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CEGL000332 Abies lasiocarpa / Rubus parviflorus Forest
Type Concept Sentence: No Data Available
Common (Translated Scientific) Name: Subalpine Fir / Thimbleberry Forest
Colloquial Name: No Data Available
Hierarchy Level: Association
Type Concept: This association occurs in Colorado and New Mexico, with an extension into the White Mountains of Arizona, at between 2470 and 3201 m (8100-10,500 feet) elevation. It is located on moist, protected sites on northwest and northeast aspects at streamsides, in draws, and on middle and lower slopes with gradients that vary between 20 and 60%. This type is at the lower boundary of the spruce-fir zone on wet sites. Soils are often extremely cobbly and are classified as Cryoboralfs and Cryoborolls at the warm extreme of the cryic soil temperature regime. Parent materials vary from limestones and limy-shales to volcanics. Abies lasiocarpa is the dominant tree species, with some codominance of Picea engelmannii. Pseudotsuga menziesii and Populus tremuloides are major seral species. Both tall- and low-shrub layers are present. Rubus parviflorus and Acer glabrum are the most common species. The herbaceous cover is rich in species diversity, suggestive of mesic conditions. Bromus ciliatus, Geranium richardsonii, Goodyera oblongifolia, and Orthilia secunda are commonly found.
Diagnostic Characteristics: Abies lasiocarpa dominates the overstory; Picea engelmannii is usually subordinate. Pseudotsuga menziesii and Populus tremuloides are major seral trees. Undergrowth is rich in species diversity. Except for the Vaccinium myrtillus phase, Vaccinium spp. are lacking; Rubus parviflorus is an indicator species for the undergrowth.
Rationale for Nominal Species or Physiognomic Features: No Data Available
Classification Comments: Two phases are recognized (Moir and Ludwig 1979, Stuever and Hayden 1997a). In the Vaccinium myrtillus phase, the low shrub Vaccinium myrtillus is codominant with Rubus parviflorus. The Acer glabrum phase is characterized by Acer glabrum in the tall-shrub stratum, and the absence of Vaccinium myrtillus in the low-shrub stratum. In the Vaccinium myrtillus phase, Abies concolor is minor or accidental, whereas in the Acer glabrum phase, it is a major seral species.
Similar NVC Types: No Data Available
note: No Data Available
Physiognomy and Structure: No Data Available
Floristics: Abies lasiocarpa is the dominant tree species, with some codominance of Picea engelmannii. Pseudotsuga menziesii and Populus tremuloides are major seral species. Both tall- and low-shrub layers are present. Rubus parviflorus and Acer glabrum are the most common species. Other frequently occurring shrubs are Holodiscus dumosus, Lonicera involucrata, Lonicera utahensis, Ribes pinetorum, Paxistima myrsinites, and Robinia neomexicana. The herbaceous cover is rich in species diversity, suggestive of mesic conditions. Bromus ciliatus, Geranium richardsonii, Goodyera oblongifolia, and Orthilia secunda (= Ramischia secunda) are commonly found. Also present are Arnica cordifolia, Artemisia franserioides, Clematis columbiana (= Clematis pseudoalpina), Prosartes trachycarpa (= Disporum trachycarpum), Chamerion angustifolium (= Epilobium angustifolium), Erigeron eximius (= Erigeron superbus), Oreochrysum parryi (= Haplopappus parryi), Lathyrus spp., Ligusticum porteri, Osmorhiza depauperata, Pteridium aquilinum, Packera cardamine (= Senecio cardamine), Maianthemum racemosum (= Smilacina racemosa), Maianthemum stellatum (= Smilacina stellata), Thalictrum fendleri, Viola canadensis, and Zigadenus elegans.
Moir and Ludwig (1979), who originally described this type, recognize two phases. The first is a Vaccinium myrtillus phase, characterized by Vaccinium being codominant with Rubus parviflorus. This phase is similar to the Abies lasiocarpa / Vaccinium myrtillus, Rubus parviflorus phase habitat type, and occurs on cooler sites with deeper snowpack. The second phase is an Acer glabrum phase. Acer is consistent in the tall-shrub stratum, and Vaccinium myrtillus is absent in the low-shrub stratum with Abies concolor as a major seral species, whereas Abies concolor is only accidental or minor in the Vaccinium myrtillus phase.
