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A2108 Pinus monophylla - Juniperus osteosperma Shrubby Woodland Alliance
Type Concept Sentence: This woodland alliance is characterized by diagnostic tree species Pinus monophylla that forms an open to dense tree layer often with Juniperus osteosperma or, less frequently, Juniperus californica in southern California, with an understory that is characterized by shrubs that typically form an open to moderately dense layer. It is found on dry mountain slopes, foothills, plateaus and ridges of the Great Basin region and eastern foothills of the Sierra Nevada, and south in scattered locations throughout southern California, including desert mountain ranges.
Common (Translated Scientific) Name: Singleleaf Pinyon - Utah Juniper Shrubby Woodland Alliance
Colloquial Name: Great Basin Singleleaf Pinyon - Utah Juniper Shrubby Woodland
Hierarchy Level: Alliance
Type Concept: This woodland alliance is found on dry mountain ranges of the Great Basin region and eastern foothills of the Sierra Nevada, and south in scattered locations throughout southern California, including desert mountain ranges. Pinus monophylla forms an open to dense tree layer often with Juniperus osteosperma or, less frequently, Juniperus californica in southern California. Juniperus osteosperma may also dominate stands as long as there is significant presence of Pinus monophylla that characterizes the stand as a pinyon-juniper stand and not the more xeric, typically lower elevation ~Juniperus osteosperma Great Basin Shrubby Woodland Alliance (A4370)$$. At higher elevations and on relatively mesic sites, such as along drainages, Juniperus scopulorum may be present and sometimes dominant. Other conifers are absent or accidental. The understory is characterized by shrubs that typically form an open to moderately dense layer. Similar scattered shrubs frequently characterize sparse and rocky understory woodlands and so those types are included in this alliance. Characteristic shrubs are many, including Amelanchier utahensis, Arctostaphylos patula, Arctostaphylos pungens, Artemisia arbuscula, Artemisia nova, Artemisia tridentata ssp. tridentata, Artemisia tridentata ssp. vaseyana, Ceanothus greggii, Cercocarpus intricatus, Cercocarpus ledifolius, Coleogyne ramosissima, Ephedra spp., Ericameria spp., Glossopetalon spinescens, Gutierrezia sarothrae, Peraphyllum ramosissimum, Prunus virginiana, Purshia stansburiana, Quercus gambelii, Quercus turbinella, Ribes velutinum, Shepherdia rotundifolia, and Symphoricarpos oreophilus. The herbaceous layer is sparse to moderate and composed of grasses and often diverse forbs. Common species may include Achnatherum hymenoides, Bouteloua gracilis, Hesperostipa comata, Poa fendleriana, Poa secunda, Pleuraphis jamesii, and Pseudoroegneria spicata. The non-native, invasive annual grass Bromus tectorum may become abundant in disturbed stands and dominate the herbaceous layer of highly disturbed stands. This alliance also includes sparse understory stands often with scattered shrubs and grasses. Stands occur on warm, dry sites on mountain slopes, foothills, plateaus and ridges in ranges of the Great Basin and southern California, usually on erosional terrain or upper alluvial slopes. Elevations range from 1000 to 2800 m. The climate is semi-arid, with approximately 20-45 cm of precipitation annually. Substrates are variable but often rocky with shallow soil.
Diagnostic Characteristics: This alliance is characterized by diagnostic tree species Pinus monophylla that forms an open to dense tree layer often with Juniperus osteosperma or, less frequently, Juniperus californica in southern California. Juniperus osteosperma may also dominate stands as long as there is significant presence of Pinus monophylla with which to characterize the stand as a pinyon-juniper stand and not the more xeric, typically lower elevation ~Juniperus osteosperma / Shrub Understory Woodland Alliance (A3496)$$. Understory diagnostic species include Amelanchier utahensis, Arctostaphylos patula, Arctostaphylos patula, Artemisia nova, Artemisia tridentata, Cercocarpus intricatus, Cercocarpus ledifolius, Coleogyne ramosissima, Purshia stansburiana, Purshia tridentata, Quercus gambelii, Quercus turbinella, Shepherdia rotundifolia, and Symphoricarpos oreophilus. This alliance also includes sparse understory stands often with scattered shrubs and grasses. The lack of understory can be caused by harsh substrate (rockland, shale badland) or dense tree canopy resulting from lack of fire that shades out understory.
Rationale for Nominal Species or Physiognomic Features: No Data Available
Classification Comments: Scattered shrubs frequently characterize sparse and rock understory stands and so are included in this alliance.
Similar NVC Types: No Data Available
note: No Data Available
Physiognomy and Structure: These are open to dense woodlands (10-70% cover), dominated by needle-leaved evergreen trees of low stature (<20 m). Scale-leaved evergreen trees or tall shrubs (2-10 m tall) are often present and may be codominant. Generally, evergreen or cold-deciduous shrubs occupy the interstices between trees, interspersed with cespitose graminoids. In total, the ground layer is usually of low to moderate cover (20-40%). This alliance also includes sparse understory stands often with scattered shrubs and grasses.
