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CEGL008405 Liriodendron tulipifera - Pinus strobus - (Tsuga canadensis) / Carpinus caroliniana / Amphicarpaea bracteata Forest

Type Concept Sentence: No Data Available


Common (Translated Scientific) Name: Tuliptree - Eastern White Pine - (Eastern Hemlock) / American Hornbeam / American Hog-peanut Forest

Colloquial Name: Central Appalachian Montane Small-Stream Floodplain Forest

Hierarchy Level:  Association

Type Concept: This association comprises small-stream floodplain forests of the shale lowlands of Virginia''s Ridge and Valley and parts of the Northern Blue Ridge. Sites encompass the bottoms and toeslope terraces of small, high-order, more-or-less high-gradient mountain streams at elevations less than 760 m (2500 feet). Liriodendron tulipifera and Pinus strobus are the most constant and abundant canopy trees, but Tsuga canadensis, Quercus alba, and Betula lenta are locally codominant. Many additional species occur as minor canopy or understory trees. Carpinus caroliniana and Cornus florida are dominant in the shrub layer. It has an extremely diverse though not particularly dense herbaceous stratum, with a mixture of wetland and upland species.

Diagnostic Characteristics: No Data Available

Rationale for Nominal Species or Physiognomic Features: No Data Available

Classification Comments: In three studies of national forest landscapes conducted by Virginia Division of Natural Heritage ecologists (Fleming and Moorhead 1996, Rawinski et al. 1996, Fleming and Moorhead 2000), vegetation of small-stream floodplains had the highest species richness of any classified community type, as well as the single plot with the highest richness recorded in the Virginia mountains (Plot PM28, Peters Mountain, Alleghany County, n=122). Twelve plots were classified as this association in the Appalachian Trail classification study (Fleming and Patterson 2009a).

Similar NVC Types: No Data Available
note: No Data Available

Physiognomy and Structure: No Data Available

Floristics: Microtopographic diversity supports a wide range of mesophytic and wetland species, and contributes to notably high species richness (range = 46 to 122 taxa per 400 m2; mean = 73) in plot-sampled stands. Liriodendron tulipifera and Pinus strobus are the most constant and abundant canopy trees, but Tsuga canadensis, Quercus alba, and Betula lenta are locally codominant. Many additional species occur as minor canopy or understory trees, including Acer rubrum, Carya tomentosa (= Carya alba), Carya ovata, Carya cordiformis, Fraxinus americana, Juglans nigra, Magnolia acuminata, Platanus occidentalis, Quercus rubra, and Ulmus americana. Carpinus caroliniana and Cornus florida are dominant in the shrub layer, frequently extending into the lowest (6-10 m tall) tree layer. Other characteristic shrubs are Lindera benzoin, Hamamelis virginiana, Ostrya virginiana, Viburnum prunifolium, Viburnum acerifolium, and Asimina triloba. Climbing lianas, including Parthenocissus quinquefolia, Toxicodendron radicans, Smilax rotundifolia, and Vitis spp., are often common. The herb layer is usually very diverse although not particularly dense or lush. Some of the more constant species in plot-sampled stands include Amphicarpaea bracteata, Botrychium virginianum, Polystichum acrostichoides, Geranium maculatum, Eurybia divaricata (= Aster divaricatus), Desmodium nudiflorum, Actaea racemosa (= Cimicifuga racemosa), Galium triflorum, Stellaria pubera, and Arisaema triphyllum. Herbs that are associated with more frequently flooded streambanks and other wet microhabitats include Carex torta, Dichanthelium clandestinum, Glyceria striata, Impatiens capensis, Leersia virginica, Lycopus virginicus, Ranunculus recurvatus, Thalictrum pubescens, Verbesina alternifolia, and Viola cucullata.

Dynamics:  No Data Available

Environmental Description:  Sites encompass the bottoms and toeslope terraces of small, high-order, more-or-less high-gradient mountain streams at elevations less than 760 m (2500 feet). Hydrologic regime is probably similar to that reported for Passage and Mill creeks in the Massanutten Mountains, where annual water table fluctuations are dramatic and small-scale fluvial landforms greatly influence species distributions (Hupp 1982, 1986, Olson and Hupp 1986). The lowest terraces of these narrow bottoms are probably flooded briefly once a year, while higher terraces are inundated only during rare, catastrophic floods. Slope ranges from 0-10° and aspect is variable. Microtopography is complex and includes floodplains, terraces, fans at tributary hollow mouths, and various bank features. Coarse woody debris, boulders, and cobbles transported by flood events are usually prominent components of the surface substrate. Soils are highly variable and range from well-drained to poorly drained. Soils collected from plot-sampled stands were strongly acidic (mean pH = 5.2) with moderately low base status.