Moir and Ludwig (1979), who originally described this type, recognize two phases. The first is a Vaccinium myrtillus phase, characterized by Vaccinium being codominant with Rubus parviflorus. This phase is similar to the Abies lasiocarpa / Vaccinium myrtillus, Rubus parviflorus phase habitat type, and occurs on cooler sites with deeper snowpack. The second phase is an Acer glabrum phase. Acer is consistent in the tall-shrub stratum, and Vaccinium myrtillus is absent in the low-shrub stratum with Abies concolor as a major seral species, whereas Abies concolor is only accidental or minor in the Vaccinium myrtillus phase.
Dynamics: Abies lasiocarpa is almost uniformly susceptible to Fomes annosus and Melampsorella caryophyllacearum (yellow witches'' broom rust). Picea is susceptible to Engelmann spruce bark beetle (Dendroctonus rufipennis), which may limit or temporarily suppress its codominance with the Abies according to Jones (1974).
Crane (1982) places this association in fire group fourteen, which are the mesic, low to mid-elevation subalpine fir habitat types in southwestern Colorado. There does not appear to be any specific information on fire frequency for forests in this fire group. Fire disturbance is relatively light, burning irregularly in patchy distribution on this moist type according to Moir and Ludwig (1979).
Following a stand-replacing fire, the community reverts to a grass and forb stage, followed shortly by aspen resprouting rapidly. The grass stage will be prolonged by repeated fires. The most likely conifers in an early stand are Douglas-fir and Engelmann spruce, although fir may be present. As the aspen grows into mature trees, the slower-growing conifers will remain understory trees. A low surface fire will kill some of the aspen but will not stimulate much sprouting. If the Douglas-fir has developed thickened bark, it should easily withstand a low surface fire. However, many of the spruce and most of the firs may be killed. The main effect of a low surface fire is to open the stand for seedling establishment and to lessen the importance of aspen. A severe fire would return the site to grass and forbs. If no fires occur, a mixed stand will develop on the site. Eventually, Douglas-fir could be lost from the site because of shading, resulting in a climax of fir and possibly spruce. This climax forest will maintain itself and low surface fires should only remove duff and allow regeneration.
Crane (1982) places this association in fire group fourteen, which are the mesic, low to mid-elevation subalpine fir habitat types in southwestern Colorado. There does not appear to be any specific information on fire frequency for forests in this fire group. Fire disturbance is relatively light, burning irregularly in patchy distribution on this moist type according to Moir and Ludwig (1979).
Following a stand-replacing fire, the community reverts to a grass and forb stage, followed shortly by aspen resprouting rapidly. The grass stage will be prolonged by repeated fires. The most likely conifers in an early stand are Douglas-fir and Engelmann spruce, although fir may be present. As the aspen grows into mature trees, the slower-growing conifers will remain understory trees. A low surface fire will kill some of the aspen but will not stimulate much sprouting. If the Douglas-fir has developed thickened bark, it should easily withstand a low surface fire. However, many of the spruce and most of the firs may be killed. The main effect of a low surface fire is to open the stand for seedling establishment and to lessen the importance of aspen. A severe fire would return the site to grass and forbs. If no fires occur, a mixed stand will develop on the site. Eventually, Douglas-fir could be lost from the site because of shading, resulting in a climax of fir and possibly spruce. This climax forest will maintain itself and low surface fires should only remove duff and allow regeneration.
Environmental Description: This association occurs at between 2470 and 3201 m (8100-10,500 feet) elevation. It is located on moist, protected sites on northwest and northeast aspects. It is found at streamsides, in draws, and on middle and lower slopes with gradients that vary between 20 and 60% (Fitzhugh et al. 1987). This type is at the lower boundary of the spruce-fir zone on wet sites (Moir and Ludwig 1979). Soils are often extremely cobbly and are classified as Cryoboralfs and Cryoborolls at the warm extreme of the cryic soil-temperature regime. Parent materials vary from limestones and limey-shales to volcanics.
Geographic Range: This association occurs in Colorado and New Mexico, with an extension into the White Mountains of Arizona.