Floristics: Pinus monophylla forms an open to dense tree layer often with Juniperus osteosperma or, less frequently, Juniperus californica in southern California. Juniperus osteosperma may also dominate stands as long as there is significant presence of Pinus monophylla that characterizes the stand as a pinyon-juniper stand and not the more xeric, typically lower elevation ~Juniperus osteosperma / Shrub Understory Woodland Alliance (A3496)$$. At higher elevations and on relatively mesic sites, such as along drainages, Juniperus scopulorum may be present and sometimes dominant. Other conifers such as Abies concolor, Pinus aristata, Pinus flexilis, Pinus jeffreyi, and Pinus ponderosa are absent or accidental with very low cover. The understory is characterized by shrubs that typically form an open to moderately dense layer. Similar scattered shrubs frequently characterize sparse and rocky understory woodlands and so those types are included in this alliance. Characteristic shrubs are many, including Amelanchier utahensis, Arctostaphylos patula, Arctostaphylos pungens, Artemisia arbuscula, Artemisia nova, Artemisia tridentata ssp. tridentata, Artemisia tridentata ssp. vaseyana, Ceanothus greggii, Cercocarpus intricatus, Cercocarpus ledifolius, Coleogyne ramosissima, Ephedra spp., Ericameria spp., Eriogonum fasciculatum, Glossopetalon spinescens, Gutierrezia sarothrae, Peraphyllum ramosissimum, Prunus virginiana, Purshia stansburiana, Quercus gambelii, Quercus turbinella, Ribes velutinum, Shepherdia rotundifolia, and Symphoricarpos oreophilus. Along the eastern slopes of the Sierra Nevada or Transverse Ranges of southern California, Arctostaphylos glauca, Ceanothus cuneatus, Eriogonum fasciculatum, Quercus chrysolepis, and Yucca schidigera may also occur. The herbaceous layer is sparse to moderate and composed of grasses and often diverse forbs. Common species may include Achnatherum hymenoides, Bouteloua gracilis, Hesperostipa comata, Poa fendleriana, Poa secunda, Pleuraphis jamesii, and Pseudoroegneria spicata. Although forb cover is generally sparse, it may be very diverse. Common forbs include Astragalus spp., Cryptantha cinerea var. jamesii (= Cryptantha jamesii), Eriogonum caespitosum, Eriogonum umbellatum, Gayophytum ramosissimum, Gilia ochroleuca, Hymenoxys richardsonii, Lomatium foeniculaceum ssp. macdougalii (= Lomatium macdougalii), Machaeranthera canescens, Penstemon linarioides, Phlox spp., and Sphaeralcea coccinea. Disturbed stands may have high cover of the introduced annual grass Bromus tectorum or Halogeton glomeratus, an introduced forb. This alliance also includes sparse understory stands often with scattered shrubs and grasses. The lack of understory can be caused by harsh substrate (rockland, shale badland) or dense tree canopy that shades out the understory.
Dynamics: Pinus monophylla is a slow-growing, long-lived tree and stands appear somewhat static over time compared to more productive forests. Regeneration occurs primarily under nurse shrubs or adult trees, where canopy shading minimizes drought stress for seedlings (Koniak 1985). These woodlands are expanding into adjacent steppe grasslands in many areas, reportedly in connection with livestock grazing and altered fire regimes. Many of these woodlands have been intensively altered to enhance livestock forage.
Environmental Description: These woodlands occur on warm, dry sites on mountain slopes, foothills, plateaus and ridges in ranges of the Great Basin and southern California, usually on erosional terrain or upper alluvial slopes. Elevations range from 1000 to 2800 m. The climate of the region is characterized as semi-arid, with warm summers and cold winters. Average annual precipitation is 27 cm (20-45 cm), and shows a peak during April through June, when more than 30% of the annual total is received. Summers are typically dry and there is usually extreme variation in yearly totals. In the Great Basin, these woodlands occur on intermediate slopes above pools of cold-air drainage. Upper elevation limits are determined by local climate and/or the presence of competing tree species. Soils are variable, but generally coarse-textured and well-drained and derived from gneiss, schist, quartzite, and amphibolite. Soil pH is usually nearly neutral or alkaline.
Geographic Range: The core distribution of this woodland alliance is the Great Basin, but it extends out into adjacent ecoregions primarily west into the foothills of the Sierra Nevada, south on scattered mountain ranges of the Mojave Desert and east into the transition zone with the Colorado Plateau. A second substantial range occurs along interior slopes in the Transverse and Peninsular ranges of southern California.
Nations: US
States/Provinces: AZ, CA, ID, NV, UT
Plot Analysis Summary:
http://vegbank.org/natureserve/ELEMENT_GLOBAL.2.866494
Confidence Level: Low
Confidence Level Comments: No Data Available
Grank: GNR
Greasons: No Data Available
Concept Lineage: This alliance is composed of 26 sparse or shrub understory associations from Old Alliance II.A.4.N.a. Pinus monophylla - (Juniperus osteosperma) Woodland Alliance (A.543), two associations from Old Alliance II.A.4.N.a. Juniperus osteosperma Woodland Alliance (A.536), and one association from Old Alliance III.A.5.N.e. Juniperus osteosperma Wooded Shrubland Alliance (A.2541). A.543 (in part), A.536 (in part), and A.2541 (in part).
Predecessors: No Data Available
Obsolete Names: No Data Available
Obsolete Parents: No Data Available
Synonomy: >< Pinus monophylla (Singleleaf pinyon woodlands) Alliance (Sawyer et al. 2009) [87.040.00]
= Pinus monophylla-(Juniperus osteosperma) Woodland Alliance (Evens et al. 2014)
>< Singleleaf Pinyon Series (Sawyer and Keeler-Wolf 1995)
>< Singleleaf Pinyon-Utah Juniper Series (Sawyer and Keeler-Wolf 1995)
>< Utah Juniper Series (Sawyer and Keeler-Wolf 1995)
= Pinus monophylla-(Juniperus osteosperma) Woodland Alliance (Evens et al. 2014)
>< Singleleaf Pinyon Series (Sawyer and Keeler-Wolf 1995)
>< Singleleaf Pinyon-Utah Juniper Series (Sawyer and Keeler-Wolf 1995)
>< Utah Juniper Series (Sawyer and Keeler-Wolf 1995)
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