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A3576 Pinus edulis - Juniperus monosperma Shrubby Woodland Alliance

Type Concept Sentence: This pinyon-juniper woodland and savanna alliance is characterized diagnostic tree species Pinus edulis (>5% cover) that forms a very open to moderately dense tree layer often with Juniperus monosperma present to codominant (or Juniperus scopulorum at higher elevations) with an understory dominated by an open to dense shrub or dwarf-shrub layer. It occurs on warm, dry mountains and foothills in southern Colorado east of the Continental Divide and in mountains and plateaus of northern and central New Mexico, extending east on breaks in the southeastern Great Plains.


Common (Translated Scientific) Name: Two-needle Pinyon - One-seed Juniper Shrubby Woodland Alliance

Colloquial Name: Two-needle Pinyon - One-seed Juniper Shrubby Woodland

Hierarchy Level:  Alliance

Type Concept: These woodlands are characterized by diagnostic tree species Pinus edulis that forms an open to dense tree layer often with Juniperus monosperma. Juniperus monosperma may also dominate stands as long as there is significant presence of Pinus edulis (not accidental) to characterize the stand as a pinyon-juniper stand and not the more xeric, typically lower elevation Juniperus osteosperma woodland. At higher elevations and relatively mesic sites, such as along drainages, Juniperus scopulorum may be present and sometimes dominant. In southern transitional areas with ~Madrean Pinyon - Juniper Woodland Group (G200)$$ in central New Mexico, Juniperus deppeana may be present but not dominant. Other conifers are absent or accidental with very low cover. The understory is characterized by shrubs that typically form an open to moderately dense layer. Characteristic shrubs are many and include Artemisia bigelovii, Artemisia tridentata, Atriplex canescens, Cercocarpus montanus, Chrysothamnus viscidiflorus, Ericameria nauseosa, Gutierrezia sarothrae, Holodiscus dumosus, Quercus gambelii, Quercus x pauciloba, Ribes cereum, Rhus trilobata, and Symphoricarpos oreophilus. Similar scattered shrubs frequently characterize sparse and rocky understory woodland stands and so those types are included in this alliance. The herbaceous layer is sparse to moderate and composed of grasses and often with diverse but low cover of forbs. Common species may include Achnatherum hymenoides, Achnatherum nelsonii, Achnatherum scribneri, Andropogon hallii, Bouteloua gracilis, Festuca arizonica, Hesperostipa comata, Hesperostipa neomexicana, Muhlenbergia montana, and Pleuraphis jamesii. This woodland alliance occurs on warm, dry mountains and foothills in southern Colorado east of the Continental Divide and in mountains and plateaus of northern and central New Mexico, extending east on breaks in the southeastern Great Plains. Stands typically occur on nearly level to steep (to 80%), rocky slopes on hillsides and ridgetops on all aspects except in elevational extremes for a given latitude where low-elevation stands are restricted to the more mesic north slopes; canyons and high-elevation stands occur on south aspects. Sites are typically dry with shallow, rocky, calcareous and alkaline soils. Other sites include eroded "badlands," lava flows, and scree slopes.

Diagnostic Characteristics: This woodland alliance occurs in foothills in southern Colorado east of the Continental Divide and in mountains and plateaus of northern and central New Mexico, extending east on breaks in the southeastern Great Plains. The diagnostic tree species Pinus edulis forms an open to dense tree layer often with Juniperus monosperma. Juniperus monosperma or Juniperus scopulorum may also dominate stands as long as there is significant presence of Pinus edulis (not accidental) to characterize the stand as a pinyon-juniper stand. The understory is characterized by shrubs. Characteristic shrubs are many. The herbaceous layer is sparse to moderate and composed of grasses and often diverse forbs. This alliance also includes sparse understory stands often with scattered shrubs and grasses.

