Print Report
A3577 Pinus edulis - Juniperus monosperma Grassy Woodland Alliance
Type Concept Sentence: This pinyon-juniper woodland and savanna alliance is characterized diagnostic tree species Pinus edulis (>5% cover) that forms a very open to moderately dense tree layer often with Juniperus monosperma present to codominant (or Juniperus scopulorum at higher elevations) with an understory dominated by an open to dense layer of perennial grasses and lacking significant cover of shrubs (<10%). It occurs on warm, dry mountains and foothills in southern Colorado east of the Continental Divide and in mountains and plateaus of northern and central New Mexico, extending east on breaks in the southeastern Great Plains.
Common (Translated Scientific) Name: Two-needle Pinyon - One-seed Juniper Grassy Woodland Alliance
Colloquial Name: Southern Rockies Pinyon - Juniper Grassy Woodland
Hierarchy Level: Alliance
Type Concept: This woodland and savanna alliance is characterized by diagnostic tree species Pinus edulis that forms a very open to moderately dense tree layer often with Juniperus monosperma. Juniperus monosperma may also dominate stands as long as there is significant presence of Pinus edulis (not accidental) to characterize the stand as a pinyon-juniper stand and not the more xeric, typically lower elevation Juniperus osteosperma woodland and savanna. At higher elevations and relatively mesic sites, such as along drainages, Juniperus scopulorum may be present and sometimes dominant. In southern transitional areas with ~Madrean Pinyon - Juniper Woodland Group (G200)$$ in central New Mexico, Juniperus deppeana may be present but not dominant. Other conifers are absent or accidental with very low cover. Scattered shrubs may be present but do not form a layer and do not exceed cover of herbaceous layer. The understory is characterized by an open to dense herbaceous layer composed of grasses often with diverse but low cover of forbs. Characteristic species may include Achnatherum nelsonii, Achnatherum scribneri, Andropogon hallii, Bouteloua curtipendula, Bouteloua gracilis, Festuca arizonica, Hesperostipa comata, Hesperostipa neomexicana, Leymus ambiguus, Muhlenbergia montana, and Pleuraphis jamesii. This alliance occurs on warm, dry mountains and foothills in southern Colorado east of the Continental Divide and in mountains and plateaus of northern and central New Mexico, extending east on breaks in the southeastern Great Plains. Stands typically occur on nearly level to steep (to 80%), rocky slopes on hillsides and ridgetops on all aspects except in elevational extremes for a given latitude where low-elevation stands are restricted to the more mesic north slopes; canyons and high-elevation stands occur on south aspects. Sites are typically dry with shallow, rocky, calcareous and alkaline soils. Other sites include eroded "badlands," lava flows, scree slopes, and deep sands.
Diagnostic Characteristics: This alliance is characterized by an open to moderate tree canopy, typically with between 5 and 30% cover. The diagnostic and often dominant species is Pinus edulis. Juniperus monosperma or Juniperus scopulorum is often present to codominant and may dominate stands as long as there is significant presence of Pinus edulis (not accidental) to characterize the stand as a pinyon-juniper stand. The understory is characterized by an open to dense herbaceous layer. Characteristic species may include Achnatherum nelsonii, Achnatherum scribneri, Andropogon hallii, Bouteloua curtipendula, Bouteloua gracilis, Festuca arizonica, Hesperostipa comata, Hesperostipa neomexicana, Leymus ambiguus, Muhlenbergia montana, and Pleuraphis jamesii.
Rationale for Nominal Species or Physiognomic Features: No Data Available
Classification Comments: Several of the associations in this alliance need further review to clarify whether to add Juniperus monosperma to the name and if there is a Colorado Plateau analog codominated by Juniperus osteosperma. ~Pinus edulis / Bouteloua curtipendula Open Woodland (CEGL000777)$$ and ~Pinus edulis - (Juniperus monosperma) / Festuca arizonica Open Woodland (CEGL000783)$$ are broadly defined as being codominated by Juniperus monosperma or Juniperus osteosperma. They may need to be split into new associations to include stands inside the distributional range of Juniperus osteosperma and placed in ~Colorado Plateau Pinyon - Juniper Woodland Group (G900)$$. Colorado stands have 5-25% tree cover. Stands with <5% cover may better be placed in similar grass associations (e.g., CEGL001703). On New Mexico sites, this type typically has 10-30% tree cover with some communities having over 30% tree cover.
