifer forests occur predominantly in the southern Rocky Mountains where they are often the matrix forests of the upper montane and subalpine zones. These are closed-canopy forests (60-100% cover) dominated by Abies lasiocarpa and/or Picea engelmannii. Abies lasiocarpa may not be present where sites are too dry or cold. Canopy associates are variable depending on location. Common associates include Abies concolor, Picea pungens, Pinus strobiformis, Populus tremuloides, and Pseudotsuga menziesii. Forest understories are highly variable across the range of this alliance and can be dominated by grasses, dry sedges, mesic forbs or shrubs. The shrub layer is sparse to moderately dense. Important shrubs may include Acer glabrum, Lonicera utahensis, Mahonia repens, Paxistima myrsinites, Physocarpus malvaceus, Ribes spp., Rosa spp., Rubus parviflorus, Symphoricarpos spp., and Vaccinium spp. Forb cover is often sparse, but diverse with species including Arnica spp., Aquilegia spp., Caltha leptosepala, Cardamine cordifolia, Chamerion angustifolium, Erigeron eximius, Lupinus argenteus, Mertensia spp., Mitella pentandra, Orthilia secunda, Osmorhiza berteroi, Oxypolis fendleri, Packera spp., Pyrola spp., and Stellaria spp. Graminoid cover is generally low but may include Achnatherum lettermanii, Bromus carinatus, Carex geyeri, Carex rossii, Festuca idahoensis, Luzula parviflora, and Trisetum spicatum. Associations occur on cool, relatively mesic to moist sites with gentle to steep slopes on all aspects (especially northerly) and also on lower slopes of canyon drainages and riparian areas. Elevations range from 2200-3500 m.
Diagnostic Characteristics: This alliance is characterized by open to closed forest canopies dominated by Abies lasiocarpa and/or Picea engelmannii singly or in combination with associated species of more southern affinities. Canopy cover is typically 60-100%, but may be lower in some occurrences. Shrub and herbaceous composition is highly variable depending on location. This alliance represents the matrix forest vegetation of the upper montane and subalpine zones of the southern Rocky Mountains.
Rationale for Nominal Species or Physiognomic Features: No Data Available
Classification Comments: Many of the spruce - fir associations of this ecological group were formally grouped into one alliance, but split apart to reflect northern and southern distributions. There are some associations placed here, such as ~Abies lasiocarpa - Picea engelmannii / Acer glabrum Forest (CEGL000294)$$, which have very broad northern and southern distributions and may require further review to be parsed into two different associations.
Similar NVC Types: No Data Available
note: No Data Available
Physiognomy and Structure: These forests are dominated by needle-leaved evergreen trees up to 45 m in height and of high cover (60-100%). Although cold-deciduous trees are relatively rare, they can be prominent in some regional variants or seral stands. Stands may be so tightly stocked that little light reaches the forest floor and understory layers are depauperate. In stands with somewhat more open canopies, a moderately dense shrub layer may be present and dominated by ericaceous or cold-deciduous species. The herbaceous layer is dominated by perennial forbs or sod-forming graminoids, and herbaceous cover increases with increasing light availability and/or soil moisture. There is often significant cover of mosses and sometimes lichens on the forest floor and on downed woody material.
