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A3573 Pinus edulis - Juniperus osteosperma Colorado Plateau Shrubby Open Woodland Alliance

Type Concept Sentence: This open woodland and scrub alliance is characteristic of exposed rocky mesatops and canyon slopes and rims in the Colorado Plateau and is characterized by diagnostic tree species Pinus edulis that forms a very open to moderately dense, short tree layer often with Juniperus osteosperma and an understory lacking or dominated by an open to moderately dense layer of shrubs (>10% cover) or, if less, then exceeding cover of grasses.


Common (Translated Scientific) Name: Two-needle Pinyon - Utah Juniper Colorado Plateau Shrubby Open Woodland Alliance

Colloquial Name: Colorado Plateau Two-needle Pinyon - Utah Juniper Open Woodland

Hierarchy Level:  Alliance

Type Concept: This open woodland and scrub alliance is characteristic of exposed rocky mesatops and canyon slopes and rims on the Colorado Plateau, but extends upslope into foothills and on xeric sites. The vegetation is sparse to moderately dense (10-80% total vegetation cover) and is dominated by dwarfed (usually <3 m tall) Pinus edulis and/or Juniperus osteosperma trees that form extensive tall shrublands or scrublands in the canyon country of the Colorado Plateau region. Both tree species are present with 3-25% cover. The understory is lacking or characterized by an open to moderately dense layer of shrubs (>10% cover) or, if less, then exceeding cover of grasses. Shrubs, if present, often include Artemisia bigelovii, Atriplex canescens, Cercocarpus intricatus, Cercocarpus montanus, Chrysothamnus viscidiflorus, Coleogyne ramosissima, Ephedra torreyana, Ephedra viridis, Fendlera rupicola, Glossopetalon spinescens var. aridum, Petradoria pumila, Purshia stansburiana, Quercus havardii var. tuckeri, or Quercus turbinella. Herbaceous layers are typically sparse and composed of xeric grasses and forbs. Opuntia fragilis or cushion plants such as Arenaria eastwoodiae and Paronychia sessiliflora may characterize the understory and exceed shrub cover. Sites are variable but generally xeric. Stands occur on dry canyon rims, ridges, hills, benches, hogbacks, mesas, elevated plains and occasionally in intermittent drainages. Elevations generally range from 1400 to 2165 m (4600-7100 feet), but may extend up to 2480 m (8135 feet). Stands typically occur on gentle to moderately steep slopes on all aspects, but range from flat to steep slopes (0-30%). The soils are variable but generally shallow, poorly developed and skeletal, ranging from well-drained clay loam or sandy clay to rapidly drain loamy sand. The unvegetated ground surface is typically composed of litter, bedrock, and bare soil. Because this is a relatively xeric woodland, bare soil and rocks may cover up to 70% of the unvegetated surface, although biological soil crusts provide up to 40% cover.

Diagnostic Characteristics: This is the common pinyon-juniper woodland and scrub found throughout much of the Colorado Plateau. The tree canopy is characterized by a sparse to open tree canopy that is relatively short (usually <3 m tall) but may extend up to 10 m. The diagnostic tree species are Pinus edulis and Juniperus osteosperma, either of which may dominate woodland stands as long as there is significant presence of Pinus edulis (not accidental) to characterize as a pinyon-juniper type. The sparse to moderately dense understory is an open shrub layer. The herbaceous layer is typically sparse and composed of grasses and often diverse forbs, especially cushion plants.

Rationale for Nominal Species or Physiognomic Features: No Data Available

Classification Comments: This alliance is similar to ~Pinus edulis - Juniperus osteosperma / Shrub Understory Foothill & Lower Montane Dry-Mesic Woodland Alliance (A3571)$$, but trees are sometime stunted by harsh, xeric environments and have a sparse to open woody canopy, and lack some of the upper foothill, lower montane floristic species such as Quercus gambelii, when compared to higher elevation foothill pinyon-juniper woodland stands. Additional review is needed to verify association placement in these alliances. Scattered shrubs frequently characterize sparse and rocky understory stands and so are included in this alliance.

