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G529 Downingia spp. - Callitriche spp. - Eryngium spp. North Pacific Vernal Pool Group

Type Concept Sentence: This group includes wet meadows that form concentric rings around shallow ephemeral pools on hard pan or bedrock with a wide variety of characteristic species such as Callitriche marginata, Downingia elegans, Elatine spp., Eryngium petiolatum, Navarretia leucocephala ssp. diffusa, Pilularia americana, or Triteleia hyacinthina. These vernal pools are found throughout intermountain valleys of Oregon, the San Juan and Gulf islands of Washington and British Columbia, and on the Columbia Plateau in Washington, Oregon, and northern Nevada.


Common (Translated Scientific) Name: Calicoflower species - Water-starwort species - Eryngo species North Pacific Vernal Pool Group

Colloquial Name: Oregon-Washington-British Columbia Vernal Pool

Hierarchy Level:  Group

Type Concept: This group includes herbaceous wetlands that surround and occur within shallow ephemeral water bodies found in depressions among grasslands, shrub-steppe and open woodlands throughout intermountain valleys of Oregon, the San Juan and Gulf islands of Washington and British Columbia, and exposed volcanic scablands of the Columbia Plateau in Washington, Oregon, and northern Nevada. Due to drawdown characteristics, vernal pools typically form concentric rings of similar vegetation. Given their relative isolation in upland-dominated landscapes, many endemic plant species are found in vernal pools. Characteristic species are predominantly annual and diverse. Northern scabland vernal pools share about a third of the species found in northern California vernal pools, but they do not share many of the more common dominant species. Currently very little quantitative plot data have been made available. Eryngium petiolatum and Plagiobothrys figuratus are known dominants that do not occur in California; however, it is assumed many more species could be used to differentiate between these similar groups. Characteristic species in these communities include Callitriche marginata, Callitriche spp., Camissonia tanacetifolia, Deschampsia danthonioides, Downingia elegans, Elatine spp., Eleocharis spp., Epilobium densiflorum, Eryngium petiolatum, Eryngium vaseyi, Grindelia nana, Isoetes orcuttii, Juncus uncialis, Myosurus minimus, Navarretia leucocephala ssp. diffusa, Pilularia americana, Plagiobothrys spp., Plagiobothrys figuratus, Plagiobothrys scouleri, Polyctenium williamsiae, Polygonum polygaloides ssp. confertiflorum, Polygonum polygaloides ssp. polygaloides, Psilocarphus brevissimus, Psilocarphus elatior, Psilocarphus oregonus, Trifolium cyathiferum, Triteleia hyacinthina, and Veronica peregrina. In northern Nevada, most of the species by biomass are perennials and include Carex douglasii, Juncus arcticus ssp. littoralis, Muhlenbergia richardsonis, and species of Eleocharis, Polygonum, Rumex, and Polyctenium. Hardpan vernal pools occur on soils with an indurated clay or cemented (Si or Fe) layer that retains water inputs throughout some portion of the spring, and that typically dry down completely into early summer months. In the Sand Juan and Gulf islands, they are created in small depressions in bedrock. Thus this group only occurs where there is hummocky micro-relief. These wetlands tend to be acidic wetlands. On the Columbia Plateau many pools are located on massive basalt flows, andesite or rhyodacite caprock. Inundation is highly irregular, sometimes not occurring for several years. Depressions usually (but not always) fill with water during winter and spring. They are generally dry again within nine months, though in exceptional times they can remain inundated for two years in a row. Water is from rainfall and snowmelt in relatively small closed basins, on average probably no more than 5-15 times the area of the ponds themselves. Pools are depressions with no outflows. Soils when present are typically silty clay, sometimes with sandy margins. Ponds range from very small (3 sqm) to large depressions (1600 sqm).

Diagnostic Characteristics: Single strata, herbaceous, seasonal dry, seasonal wet wetlands.

Rationale for Nominal Species or Physiognomic Features: No Data Available

Classification Comments: The associations that occur "throughout intermountain valleys of California" are not included here but are in ~Californian Vernal Pool Group (G530)$$. Plot data are needed to better differentiate Oregon, Washington, and British Columbian vernal pools from California pools.

Similar NVC Types: No Data Available
note: No Data Available

Physiognomy and Structure: Annual and perennial herbaceous.

