Print Report

A0816 Artemisia filifolia Great Plains Sand Prairie Scrub Alliance

Type Concept Sentence: This alliance includes Artemisia filifolia-dominated shrublands occurring mostly in the central and southern Great Plains from Nebraska and Colorado, south to the Trans-Pecos of western Texas and northern Chihuahuan Desert, extending northwest into the Colorado Plateau. These shrublands typically occur on flat, hummocky, or rolling terrain, as well as on partially stabilized dunes and sandsheets. Vegetation cover is sparse to moderately dense, with a shrub stratum approximately 1 m tall, dominated by Artemisia filifolia, interspersed with areas of bare substrate and scattered tall or midgrasses.


Common (Translated Scientific) Name: Sand Sagebrush Great Plains Sand Prairie Scrub Alliance

Colloquial Name: Great Plains Sand Sagebrush Sand Prairie Scrub

Hierarchy Level:  Alliance

Type Concept: This alliance includes Artemisia filifolia-dominated shrublands occurring mostly in the central and southern Great Plains from Colorado and Wyoming, south to the Trans-Pecos of western Texas and northern Chihuahuan Desert, extending northwest into the Colorado Plateau. Vegetation cover is sparse to moderately dense, with a shrub stratum approximately 1 m tall, dominated by Artemisia filifolia, interspersed with areas of bare substrate and scattered tall or midgrasses. Species composition varies with geography, precipitation, disturbance, and soil texture. Associated species may include Andropogon hallii, Artemisia frigida, Bouteloua curtipendula, Bouteloua gracilis, Carex duriuscula, Calamovilfa gigantea, Calamovilfa longifolia, Calylophus serrulatus, Carex inops ssp. heliophila, Helianthus petiolaris, Hesperostipa comata, Heterotheca villosa var. villosa, Ipomoea leptophylla, Lathyrus polymorphus, Lygodesmia juncea, Opuntia spp., Penstemon buckleyi, Prosopis glandulosa, Prunus angustifolia, Psoralidium lanceolatum, Schizachyrium scoparium, Sporobolus giganteus, Sporobolus cryptandrus, and Yucca glauca. Communities associated with gypsum dunes have many gypsophiles or gypsum endemics. Colorado Plateau shrub associates include Ericameria nauseosa, Ephedra torreyana, Ephedra viridis, Gutierrezia sarothrae, Atriplex canescens, and the graminoids Muhlenbergia pungens, Sporobolus cryptandrus, Bouteloua eriopoda, and Achnatherum hymenoides. Some examples found in Texas and Oklahoma may be dominated by Quercus havardii. Degraded examples of this alliance may be dominated by Prunus angustifolia. Some examples in Oklahoma and Texas may be codominated by Quercus havardii. These shrublands typically occur on flat, hummocky, or rolling terrain, as well as on partially stabilized dunes and sandsheets. Soils supporting these communities have low water retention and nutrient availability, and are typically sand or loamy sand, primarily of eolian origin, but include sand deposits derived from sandstone residuum and cinder deposits. Less xeric sites tend to be more grass-dominated. In western Kansas and eastern Colorado, this alliance is found downwind of major waterways where alluvial sand is blown. In Texas these shrublands occur over sandy soils in the Rolling and High Plains and on gypsum dunes in the Trans-Pecos. On the Colorado Plateau, stands occur on a variety of sites including pockets of sand below sandstone cliffs, dunes and sheets of sand or cinder, floodplain terraces and alluvial fans. Timing and amount of growing-season precipitation can greatly affect species abundance from year to year. Drought or overgrazing stands of this alliance will reduce vegetation cover and can allow the wind to cause blowouts or active dunes.

Diagnostic Characteristics: Shrublands dominated by Artemisia filifolia within the central and southern Great Plains. Degraded examples may be dominated by Prunus angustifolia and some examples in Oklahoma and Texas may be codominated by Quercus havardii.

Rationale for Nominal Species or Physiognomic Features: No Data Available

Classification Comments: This alliance contains all the Artemisia filifolia-dominated associations and ranges across the entire Great Plains. More information may suggest a north-south separation. Name of the alliance needs to include better diagnostic species, as this alliance has a name very similar to A3181 (G312). For now, a geographic modifier is used to distinguish it. Stands in the Black Hills region should be assigned to a type in G889, or potentially to G331/A4381. According to Gerry Steinauer (pers. comm. 2019) sandsage in the the southwest corner should go south as it has several southern species associated with it. The Sandsage north of the Platte looses many of these species and it''s not at all clear whether it is true sandsage prairie or just Sandhills prairie with sandsage plants in it (see CEGL002177 for a treatment of that type). At this time, we place Nebraska sandsage in the central - southern great plains and stands north of the Platte are considered outliers. Nebraska sand prairies should be in the northern great plains group (G889).