Geographic Range: This community is widely but locally distributed in the Blue Ridge and Ridge and Valley regions of northwestern and west-central Virginia. Occurrences in Maryland, Pennsylvania, and West Virginia are likely.

Nations: US

States/Provinces:  VA, WV




Confidence Level: Moderate

Confidence Level Comments: No Data Available

Grank: G3

Greasons: No Data Available


Concept Lineage: No Data Available

Predecessors: No Data Available

Obsolete Names: No Data Available

Obsolete Parents: No Data Available

Synonomy: = Liriodendron tulipifera - Pinus strobus - (Tsuga canadensis) / Carpinus caroliniana / Amphicarpaea bracteata Forest (Fleming and Patterson 2009a)
= Liriodendron tulipifera - Pinus strobus - (Tsuga canadensis) / Carpinus caroliniana / Amphicarpaea bracteata Forest (Fleming and Coulling 2001)
> Pinus strobus - Tsuga canadensis - Quercus alba / Carpinus caroliniana Association (Rawinski et al. 1996)
> Quercus alba - Pinus strobus / Carpinus caroliniana / Amphicarpaea bracteata Forest (Fleming and Moorhead 2000)
< Yellow-Poplar - White Oak - Northern Red Oak: 59 (Eyre 1980)

Concept Author(s): K.D. Patterson and G.P. Fleming

Author of Description: G. Fleming and P. Coulling

Acknowledgements: No Data Available

Version Date: 02-19-10

  • Eyre, F. H., editor. 1980. Forest cover types of the United States and Canada. Society of American Foresters, Washington, DC. 148 pp.
  • Fleming, G. P., K. D. Patterson, and K. Taverna. 2017. The natural communities of Virginia: A classification of ecological community groups and community types. Third approximation. Version 3.0. Virginia Department of Conservation and Recreation, Division of Natural Heritage, Richmond, VA. [http://www.dcr.virginia.gov/natural-heritage/natural-communities/]
  • Fleming, G. P., and K. D. Patterson. 2009a. A vegetation classification for the Appalachian Trail: Virginia south to Georgia. Virginia Department of Conservation and Recreation, Division of Natural Heritage. In-house analysis, March 2009.
  • Fleming, G. P., and K. Taverna. 2006. Vegetation classification for the National Capitol Region parks, western region. Regional (VA-WVA-MD-DC) analysis prepared for NatureServe and USGS-NPS Vegetation Mapping Program, March 2006. Virginia Department of Conservation and Recreation, Division of Natural Heritage, Richmond.
  • Fleming, G. P., and P. P. Coulling. 2001. Ecological communities of the George Washington and Jefferson national forests, Virginia. Preliminary classification and description of vegetation types. Virginia Department of Conservation and Recreation, Division of Natural Heritage, Richmond, VA. 317 pp.
  • Fleming, G. P., and W. H. Moorhead, III. 2000. Plant communities and ecological land units of the Peter''s Mountain area, James River Ranger District, George Washington and Jefferson national forests, Virginia. Natural Heritage Technical Report 00-07. Virginia Department of Conservation and Recreation, Division of Natural Heritage, Richmond. Unpublished report submitted to the USDA Forest Service. 195 pp. plus appendices.
  • Hupp, C. R. 1982. Stream-grade variation and riparian forest ecology along Passage Creek, Virginia. Bulletin of the Torrey Botanical Club 109:488-499.
  • Hupp, C. R. 1986. Upstream variation in bottomland vegetation patterns, northwestern Virginia. Bulletin of the Torrey Botanical Club 4:421-430.
  • Olson, C. G., and C. R. Hupp. 1986. Coincidence and spatial variability of geology, soils, and vegetation, Mill Run watershed, Virginia. Earth Surface Processes and Landforms 11:619-29.
  • Rawinski, T. J., K. N. Hickman, J. Waller-Eling, G. P. Fleming, C. S. Austin, S. D. Helmick, C. Huber, G. Kappesser, F. C. Huber, Jr., T. Bailey, and T. K. Collins. 1996. Plant communities and ecological land units of the Glenwood Ranger District, George Washington and Jefferson national forests, Virginia. Virginia Department of Conservation and Recreation, Division of Natural Heritage. Natural Heritage Technical Report 96-20. Richmond. 65 pp. plus appendices.
  • Southeastern Ecology Working Group of NatureServe. No date. International Ecological Classification Standard: International Vegetation Classification. Terrestrial Vegetation. NatureServe, Durham, NC.