Similar NVC Types: No Data Available
note: No Data Available

Physiognomy and Structure: Vegetation included in this alliance has a sparse to moderately dense tree canopy that is typically <3 m tall, but may extent to 10 m on favorable sites. Stands are either solely dominated by evergreen needle-leaved trees or may be codominated by scale-leaved evergreen trees. A sparse to moderately dense shrub layer (0.5-3 m tall) may be present. If present, the shrub layer ranges from a single species to a diverse mix of broad-leaved and microphyllous deciduous or evergreen shrubs that are usually less than 3 m tall. A sparse to moderate ground layer dominated by perennial graminoids is usually present. Perennial forbs and cacti are often scattered throughout the stands. Annual forbs and grasses may be seasonally present.

Floristics: This vegetation of this open woodland and scrub alliance is sparse to moderately dense (10-80% total vegetation cover) and is dominated by dwarfed (usually <3 m tall) Pinus edulis and/or Juniperus osteosperma trees that form extensive tall shrublands or scrublands in the canyon country of the Colorado Plateau region. Both tree species are present with 3-25% cover. The understory may be lacking or be characterized by an open to moderately dense layer of shrubs (>10% cover) or, if less, then exceeding cover of grasses. Shrubs, if present, often include Artemisia bigelovii, Atriplex canescens, Cercocarpus intricatus, Cercocarpus montanus, Chrysothamnus viscidiflorus, Coleogyne ramosissima, Ephedra torreyana, Ephedra viridis, Fendlera rupicola, Glossopetalon spinescens var. aridum (= Glossopetalon nevadense), Petradoria pumila, Purshia stansburiana, Quercus havardii var. tuckeri, or Quercus turbinella. Shrub mixes often occur on steeper talus slopes. Herbaceous layers are typically sparse and composed of xeric grasses and diverse forbs, especially cushion plants. Opuntia fragilis or cushion plants, such as Arenaria eastwoodiae, Enceliopsis nudicaulis, Erigeron compactus, Erigeron pumilus, Eriogonum alatum, Frasera albomarginata, Heterotheca villosa, Hymenopappus filifolius, Paronychia sessiliflora, Polygala subspinosa, Stenotus armerioides, Tetraneuris acaulis, Tetraneuris torreyana (= Tetraneuris depressa), Townsendia aprica, and Townsendia incana, may characterize the understory and exceed shrub cover. Common perennial grasses include Achnatherum hymenoides, Aristida purpurea, Bouteloua gracilis, Pleuraphis jamesii, and Poa fendleriana. Other common forbs, such as Arenaria fendleri, Artemisia ludoviciana, Descurainia pinnata, Eriogonum ovalifolium, Heterotheca villosa, Lappula occidentalis, Machaeranthera grindelioides, Oenothera pallida, Penstemon linarioides, and Streptanthus cordatus, may be present with low cover. Stands of this community sampled at higher elevation have northern aspects and more dense vegetation cover. Biological soil crusts may have significant cover in sites derived from loess.

Dynamics:  Pinus edulis is extremely drought-tolerant and slow-growing (Little 1987, Powell 1988b, Muldavin et al. 1998c). It is also non-sprouting and may be killed by fire (Wright et al. 1979). Many xeric stands in this alliance are too sparse to burn. However, the effect of fire on a stand is largely dependent on the tree height and density, fine-fuel load on the ground, weather conditions, and season (Dwyer and Pieper 1967, Wright et al. 1979). Trees are more vulnerable in open stands where fires frequently occur in the spring, when the relative humidity is low, wind speeds are over 10-20 mph, and there are adequate fine fuels to carry fire (Wright et al. 1979). Under other conditions, burns tend to be spotty with low tree mortality. Large trees are generally not killed unless fine fuels, such as tumbleweeds, have accumulated beneath the tree to provide ladder fuels for the fire to reach the crown (Jameson 1962). Closed-canopy stands rarely burn because they typically do not have enough understory or wind to carry a fire (Wright et al. 1979).