Rationale for Nominal Species or Physiognomic Features: No Data Available

Classification Comments: In northwestern New Mexico, the distributional ranges of Juniperus monosperma and Juniperus osteosperma overlap. This alliance does not occur within the range of Juniperus osteosperma with this transition zone from the southern Rocky Mountain pinyon-juniper woodland alliances to the Colorado Plateau pinyon-juniper woodland alliances. Stands with sparse or rocky understory are included in this alliance because scattered similar shrub species are frequently present and characterize the stand at lower cover. The following associations in this alliance need further review and classification action: ~Pinus edulis - Juniperus scopulorum / Holodiscus dumosus Woodland (CEGL002802)$$ is currently known from Black Canyon of the Gunnison, Curecanti National Recreation Area and Great Sand Dunes in Colorado and El Malpais National Monument in New Mexico. More survey and classification work are needed to fully characterize this type. It may need to be split with a new association to include stands inside the distributional range of Juniperus osteosperma and placed in ~Colorado Plateau Pinyon - Juniper Woodland Group (G900)$$.

Similar NVC Types: No Data Available
note: No Data Available

Physiognomy and Structure: Vegetation included in this alliance has a moderately sparse to moderately dense tree canopy that is typically 3-10 m tall. Stands are either solely dominated by evergreen needle-leaved trees or may be codominated by broad-leaved or scale-leaved evergreen trees. An open to moderately dense shrub layer (0.5-3 m tall) is present and ranges from a single species to a diverse mix of broad-leaved and microphyllous deciduous or evergreen shrubs that are usually less than 3 m tall. A sparse to moderate ground layer dominated by perennial graminoids is usually present. Perennial forbs and cacti are often scattered throughout the stands. Annual forbs and grasses may be seasonally present. This alliance also includes sparse understory stands often with scattered shrubs and grasses.

Floristics: These woodlands are characterized by diagnostic tree species Pinus edulis that forms an open to dense tree layer often with Juniperus monosperma. Juniperus monosperma may also dominate stands as long as there is significant presence of Pinus edulis (not accidental) to characterize the stand as a pinyon-juniper stand and not the more xeric, typically lower elevation Juniperus osteosperma woodland. At higher elevations and relatively mesic sites, such as along drainages, Juniperus scopulorum may be present and sometimes dominant. In southern transitional areas with ~Madrean Pinyon - Juniper Woodland Group (G200)$$ in central New Mexico, Juniperus deppeana may be present but not dominant. Other conifers are absent or accidental with very low cover. The understory is characterized by shrubs that typically form an open to moderately dense layer. Characteristic shrubs are many and include Artemisia bigelovii, Artemisia tridentata, Atriplex canescens, Cercocarpus montanus, Chrysothamnus viscidiflorus, Ericameria nauseosa, Fallugia paradoxa, Gutierrezia sarothrae, Holodiscus dumosus, Quercus gambelii, Quercus x pauciloba, Ribes cereum, Rhus trilobata, and Symphoricarpos oreophilus. Similar scattered shrubs frequently characterize sparse and rocky understory woodland stands and so those types are included in this alliance. The herbaceous layer is sparse to moderate and composed of grasses and often with diverse but low cover of forbs. Common species may include Achnatherum hymenoides, Achnatherum nelsonii, Achnatherum scribneri, Andropogon hallii, Achnatherum hymenoides, Bouteloua gracilis, Festuca arizonica, Hesperostipa comata, Hesperostipa neomexicana, Koeleria macrantha, Muhlenbergia montana, Piptatheropsis micrantha (= Piptatherum micranthum), Pleuraphis jamesii and Sporobolus spp. Many forb species occur, but few have much cover. Commonly present forbs include species of Artemisia, Eriogonum, Heterotheca, Mirabilis, Penstemon, Phlox, Senecio, and Zinnia. Annual grasses and forbs are seasonally present. Adjacent vegetation at higher elevations is typically woodland or forest dominated by Pinus ponderosa. Adjacent vegetation at lower elevations is often Juniperus spp.-dominated woodland and savanna, Artemisia spp.-dominated shrubland, or grassland.

Dynamics:  Pinus edulis is extremely drought-tolerant and slow-growing (Little 1987, Powell 1988b, Muldavin et al. 1998c). It is also non-sprouting and may be killed by fire (Wright et al. 1979). The effect of fire on a stand is largely dependent on the tree height and density, fine-fuel load on the ground, weather conditions, and season (Dwyer and Pieper 1967, Wright et al. 1979). Trees are more vulnerable in open stands where fires frequently occur in the spring, when the relative humidity is low, wind speeds are over 10-20 mph, and there are adequate fine fuels to carry fire (Wright et al. 1979). Under other conditions, burns tend to be spotty with low tree mortality. Large trees are generally not killed unless fine fuels, such as tumbleweeds, have accumulated beneath the tree to provide ladder fuels for the fire to reach the crown (Jameson 1962). Closed-canopy stands rarely burn because they typically do not have enough understory or wind to carry a fire (Wright et al. 1979).