Similar NVC Types: No Data Available
note: No Data Available
Physiognomy and Structure: Vegetation included in this alliance has a moderately sparse to moderately dense tree canopy (5-50% cover) that is typically 3-10 m tall. Stands are either solely dominated by evergreen needle-leaved trees or may be codominated by broad-leaved or scale-leaved evergreen trees. An open to moderate ground layer dominated by perennial graminoids is present. Perennial forbs and cacti are often scattered throughout the stands. Annual forbs and grasses may be seasonally present. Scattered broad-leaved and microphyllous deciduous or evergreen shrubs may be present but do not form a layer (<10% cover).
Floristics: This woodland and savanna alliance is characterized by diagnostic tree species Pinus edulis that forms a very open to moderately dense, short (<20 m tall) tree layer often with Juniperus monosperma. Juniperus monosperma may also dominate stands as long as there is significant (>5% cover) presence of Pinus edulis (not accidental) to characterize the stand as a pinyon-juniper stand and not the more xeric, typically lower elevation Juniperus osteosperma woodland and savanna. At higher elevations and relatively mesic sites, such as along drainages, Juniperus scopulorum may be present and sometimes dominant. In southern transitional areas with ~Madrean Pinyon - Juniper Woodland Group (G200)$$ in central New Mexico, Juniperus deppeana may be present but not dominant. Other conifers are absent or accidental with very low cover. Scattered shrubs such as Artemisia bigelovii, Artemisia tridentata, Cercocarpus montanus, Gutierrezia sarothrae, Krascheninnikovia lanata, Opuntia spp., and Rhus trilobata may be present (<10% total cover) but do not form a layer and do not exceed cover of herbaceous layer. The understory is characterized by an open to dense herbaceous layer composed of perennial grasses often with diverse but low cover of forbs. Characteristic species may include Achnatherum hymenoides, Achnatherum nelsonii, Achnatherum scribneri, Aristida spp., Andropogon hallii, Bouteloua curtipendula, Bouteloua gracilis, Festuca arizonica, Koeleria macrantha, Hesperostipa comata, Hesperostipa neomexicana, Leymus ambiguus, Muhlenbergia montana, Piptatheropsis micrantha (= Oryzopsis micrantha), Pleuraphis jamesii, and Sporobolus spp. Many forb species occur, but few have much cover. Commonly present forbs include species of Artemisia, Eriogonum, Heterotheca, Mirabilis, Penstemon, Phlox, Senecio, and Zinnia. Annual grasses and forbs are seasonally present. Adjacent vegetation at higher elevations is typically woodland or forest dominated by Pinus ponderosa. Adjacent vegetation at lower elevations is often Juniperus spp.-dominated woodland and savanna, Artemisia spp.-dominated shrubland, or grassland.
Dynamics: Pinus edulis is extremely drought-tolerant and slow-growing (Little 1987, Powell 1988b, Muldavin et al. 1998c). It is also non-sprouting and may be killed by fire (Wright et al. 1979). The effect of fire on a stand is largely dependent on the tree height and density, fine-fuel load on the ground, weather conditions, and season (Dwyer and Pieper 1967, Wright et al. 1979). Trees are more vulnerable in open stands where fires frequently occur in the spring, when the relative humidity is low, wind speeds are over 10-20 mph, and there are adequate fine fuels to carry fire (Wright et al. 1979). Under other conditions, burns tend to be spotty with low tree mortality. Large trees are generally not killed unless fine fuels, such as tumbleweeds, have accumulated beneath the tree to provide ladder fuels for the fire to reach the crown (Jameson 1962). Closed-canopy stands rarely burn because they typically do not have enough understory or wind to carry a fire (Wright et al. 1979).
Although Pinus edulis is drought-tolerant, prolonged droughts will weaken trees and promote mortality by secondary agents. Periodic die-offs of pinyon pine caused by insects, such as the pinyon Ips beetle (Ips confusus), or fungal agents, such as blackstain root-rot (Leptographium wageneri), tend to be correlated with droughts (Anhold 2005). These mortality events may be localized or widespread but can result in 50 to 90% mortality of Pinus edulis (Harrington and Cobb 1988).