Similar NVC Types: No Data Available
note: No Data Available

Physiognomy and Structure: Vegetation included in this alliance has a sparse to moderately dense cover of microphyllous evergreen shrubs less than 1.5 m tall. The sparse to moderately dense graminoid layer is dominated by tall, medium-tall or short bunch grasses. Forb cover is generally sparse. Scattered broad-leaved deciduous tall shrubs may be present.

Floristics: This alliance includes Artemisia filifolia-dominated shrublands that occur on sandy sites in the central and southern Great Plainsthe Chihuahuan Desert, and the Colorado Plateau. The vegetation is characterized by a sparse to moderately dense woody layer approximately 1 m tall that is dominated by the microphyllous evergreen shrub Artemisia filifolia. These shrubs usually do not grow as clumps but as individuals with the interstices most often dominated by a sparse to moderately dense layer of tall, mid or short grasses (Bruner 1931, Ramaley 1939a, 1939b, Steinauer 1989, Dick-Peddie 1993). Associated shrub and dwarf-shrub composition varies with geography, precipitation, disturbance, and soils. In the Great Plains, some stands have Prunus angustifolia or Quercus havardii as a codominants in the shrub layer. This species often grows taller than Artemisia filifolia and may form thickets (McGregor and Barkley 1986). In northern stands, Artemisia frigida is more common, and Chihuahuan Desert stands may include Dalea lanata, Psorothamnus scoparius, Tiquilia hispidissima, or Yucca elata. On the Colorado Plateau, Ericameria nauseosa, Ephedra torreyana, Ephedra viridis, Gutierrezia sarothrae, or Atriplex canescens may codominate. Coleogyne ramosissima is typically absent. Species of Opuntia and Yucca are common in many stands throughout its range.
The sparse to moderately dense herbaceous layer is typically dominated by graminoids. The most abundant and widespread species are Achnatherum hymenoides (= Oryzopsis hymenoides), Andropogon hallii, Bouteloua gracilis, Bouteloua curtipendula, Calamovilfa longifolia, Schizachyrium scoparium, or Sporobolus cryptandrus. Carex inops ssp. heliophila, Carex duriuscula (= Carex eleocharis), and Hesperostipa comata (= Stipa comata) are more common in northern stands, and Bouteloua breviseta, Bouteloua eriopoda, Bouteloua trifida, Sporobolus flexuosus, Sporobolus giganteus, and Sporobolus nealleyi are restricted to southern stands. Muhlenbergia pungens, Sporobolus cryptandrus, Bouteloua eriopoda, and Achnatherum hymenoides are important graminoids on the Colorado Plateau. Forbs are typically not abundant in these communities. Associated species include Calylophus serrulatus, Heterotheca villosa var. villosa, Helianthus petiolaris, Ipomoea leptophylla, Lathyrus polymorphus, Lepidium montanum, Lygodesmia juncea, Mentzelia spp., Penstemon buckleyi, and Psoralidium lanceolatum. Communities associated with gypsum dunes have many gypsophiles or gypsum endemics.

Dynamics:  These shrublands occur as any one of several stages in a successional sequence. Drought or overgrazing stands of this alliance will reduce vegetation cover and can allow the wind to cause blowouts or active dunes (Ramaley 1939b). Ramaley (1939b) describes the succession in Colorado from loose sand to a sandhills - mixed community dominated by Muhlenbergia pungens. It then may proceed to an Artemisia filifolia (sand sage) community or skip this stage and succeed to the sand prairie, late seral community dominated by Hesperostipa comata, Calamovilfa longifolia, and Andropogon hallii. This can happen relatively quickly with adequate precipitation and rest from grazing. Ramaley (1939b) also reported that unless protected from overgrazing and fires, the sand sage community will not succeed into the sand prairie community. However, in regions with marginal precipitation, such as occurs over much of eastern Colorado, the sand sage community may be the last successional stage (Ramaley 1939b).