Although Pinus edulis is drought-tolerant, prolonged droughts will weaken trees and promote mortality by secondary agents. Periodic die-offs of pinyon pine caused by insects, such as the pinyon Ips beetle (Ips confusus), or fungal agents, such as blackstain root-rot (Leptographium wageneri), tend to be correlated with droughts (Anhold 2005). These mortality events may be localized or widespread but can result in 50 to 90% mortality of Pinus edulis (Harrington and Cobb 1988).

Climatic and other factors have resulted in denser and expanded pinyon-juniper stands throughout the Colorado Plateau and Great Basin. Denser stands are more susceptible to attack by insects and disease (Anhold 2005). In addition, altered fire regimes, cutting trees for fencing or firewood, and improper grazing by livestock have significant impacts on the quality of sites. Grazing by livestock can modify the fire regime by removing the fine fuels that carry fire. Fire, livestock grazing, and trampling by recreationalists and vehicles disturb cryptogamic soil crusts that help maintain soil structure, reduce soil erosion, provide habitat for plants and preserve biological diversity (Ladyman and Muldavin 1996). More study is needed to understand and manage these woodlands ecologically.

Environmental Description:  This woodland alliance occurs on warm, dry mountains and foothills in southern Colorado east of the Continental Divide and in mountains and plateaus of northern and central New Mexico, extending east on breaks in the southeastern Great Plains. Elevations range from 1406-2510 m. Climate is semi-arid and droughts are not uncommon. Summers are generally hot, and winters range from mild with cold periods and occasional snows in southern New Mexico and Arizona to the more typical extended periods of freezing temperatures. The seasonality of precipitation varies from east to west with summer rain more common in the southern and eastern portion of the alliance''s range and winter precipitation more common in the western portion of the range. Mean annual precipitation ranges from 30-46 cm. Stands typically occur on nearly level to steep (to 80%), rocky slopes on hillsides and ridgetops. Aspect does not seem important except in elevational extremes for a given latitude where low-elevation stands are restricted to the more mesic north slopes, and canyons and high-elevation stands occur on south aspects. Sites are typically dry with shallow, rocky, calcareous and alkaline soils. Other sites include eroded "badlands," lava flows, scree slopes, and deep sands. Soil textures range from sandy loam to clay and are typically derived from limestone, sandstone or shale. Other parent materials include andesite, basalt, granite, quartzite, monzonite, rhyolite and mixed alluvium. This alliance also includes sparse understory stands often with scattered shrubs and grasses. The lack of understory can be caused by harsh substrate (rockland, shale badland) or dense tree canopy resulting from lack of fire that shades out understory.

Geographic Range: This woodland alliance occurs on warm, dry mountains and foothills in southern Colorado east of the Continental Divide and in mountains and plateaus of northern and central New Mexico, extending east on breaks in the southeastern Great Plains.

Nations: US

States/Provinces:  AZ, CO, NM, OK?




Confidence Level: Low

Confidence Level Comments: No Data Available

Grank: GNR

Greasons: No Data Available


Concept Lineage: This alliance is composed of five sparse or shrub understory associations from Old Alliance II.A.4.N.a. Pinus edulis - (Juniperus spp.) Woodland Alliance (A.516).

Predecessors: No Data Available

Obsolete Names: No Data Available

Obsolete Parents: No Data Available

Synonomy: ? Colorado Pinyon-Alligator Juniper Series (Dick-Peddie 1993)
? Colorado Pinyon-Mixed Juniper Series (Dick-Peddie 1993)
? Colorado Pinyon-One Seeded Juniper Series (Dick-Peddie 1993)
? Colorado Pinyon-Rocky Mountain Juniper Series (Dick-Peddie 1993)

Concept Author(s): K.A. Schulz, in Faber-Langendoen et al. (2013)

Author of Description: K.A. Schulz

Acknowledgements: No Data Available

Version Date: 03-14-14

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