Floristics: Despite the wide distribution of these subalpine forests, their tree canopy characteristics are remarkably similar across their range of distribution. Abies lasiocarpa and/or Picea engelmannii are dominant singly or together. Both species are typically present in most stands. In some stands, Picea engelmannii may be absent altogether, or Abies lasiocarpa may only occur as seedlings and saplings. Picea engelmannii will often be prominent on more moist sites or in more mature stands. Populus tremuloides is a major seral species, and Pinus contorta occurs less frequently. Other common canopy associates may include Abies concolor, Picea pungens, and Pseudotsuga menziesii. The shrub and herbaceous layers of these forest associations are highly variable. Forest understories can be dominated by grasses or dry sedges in the more open forests, or by ericaceous shrubs or moist forbs in closed-canopy, moist sites. Shrub layers where they do occur are typically less than 1 m tall. Important to dominant shrubs include Acer glabrum, Jamesia americana, Juniperus communis, Mahonia repens, Physocarpus monogynus, Ribes spp., Shepherdia canadensis, Vaccinium cespitosum, Vaccinium myrtillus, and Vaccinium scoparium. Forb cover is often sparse, but diverse with species including Arnica spp., Aquilegia spp., Caltha leptosepala, Cardamine cordifolia, Chamerion angustifolium, Erigeron eximius, Lupinus argenteus, Mertensia spp., Mitella pentandra, Orthilia secunda, Osmorhiza berteroi, Oxypolis fendleri, Packera spp., Pyrola spp., and Stellaria spp. Graminoid cover is generally low but may include Achnatherum lettermanii (= Stipa lettermanii), Bromus carinatus, Carex geyeri, Carex rossii, Festuca idahoensis, Luzula parviflora, and Trisetum spicatum.
Dynamics: Picea engelmannii can be very long-lived, reaching 500 years of age. Abies lasiocarpa decreases in importance relative to Picea engelmannii with increasing distance from the region of Montana and Idaho where maritime air masses influence the climate. Fire is an important disturbance factor, but fire regimes have a long return interval and so are often stand-replacing. Picea engelmannii can rapidly recolonize and dominate burned sites, or can succeed to other species such as Pinus contorta or Populus tremuloides. Due to great longevity, Pseudotsuga menziesii may persist in stands of this alliance for long periods without regeneration. Old-growth characteristics in Picea engelmannii forests will include treefall and windthrow gaps in the canopy, with large downed logs, rotting woody material, tree seedling establishment on logs or on mineral soils unearthed in root balls, and snags. Picea engelmannii is susceptible to infestations by spruce beetle (Dendroctonus rufipennis) or spruce budworm (Choristoneura occidentalis), which can cause high mortality during outbreaks. In the Southwest, dwarf mistletoe Arceuthobium microcarpum is a common cause of mortality for the species.
Abies lasiocarpa forests develop on sites with limited, short growing seasons and relatively deep winter snowpacks. Tree growth is very slow in these habitats, and forests are rapidly colonized by much more rapidly growing shade-intolerant species, such as Pseudotsuga menziesii, Pinus contorta, or Populus tremuloides, following fire, clearcut logging, or windthrow disturbance. Abies lasiocarpa is among the most shade-tolerant trees in the Rocky Mountains, but seedlings compete poorly in greater than 50% full sunlight (Burns and Honkala 1990a). Fire is important in many of the more open sites, as well as those on steep slopes. Snow avalanches occur frequently at upper elevations, and can result in a mosaic of varying stand ages on sites affected by this disturbance type.
Abies lasiocarpa forests develop on sites with limited, short growing seasons and relatively deep winter snowpacks. Tree growth is very slow in these habitats, and forests are rapidly colonized by much more rapidly growing shade-intolerant species, such as Pseudotsuga menziesii, Pinus contorta, or Populus tremuloides, following fire, clearcut logging, or windthrow disturbance. Abies lasiocarpa is among the most shade-tolerant trees in the Rocky Mountains, but seedlings compete poorly in greater than 50% full sunlight (Burns and Honkala 1990a). Fire is important in many of the more open sites, as well as those on steep slopes. Snow avalanches occur frequently at upper elevations, and can result in a mosaic of varying stand ages on sites affected by this disturbance type.