Nations: US
States/Provinces: AZ, CO, NM
Plot Analysis Summary:
http://vegbank.org/natureserve/ELEMENT_GLOBAL.2.689915
Confidence Level: Moderate
Confidence Level Comments: No Data Available
Grank: G5
Greasons: No Data Available
Type | Name | Database Code | Classification Code |
---|---|---|---|
Class | 1 Forest & Woodland Class | C01 | 1 |
Subclass | 1.B Temperate & Boreal Forest & Woodland Subclass | S15 | 1.B |
Formation | 1.B.2 Cool Temperate Forest & Woodland Formation | F008 | 1.B.2 |
Division | 1.B.2.Nb Rocky Mountain Forest & Woodland Division | D194 | 1.B.2.Nb |
Macrogroup | 1.B.2.Nb.5 Subalpine Fir - Engelmann Spruce - Whitebark Pine Rocky Mountain Forest Macrogroup | M020 | 1.B.2.Nb.5 |
Group | 1.B.2.Nb.5.c Engelmann Spruce - Subalpine Fir - Mountain Hemlock Moist Forest & Woodland Group | G218 | 1.B.2.Nb.5.c |
Alliance | A3615 Subalpine Fir - Engelmann Spruce Southern Rocky Mountain Moist Forest Alliance | A3615 | 1.B.2.Nb.5.c |
Association | CEGL000332 Subalpine Fir / Thimbleberry Forest | CEGL000332 | 1.B.2.Nb.5.c |
Concept Lineage: No Data Available
Predecessors: No Data Available
Obsolete Names: No Data Available
Obsolete Parents: No Data Available
Synonomy: = Abies bifolia / Rubus parviflorus (Stuever and Hayden 1997a)
= Abies lasiocarpa - Picea engelmannii / Rubus parviflorus Plant Association (Johnston 1987) [in Colorado in the White River National Forest from 2530-2865 m (8300-9400 feet), the San Juan National Forest from 2680-3200 m (8800-10,500 feet), the Rio Grande National Forest from 2590-3140 m (8500-10,300 feet) and New Mexico.]
= Abies lasiocarpa / Rubus parviflorus Habitat Type (Alexander et al. 1987)
= Abies lasiocarpa / Rubus parviflorus Habitat Type (Muldavin et al. 1996)
= Abies lasiocarpa / Rubus parviflorus Habitat Type (Muldavin et al. 1986) [in the White Mountains of Arizona at 2810 m (9220 feet).]
= Abies lasiocarpa / Rubus parviflorus Habitat Type (DeVelice et al. 1986) [found in the San Juan Mountains of Colorado from 2710-2990 m (8900-9800 feet).]
= Abies lasiocarpa / Rubus parviflorus Habitat Type (Larson and Moir 1987) [found in New Mexico.]
= Abies lasiocarpa / Rubus parviflorus Habitat Type (Moir and Ludwig 1979) [located in the Mogollon Mountains of New Mexico from 2590-3140 m (8500-10,300 feet).]
? Abies lasiocarpa / Rubus parviflorus Habitat Type (Fitzhugh et al. 1987) [found in the Mogollon Mountains, Gila National Forest, Glenwood and Wilderness Ranger Districts of New Mexico from 2480 and 2940 m (8140-9640 feet).]
= Abies lasiocarpa - Picea engelmannii / Rubus parviflorus Plant Association (Johnston 1987) [in Colorado in the White River National Forest from 2530-2865 m (8300-9400 feet), the San Juan National Forest from 2680-3200 m (8800-10,500 feet), the Rio Grande National Forest from 2590-3140 m (8500-10,300 feet) and New Mexico.]
= Abies lasiocarpa / Rubus parviflorus Habitat Type (Alexander et al. 1987)
= Abies lasiocarpa / Rubus parviflorus Habitat Type (Muldavin et al. 1996)
= Abies lasiocarpa / Rubus parviflorus Habitat Type (Muldavin et al. 1986) [in the White Mountains of Arizona at 2810 m (9220 feet).]
= Abies lasiocarpa / Rubus parviflorus Habitat Type (DeVelice et al. 1986) [found in the San Juan Mountains of Colorado from 2710-2990 m (8900-9800 feet).]
= Abies lasiocarpa / Rubus parviflorus Habitat Type (Larson and Moir 1987) [found in New Mexico.]
= Abies lasiocarpa / Rubus parviflorus Habitat Type (Moir and Ludwig 1979) [located in the Mogollon Mountains of New Mexico from 2590-3140 m (8500-10,300 feet).]
? Abies lasiocarpa / Rubus parviflorus Habitat Type (Fitzhugh et al. 1987) [found in the Mogollon Mountains, Gila National Forest, Glenwood and Wilderness Ranger Districts of New Mexico from 2480 and 2940 m (8140-9640 feet).]
- Alexander, B. G., Jr., E. L. Fitzhugh, F. Ronco, Jr., and J. A. Ludwig. 1987. A classification of forest habitat types of the northern portion of the Cibola National Forest, NM. General Technical Report RM-143. USDA Forest Service, Rocky Mountain Forest and Range Experiment Station, Fort Collins, CO. 35 pp.