A 10-year grazing study on sand sage pastures in Colorado by Sims et al. (1976) and Dahl and Norris (1965) found that Bouteloua gracilis abundance increased with increased cattle grazing, whereas Calamovilfa longifolia and Hesperostipa comata decreased. With heavy grazing, Artemisia filifolia density increased because of seedling recruitment. This may be due to decreased competition with grasses. In the lightly grazed treatments, Hesperostipa comata abundance more than doubled and the Artemisia filifolia density decreased slightly. Weaver and Albertson (1956) reported Artemisia filifolia and Sporobolus cryptandrus both increasing with grazing in sandhills of Oklahoma.

In Colorado, fire frequency and extent are thought to be low in these stands because sand sage areas are usually surrounded by other communities that are too moist or too sparse to carry a fire well (Ramaley 1939b). In the Great Plains, Wright and Bailey (1980) reported that after fire Artemisia filifolia will resprout and will also reproduce vigorously as seedlings. The shrubs Prunus angustifolia and Rhus spp. also vigorously resprouted after fire (Jackson 1965). Generally, however, fire reduces the vegetation cover that protects these shrublands from blowouts.

Timing and amount of growing-season precipitation can greatly affect species abundance from year to year. Normal to wet springs with a dry summer often result in biomass being dominated by cool-season species such as Hesperostipa comata. A year with a dry spring and normal to wet summer results in biomass being dominated by warm-season species such as Andropogon hallii and Calamovilfa longifolia. Similarly, timing of grazing can have the same result. Forb abundance and diversity can be very high during summers with significantly higher than average precipitation. Panicum virgatum, Sorghastrum nutans, and Prunus pumila var. besseyi are present in low abundance in good condition stands in Colorado but are often eliminated by heavy grazing (Soil Conservation Service 1978).

Environmental Description:  Shrublands included in this alliance occur on sandy sites in the central and southern Great Plains into the Chihuahuan Desert and on the Colorado Plateau. Elevations range from 1122 to 1769 m (3680-5803 feet). The climate is semi-arid to arid, and mean annual precipitation ranges from 20-65 cm. Sites include flat to moderately sloping hummocky or rolling terrain to partially stabilized dunes. Stands can occur on any aspect. The soils are sand or loamy sand, primarily of eolian origin, but include sand deposits derived from sandstone residuum and cinder deposits. All substrates are well-drained to excessively well-drained. Water retention and nutrient availability of the soils are low because water infiltrates rapidly and percolates deeply into the coarse-textured substrate and is therefore only available to deep-rooted plants. In southwestern Kansas and southeastern Colorado, this alliance is found downwind of major waterways where alluvial sand is blown (Johnston 1987). In Texas these shrublands occur on sandy soils in the Rolling and High Plains and on gypsum dunes in the Trans-Pecos (Dick-Peddie 1993). On the Colorado Plateau, stands occur on a variety of sites including pockets of sand below sandstone cliffs, partially stabilized dunes and sheets of cinder or sand, floodplain terraces and alluvial fans. Adjacent vegetation varies by geographic location but is generally grasslands dominated by shortgrass and midgrass prairie species such as Bouteloua gracilis, Pleuraphis jamesii, Calamagrostis canadensis, and Hesperostipa comata. In desert areas it is surrounded by Chihuahuan Desert scrub dominated by Larrea tridentata.

Geographic Range: This alliance occurs on sandy sites in the central to southern Great Plains and Chihuahuan Desert from Colorado and Nebraska south south to Trans-Pecos Texas, southern New Mexico, and Arizona.

Nations: US

States/Provinces:  CO, KS, NE, NM, OK, TX, WY?




Confidence Level: Low

Confidence Level Comments: No Data Available

Grank: GNR

Greasons: No Data Available


Concept Lineage: This contains all the association from the original A.816 and one association from A.780 and one from A.1884

Predecessors: No Data Available

Obsolete Names: No Data Available

Obsolete Parents: No Data Available

Synonomy: ? Artemisia filifolia Series #303 (Johnston 1987)
? Artemisia filifolia Shrubland Alliance (Hoagland 2000)
? Artemisia filifolia shrubland alliance (Hoagland 1998a)
= SRM Cover Type #605 - Sandsage Prairie (Shiflet 1994)
? Sand Sage Community (Ramaley 1939b) [Colorado]
? Sand Sagebrush (Dick-Peddie 1993) [New Mexico]
? Sandsage Prairie (Küchler 1974)

Concept Author(s): K.A. Schulz, in Faber-Langendoen et al. (2013)

Author of Description: S. Menard

Acknowledgements: No Data Available

Version Date: 12-18-14

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