Although Pinus edulis is drought-tolerant, prolonged droughts will weaken trees and promote mortality by secondary agents. Periodic die-offs of pinyon pine caused by insects, such as the pinyon Ips beetle (Ips confusus), or fungal agents, such as blackstain root-rot (Leptographium wageneri), tend to be correlated with droughts (Anhold 2005). These mortality events may be localized or widespread but can result in 50 to 90% mortality of Pinus edulis (Harrington and Cobb 1988).

Climatic and other factors have resulted in denser and expanded pinyon-juniper stands throughout the Colorado Plateau and Great Basin. Denser stands are more susceptible to attack by insects and disease (Anhold 2005). In addition, altered fire regimes, cutting trees for fencing or firewood, and improper grazing by livestock have significant impacts on the quality of sites. Grazing by livestock can modify the fire regime by removing the fine fuels that carry fire. Fire, livestock grazing, and trampling by recreationalists and vehicles disturb cryptogamic soil crusts that help maintain soil structure, reduce soil erosion, provide habitat for plants and preserve biological diversity (Ladyman and Muldavin 1996). More study is needed to understand and manage these woodlands ecologically.

Environmental Description:  This open woodland and scrub alliance is characteristic of exposed rocky mesatops and canyon slopes and rims on the Colorado Plateau, but extends upslope into foothills and on xeric sites. Climate is semi-arid. Summers are hot, and winters are cold with occasional snows and extended periods of freezing temperatures. The seasonality of precipitation varies from east to west with summer rain more common in the southern and eastern portion of the alliance''s range and winter precipitation more common in the western portion of the range. Mean annual precipitation ranges from 20-35 cm. Sites are variable but generally xeric. Stands occur on dry canyon rims, ridges, hills, benches, Hogbacks, mesas, elevated plains and occasionally in intermittent drainages. Elevations generally range from 1400 to 2165 m (4600-7100 feet), but may extend up to 2480 m (8135 feet). Stands typically occur on gentle to moderately steep slopes on all aspects, but range from flat to steep slopes (0-30%). The soils are variable but generally shallow, poorly developed and skeletal, ranging from well-drained clay loam or sandy clay to rapidly drain loamy sand. Exposed sandstone or limestone bedrock and bare soil have high cover, and woody plants are generally rooted in cracks and joints in bedrock. Some stands may also occur on shale slopes covered by sandstone colluvium. Parent materials are variable and frequently include sandstones and shales of the Cedar Mesa, Chinle, and Dakota Formation sandstones, Kayenta Formation, Curtis Formation, Moenkopi Formation, Morrison Formation, Organ Rock shales and eolian silt deposits, and Wingate and Navajo sandstones eroded and redeposited as eolian sands. The unvegetated ground surface is typically composed of litter, bedrock, and bare soil. Because this is a relatively xeric woodland, bare soil and rocks may cover up to 70% of the unvegetated surface, although biological soil crusts provide up to 40% cover.

Geographic Range: The core distribution of this open woodland alliance is the canyon country of the Colorado Plateau.