Environmental Description: These forests occur in the southern Rocky Mountains where they are often the matrix forests of the upper montane and subalpine zones. They occur above the warmer and drier montane forests. Average temperatures are fairly uniform across the alliance''s range, with mean July and January temperatures of 12° and -10°C, respectively (Burns and Honkala 1990a). Snowpacks can be deep, but often melt quickly, and summers are cool. Summer frosts are characteristic, especially on sites where cold air pools. Elevations range from 2200-3500 m. Associations occur on cool, relatively mesic to moist sites with gentle to steep slopes on all aspects. In some locations where there is cold-air drainage, these forests may extend down in elevation into the montane zone, where they will occupy dry stream terraces, toeslopes, or mesic sites with cooler temperature regimes such as northern aspects. Parent materials and soils are variable across the distribution of the alliance. Parent materials include ash, tuff, lava, basalt, granitics, quartzite, dolomite, rhyolite, and other sedimentary rock types. Stands can also occur on colluvium or alluvium. Soils are typically not deep, poorly developed, and can have significant amounts of rock and gravel in the profile. Subalpine soils, such as found associated with these forests, often show evidence of podzolization processes and poorly decomposed organic layers are common.
Geographic Range: This alliance is occurs in the upper montane and subalpine zones of the central and southern Rocky Mountains, but extends west in the Utah High Plateaus, Great Basin, and Columbia Plateau and east into the Wyoming Basins.
Nations: US
States/Provinces: AZ, CO, ID, MT, NM, NV, UT, WY
Plot Analysis Summary:
http://vegbank.org/natureserve/ELEMENT_GLOBAL.2.899673
Confidence Level: Moderate
Confidence Level Comments: No Data Available
Grank: GNR
Greasons: No Data Available
Concept Lineage: A.164, A.168
Predecessors: No Data Available
Obsolete Names: No Data Available
Obsolete Parents: No Data Available
Synonomy: >< Abies lasiocarpa-Picea engelmannii Series (Johnston 1987)
>< Picea series (Pfister et al. 1977)
>< Engelmann Spruce - Subalpine Fir: 206 (Eyre 1980)
>< Western Needleleaf Forests: 15: Western Spruce-Fir Forest (Picea-Abies) (Küchler 1964)
>< Western Needleleaf Forests: 21: Southwestern Spruce-Fir Forest (Picea-Abies) (Küchler 1964)
>< Picea series (Pfister et al. 1977)
>< Engelmann Spruce - Subalpine Fir: 206 (Eyre 1980)
>< Western Needleleaf Forests: 15: Western Spruce-Fir Forest (Picea-Abies) (Küchler 1964)
>< Western Needleleaf Forests: 21: Southwestern Spruce-Fir Forest (Picea-Abies) (Küchler 1964)
- Burns, R. M., and B. H. Honkala, technical coordinators. 1990a. Silvics of North America: Volume 1. Conifers. Agriculture Handbook 654. USDA Forest Service, Washington, DC. 675 pp.
- Eyre, F. H., editor. 1980. Forest cover types of the United States and Canada. Society of American Foresters, Washington, DC. 148 pp.
- Faber-Langendoen, D., J. Drake, M. Hall, G. Kittel, S. Menard, C. Nordman, M. Pyne, M. Reid, M. Russo, K. Schulz, L. Sneddon, K. Snow, and J. Teague. 2013-2019b. Screening alliances for induction into the U.S. National Vegetation Classification: Part 1 - Alliance concept review. NatureServe, Arlington, VA.
- Johnston, B. C. 1987. Plant associations of Region Two: Potential plant communities of Wyoming, South Dakota, Nebraska, Colorado, and Kansas. R2-ECOL-87-2. USDA Forest Service, Rocky Mountain Region. Lakewood, CO. 429 pp.
- Küchler, A. W. 1964. Potential natural vegetation of the conterminous United States. American Geographic Society Special Publication 36. New York, NY. 116 pp.
- Pfister, R. D., B. L. Kovalchik, S. F. Arno, and R. C. Presby. 1977. Forest habitat types of Montana. General Technical Report INT-34. USDA Forest Service, Intermountain Forest and Range Experiment Station, Ogden, UT. 174 pp.