Floristics: Given their relative isolation in upland-dominated landscapes, many endemic plant species are found in vernal pools. Characteristic species are predominantly annual and diverse. Northern scabland vernal pools share about a third of the species found in northern California vernal pools, but they do not share many of the more common dominant species. Currently very little quantitative plot data have been made available. Eryngium petiolatum and Plagiobothrys figuratus are known dominants that do not occur in California; however, it is assumed many more species could be used to differentiate between these similar groups. Characteristic species in these communities include Callitriche marginata, Callitriche spp., Camissonia tanacetifolia, Deschampsia danthonioides, Downingia elegans, Elatine spp., Eleocharis spp., Epilobium densiflorum (= Boisduvalia densiflora), Eryngium petiolatum, Eryngium vaseyi, Grindelia nana, Isoetes orcuttii, Juncus uncialis, Myosurus minimus (= Myosurus x clavicaulis), Navarretia leucocephala ssp. diffusa, Pilularia americana, Plagiobothrys spp., Plagiobothrys figuratus, Plagiobothrys scouleri, Polyctenium williamsiae, Polygonum polygaloides ssp. confertiflorum, Polygonum polygaloides ssp. polygaloides, Psilocarphus brevissimus, Psilocarphus elatior, Psilocarphus oregonus, Trifolium cyathiferum, Triteleia hyacinthina, and Veronica peregrina. In northern Nevada, most of the species by biomass are perennials and include Carex douglasii, Juncus arcticus ssp. littoralis (= Juncus balticus), Muhlenbergia richardsonis, and species of Eleocharis, Polygonum, Rumex, and Polyctenium.

Dynamics:  Pools fill up during winter and spring rains and dry completely by fall.

Environmental Description:  Climate: Mediterranean climate with mild winters and dry summers; interior Washington vernal pools have colder winters. Soil/substrate/hydrology: Vernal pools form in areas with a hummocky micro-relief, within bedrock or soil underlain by a hardpan restricting water drainage. Hardpan vernal pools can have an indurated clay or cemented (Si or Fe) hardpan that retains water inputs throughout some portion of the spring, and typically the depression dries down entirely into early summer months. In the Sand Juan and Gulf islands, they are created in small depressions in bedrock. These wetlands tend to be acidic. In the interior of Washington and Oregon, many pools are located on massive basalt flows, andesite or rhyodacite caprock. Inundation is highly irregular, sometimes not occurring for several years. Depressions usually (but not always) fill with water during winter and spring. They are generally dry again within nine months, though in exceptional times they can remain inundated for two years in a row. Water is from rainfall and snowmelt in relatively small closed basins, on average probably no more than 5-15 times the area of the ponds themselves. Soils are silty clay, sometimes with sandy margins. Environmental information compiled from Chappell and Christy (1994), Bjork (1997), and Bjork and Dunwiddie (2004).

Geographic Range: Vernal pools occur in grasslands and open woodlands throughout intermountain valleys of Oregon and the Gulf and San Juan islands of Washington, and in the northern Columbia Basin and perhaps the Okanagan Valley, up to British Columbia, and into the western portion of the Great Basin in Nevada.

Nations: CA,US

States/Provinces:  BC, NV, OR, WA




Confidence Level: Low

Confidence Level Comments: No Data Available

Grank: GNR

Greasons: No Data Available


Concept Lineage: No Data Available

Predecessors: No Data Available

Obsolete Names: No Data Available

Obsolete Parents: No Data Available

Synonomy: No Data Available

Concept Author(s): C. Chappell and R. Crawford, in Faber-Langendoen et al. (2011)

Author of Description: C. Chappell, R. Crawford, J. Morefield, G. Kittel

Acknowledgements: No Data Available

Version Date: 12-02-15

  • Bjork, C. R. 1997. Vernal pools of the Columbia Plateau of eastern Washington. Report to the Washington Field Office of The Nature Conservancy. 29 pp. plus 7 appendices.
  • Bjork, C. R., and P. W. Dunwiddie. 2004. Floristics and distribution of vernal pools on the Columbia Plateau of eastern Washington. Rhodora 106(928):327-347.
  • Chappell, C., and J. Christy. 2004. Willamette Valley-Puget Trough-Georgia Basin Ecoregion Terrestrial Ecological System EO Specs and EO Rank Specs. Appendix 11 in: J. Floberg, M. Goering, G. Wilhere, C. MacDonald, C. Chappell, C. Rumsey, Z. Ferdana, A. Holt, P. Skidmore, T. Horsman, E. Alverson, C. Tanner, M. Bryer, P. Lachetti, A. Harcombe, B. McDonald, T. Cook, M. Summers, and D. Rolph. Willamette Valley-Puget Trough-Georgia Basin Ecoregional Assessment, Volume One: Report prepared by The Nature Conservancy with support from The Nature Conservancy of Canada, Washington Department of Fish and Wildlife, Washington Department of Natural Resources (Natural Heritage and Nearshore Habitat programs), Oregon State Natural Heritage Information Center and the British Columbia Conservation Data Centre.
  • Faber-Langendoen, D., J. Drake, S. Gawler, M. Hall, C. Josse, G. Kittel, S. Menard, C. Nordman, M. Pyne, M. Reid, L. Sneddon, K. Schulz, J. Teague, M. Russo, K. Snow, and P. Comer, editors. 2010-2019a. Divisions, Macrogroups and Groups for the Revised U.S. National Vegetation Classification. NatureServe, Arlington, VA. plus appendices. [in preparation]
  • Holland, V. L., and D. J. Keil. 1995. California vegetation. Kendall/Hunt Publishing Company, Dubuque, IA. 516 pp.