Climatic and other factors have resulted in denser and expanded pinyon-juniper stands. Denser stands are more susceptible to attack by insects and disease (Anhold 2005). In addition, altered fire regimes, cutting trees for fencing or firewood, and improper grazing by livestock have significant impacts on the quality of sites. Grazing by livestock can modify the fire regime by removing the fine fuels that carry fire. Fire, livestock grazing, and trampling by recreationalists and vehicles disturb cryptogamic soil crusts that help maintain soil structure, reduce soil erosion, provide habitat for plants and preserve biological diversity (Ladyman and Muldavin 1996). More study is needed to understand and manage these woodlands ecologically.
Although Pinus edulis is drought-tolerant, prolonged droughts will weaken trees and promote mortality by secondary agents. Periodic die-offs of pinyon pine caused by insects, such as the pinyon Ips beetle (Ips confusus), or fungal agents, such as blackstain root-rot (Leptographium wageneri), tend to be correlated with droughts (Anhold 2005). These mortality events may be localized or widespread but can result in 50 to 90% mortality of Pinus edulis (Harrington and Cobb 1988).
Climatic and other factors have resulted in denser and expanded pinyon-juniper stands. Denser stands are more susceptible to attack by insects and disease (Anhold 2005). In addition, altered fire regimes, cutting trees for fencing or firewood, and improper grazing by livestock have significant impacts on the quality of sites. Grazing by livestock can modify the fire regime by removing the fine fuels that carry fire. Fire, livestock grazing, and trampling by recreationalists and vehicles disturb cryptogamic soil crusts that help maintain soil structure, reduce soil erosion, provide habitat for plants and preserve biological diversity (Ladyman and Muldavin 1996). More study is needed to understand and manage these woodlands ecologically.
Environmental Description: This woodland and savanna alliance occurs on warm, dry mountains and foothills in southern Colorado east of the Continental Divide and in mountains and plateaus of northern and central New Mexico, extending east on breaks in the southeastern Great Plains. Climate is semi-arid and droughts are not uncommon. Summers are generally hot, and winters range from mild with cold periods and occasional snows in southern New Mexico and Arizona to the more typical extended periods of freezing temperatures. The seasonality of precipitation varies from east to west with summer rain more common in the southern and eastern portion of the alliance''s range and winter precipitation more common in the western portion of the range. Mean annual precipitation ranges from 30-46 cm. Stands occur on escarpments and canyons, mesas, and piedmonts to the lower montane zone. Elevation ranges from 1406-2510 m. Sites are nearly level to moderately steep, rocky slopes on hillsides and ridgetops. Aspect does not seem important except in elevational extremes for a given latitude where low-elevation stands are restricted to the more mesic north slopes; canyons and high-elevation stands occur on south aspects. Sites are typically dry with shallow, rocky, calcareous and alkaline soils. Other sites include eroded "badlands," lava flows, scree slopes, and deep sands. Soil textures range from sandy loam to clay and are typically derived from limestone, sandstone or shale. Other parent materials include andesite, basalt, granite, quartzite, monzonite, rhyolite and mixed alluvium.
Geographic Range: This woodland and savanna alliance occurs on warm, dry mountains and foothills in southern Colorado east of the Continental Divide and in mountains and plateaus of northern and central New Mexico (USFS 2007), extending east on breaks in the southeastern Great Plains.
Nations: US
States/Provinces: AZ, CO, NM, OK, TX?
Plot Analysis Summary:
http://vegbank.org/natureserve/ELEMENT_GLOBAL.2.899635
Confidence Level: Low
Confidence Level Comments: No Data Available
Grank: GNR
Greasons: No Data Available
Concept Lineage: This alliance is composed of eight herbaceous understory associations from Old Alliance II.A.4.N.a. Pinus edulis - (Juniperus spp.) Woodland Alliance (A.516).
Predecessors: No Data Available
Obsolete Names: No Data Available
Obsolete Parents: No Data Available
Synonomy: < Colorado Pinyon-Alligator Juniper Series (Dick-Peddie 1993)
< Colorado Pinyon-Mixed Juniper Series (Dick-Peddie 1993)
< Colorado Pinyon-One Seeded Juniper Series (Dick-Peddie 1993)
< Colorado Pinyon-Rocky Mountain Juniper Series (Dick-Peddie 1993)
< Colorado Pinyon-Mixed Juniper Series (Dick-Peddie 1993)
< Colorado Pinyon-One Seeded Juniper Series (Dick-Peddie 1993)
< Colorado Pinyon-Rocky Mountain Juniper Series (Dick-Peddie 1993)
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