- Bourgeron, P. S., and L. D. Engelking, editors. 1994. A preliminary vegetation classification of the western United States. Unpublished report. The Nature Conservancy, Western Heritage Task Force, Boulder, CO. 175 pp. plus appendix.
- CNHP [Colorado Natural Heritage Program]. 2006-2017. Tracked natural plant communities. Colorado Natural Heritage Program, Colorado State University, Fort Collins. [https://cnhp.colostate.edu/ourdata/trackinglist/plant_communities/]
- Crane, M. F. 1982. Fire ecology of Rocky Mountain Region forest habitat types. USDA Forest Service final report. 272 pp.
- DeVelice, R. L., J. A. Ludwig, W. H. Moir, and F. Ronco, Jr. 1986. A classification of forest habitat types of northern New Mexico and southern Colorado. General Technical Report RM-131. USDA Forest Service, Rocky Mountain Forest and Range Experiment Station, Fort Collins, CO. 59 pp.
- Fitzhugh, E. L., W. H. Moir, J. A. Ludwig, and F. Ronco, Jr. 1987. Forest habitat types in the Apache, Gila, and part of the Cibola national forests. General Technical Report RM-145. USDA Forest Service, Rocky Mountain Forest and Range Experiment Station, Fort Collins, CO. 116 pp.
- Johnston, B. C. 1987. Plant associations of Region Two: Potential plant communities of Wyoming, South Dakota, Nebraska, Colorado, and Kansas. R2-ECOL-87-2. USDA Forest Service, Rocky Mountain Region. Lakewood, CO. 429 pp.
- Jones, J. R. 1974. Silviculture of southwestern mixed conifer and aspen: The status of our knowledge. Research Paper RM-122. USDA Forest Service, Rocky Mountain Forest and Range Experiment Station, Fort Collins, CO. 44 pp.
- Larson, M., and W. H. Moir. 1987. Forest and woodland habitat types of northern New Mexico and northern Arizona. Edition 2. USDA Forest Service, Southwestern Region, Albuquerque, NM.
- Moir, W. H., and J. A. Ludwig. 1979. A classification of spruce-fir and mixed conifer habitat types of Arizona and New Mexico. Research Paper RM-207. USDA Forest Service, Rocky Mountain Forest and Range Experiment Station, Fort Collins, CO. 47 pp.
- Muldavin, E. H., R. L. DeVelice, and F. Ronco, Jr. 1996. A classification of forest habitat types of southern Arizona and portions of the Colorado Plateau. General Technical Report RM-GTR-287. USDA Forest Service, Rocky Mountain Forest and Range Experiment Station, Fort Collins, CO. 130 pp.
- Muldavin, E., R. L. DeVelice, and W. A. Dick-Peddie. 1986. Forest habitat types of the Fort Apache, San Carlos and Hualapai Indian reservations, Arizona. Final Report Cooperative Agreement 28-K3-208 Addendum, Rocky Mountain Forest and Range Experiment Station and New Mexico State University. 67 pp.
- Stuever, M. C., and J. S. Hayden. 1997a. Plant associations of Arizona and New Mexico, edition 3. Volume 1: Forests. USDA Forest Service, Southwestern Region. Habitat Typing Guides. 291 pp.
- Tirmenstein, D. 1990b. Vaccinium myrtillus. In: Fire Effects Information System [Online]. USDA Forest Service, Rocky Mountain Research Station, Fire Sciences Laboratory (Producer). [http://www.fs.fed.us/database/feis/]
- USFS [U.S. Forest Service]. 1987a. Forest and woodland habitat types (plant associations) of northern New Mexico and northern Arizona. USDA Forest Service, Southwestern Region, Albuquerque, NM. Second edition, 170 pp. plus insert.
- USFS [U.S. Forest Service]. 1987b. Forest and woodland habitat types (plant associations) of Arizona south of the Mogollon Rim and southwestern New Mexico. USDA Forest Service, Southwestern Region, Albuquerque, NM. Second edition, 168 pp. plus insert.
- Uchytil, R. J. 1991e. Abies lasiocarpa. In: Fire Effects Information System [Online]. USDA Forest Service, Rocky Mountain Research Station, Fire Sciences Laboratory (Producer). [http://www.fs.fed.us/database/feis/] (accessed May 26, 2015).
- Western Ecology Working Group of NatureServe. No date. International Ecological Classification Standard: International Vegetation Classification. Terrestrial Vegetation. NatureServe, Boulder, CO.