Nations: US

States/Provinces:  AZ, CA, CO, NM, NV?, UT, WY




Confidence Level: Moderate

Confidence Level Comments: No Data Available

Grank: GNR

Greasons: No Data Available

Type Name Database Code Classification Code
Class 1 Forest & Woodland Class C01 1
Subclass 1.B Temperate & Boreal Forest & Woodland Subclass S15 1.B
Formation 1.B.2 Cool Temperate Forest & Woodland Formation F008 1.B.2
Division 1.B.2.Nc Western North American Pinyon - Juniper Woodland & Scrub Division D010 1.B.2.Nc
Macrogroup 1.B.2.Nc.1 Singleleaf Pinyon - Utah Juniper - Western Juniper Intermountain Woodland Macrogroup M896 1.B.2.Nc.1
Group 1.B.2.Nc.1.a Two-needle Pinyon - Utah Juniper Woodland Group G900 1.B.2.Nc.1.a
Alliance A3573 Two-needle Pinyon - Utah Juniper Colorado Plateau Shrubby Open Woodland Alliance A3573 1.B.2.Nc.1.a
Association CEGL000779 Two-needle Pinyon - Utah Juniper / Littleleaf Mountain-mahogany Woodland CEGL000779 1.B.2.Nc.1.a
Association CEGL000781 Two-needle Pinyon - Utah Juniper / Blackbrush Woodland CEGL000781 1.B.2.Nc.1.a
Association CEGL000782 Two-needle Pinyon - Utah Juniper / Stansbury''s Cliffrose Woodland CEGL000782 1.B.2.Nc.1.a
Association CEGL002118 Two-needle Pinyon - Utah Juniper / Bigelow''s Sagebrush Woodland CEGL002118 1.B.2.Nc.1.a
Association CEGL002148 Two-needle Pinyon - Utah Juniper / Sparse Understory Woodland CEGL002148 1.B.2.Nc.1.a
Association CEGL002328 Two-needle Pinyon - Utah Juniper / Mixed Shrubs Talus Woodland CEGL002328 1.B.2.Nc.1.a
Association CEGL002331 Two-needle Pinyon - Utah Juniper / Black Sagebrush Woodland CEGL002331 1.B.2.Nc.1.a
Association CEGL002332 Two-needle Pinyon - Utah Juniper / Rock-goldenrod Woodland CEGL002332 1.B.2.Nc.1.a
Association CEGL002334 Two-needle Pinyon - Utah Juniper / (Roundleaf Buffaloberry, Utah Serviceberry) Wooded Shrubland CEGL002334 1.B.2.Nc.1.a
Association CEGL002335 Two-needle Pinyon - Utah Juniper / Roundleaf Buffaloberry Woodland CEGL002335 1.B.2.Nc.1.a
Association CEGL002365 Two-needle Pinyon - Utah Juniper / Pygmy Sagebrush Woodland CEGL002365 1.B.2.Nc.1.a
Association CEGL002366 Two-needle Pinyon - Utah Juniper / Saltbush species Woodland CEGL002366 1.B.2.Nc.1.a
Association CEGL002369 Two-needle Pinyon - Utah Juniper / Torrey''s Joint-fir - Bigelow''s Sagebrush Woodland CEGL002369 1.B.2.Nc.1.a
Association CEGL002370 Two-needle Pinyon - Utah Juniper / Mormon-tea - Broom Snakeweed Woodland CEGL002370 1.B.2.Nc.1.a
Association CEGL002374 Two-needle Pinyon - Utah Juniper / Brittle Prickly-pear Woodland CEGL002374 1.B.2.Nc.1.a
Association CEGL002375 Two-needle Pinyon - Utah Juniper / Cushion Plant Woodland CEGL002375 1.B.2.Nc.1.a
Association CEGL002497 Two-needle Pinyon - Utah Juniper / Tucker Sand Shinnery Oak Woodland CEGL002497 1.B.2.Nc.1.a
Association CEGL004005 Two-needle Pinyon - Utah Juniper / Cliff Fendlerbush Woodland CEGL004005 1.B.2.Nc.1.a
Association CEGL004007 Two-needle Pinyon - Utah Juniper / Sonoran Scrub Oak Woodland CEGL004007 1.B.2.Nc.1.a
Association CEGL005657 Two-needle Pinyon - Utah Juniper / Alderleaf Mountain-mahogany - Mixed Shrubs Woodland CEGL005657 1.B.2.Nc.1.a

Concept Lineage: This alliance is composed of 19 associations from Old Alliance II.A.4.N.a. Pinus edulis - (Juniperus spp.) Woodland Alliance (A.516) and one association from Old Alliance III.A.5.N.e. Pinus edulis - Juniperus osteosperma Wooded Shrubland Alliance (A.2649).

Predecessors: No Data Available

Obsolete Names: No Data Available

Obsolete Parents: No Data Available

Synonomy: < Pinus edulis Series (Francis 1986)
< Colorado Pinyon-Utah Juniper Series (Dick-Peddie 1993)

Concept Author(s): K.A. Schulz, in Faber-Langendoen et al. (2013)

Author of Description: K.A. Schulz

Acknowledgements: We have incorporated significant descriptive information previously compiled by J. Coles.

Version Date: 03-14-14

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