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A3571 Pinus edulis - Juniperus osteosperma / Shrub Understory Foothill & Lower Montane Dry-Mesic Woodland Alliance

Type Concept Sentence: This woodland alliance is characterized by diagnostic tree species Pinus edulis that forms a very open to moderately dense tree layer often with Juniperus osteosperma (sometimes with Juniperus monosperma or Juniperus scopulorum) and an understory dominated by an open to dense layer of relatively mesic shrubs or shrubs exceed cover of grasses. It occurs on dry-mesic mountain slopes, foothills, and plateaus in the Colorado Plateau extending east into the west slope of the southern Rocky Mountains.


Common (Translated Scientific) Name: Two-needle Pinyon - Utah Juniper / Shrub Understory Foothill & Lower Montane Dry-Mesic Woodland Alliance

Colloquial Name: Foothill & Lower Montane Pinyon - Juniper / Shrub Dry-Mesic Woodland

Hierarchy Level:  Alliance

Type Concept: This woodland alliance is characterized by diagnostic tree species Pinus edulis that forms an open to dense tree layer often with Juniperus osteosperma or, less frequently, Juniperus monosperma within the range of Juniperus osteosperma in northern Arizona. Juniperus osteosperma may also dominate stands as long as there is significant presence of Pinus edulis (not accidental) to characterize the stand as a pinyon-juniper stand and not the more xeric, typically lower elevation Juniperus osteosperma woodland. At higher elevations and relatively mesic sites, such as along drainages, Juniperus scopulorum may be present and sometimes dominant. Other conifers are absent or accidental with very low cover. The understory is characterized by relatively mesic shrubs that typically form an open to moderately dense layer. Diagnostic shrubs include Amelanchier utahensis, Arctostaphylos patula, Arctostaphylos pungens, Artemisia tridentata ssp. tridentata, Artemisia tridentata ssp. vaseyana, Artemisia tridentata ssp. wyomingensis, Cercocarpus ledifolius, Quercus gambelii, and Symphoricarpos oreophilus. The herbaceous layer is sparse to moderate and composed of grasses often with diverse, but low cover of forbs. Common species may include Achnatherum hymenoides, Bouteloua gracilis, Hesperostipa comata, Hesperostipa neomexicana, Poa fendleriana, Poa secunda, Pleuraphis jamesii, and Pseudoroegneria spicata. The non-native, invasive annual grass Bromus tectorum may become abundant in disturbed stands and dominate the herbaceous layer of highly disturbed stands. These woodlands occur on warm, dry-mesic sites on mountain slopes and foothills and in the higher plateaus and mountains in the Colorado Plateau and the west slope of the southern Rocky Mountains. Stands occur on flat to moderate slopes along drainages, on mesatops, and on moderate to steep, sometimes rocky slopes of foothills, mountains and canyons, especially in draws where soil moisture is concentrated, or on northern aspects or where shaded by upper canyon walls. Aspects are variable and elevations range from 1500-2770 m (4920-9090 feet). The soils are variable but tend to be shallow, gravelly and rapidly drained, ranging from sandy loams to silt clay loams. Litter from shrubs may be extensive (over 50% cover). Parent materials include alluvial, colluvial or eolian deposits derived from sandstone, shale, limestone, granite quartzite and rhyolite.

Diagnostic Characteristics: This woodland alliance occurs in the Colorado Plateau and extends east into the west slope of the southern Rocky Mountains within the distributional range of Juniperus osteosperma. The diagnostic tree species are Pinus edulis and Juniperus osteosperma, either of which may dominate woodland stands as long as there is significant presence of Pinus edulis (not accidental) to characterize as a pinyon-juniper type. The understory is characterized by relatively mesic shrubs. Diagnostic shrubs include Amelanchier utahensis, Arctostaphylos patula, Arctostaphylos pungens, Artemisia tridentata ssp. tridentata, Artemisia tridentata ssp. vaseyana, Artemisia tridentata ssp. wyomingensis, Cercocarpus ledifolius, Quercus gambelii, and Symphoricarpos oreophilus. The herbaceous layer is sparse to moderate and composed of grasses often with diverse, but low cover of forbs.

Rationale for Nominal Species or Physiognomic Features: No Data Available

Classification Comments: In northwestern New Mexico outside the distributional range of Juniperus osteosperma, this alliance transitions into the southern Rocky Mountain pinyon-juniper woodland alliances. Stands with sparse or rock understory are included in this alliance because scattered similar shrub species are frequently present and characterize the stand at lower cover. Additional review is needed to verify alliance placement between this alliance and ~Pinus edulis - Juniperus osteosperma Colorado Plateau Shrubby Open Woodland Alliance (A3573)$$.

~Pinus edulis - Juniperus spp. / Artemisia arbuscula Woodland (CEGL004008)$$ needs further review and classification action - possibly archive because of lack of information. Also review lower elevation stands dominated by Artemisia tridentata ssp. wyomingensis and possibly create a new association and place in ~Pinus edulis - Juniperus osteosperma Colorado Plateau Shrubby Open Woodland Alliance (A3573)$$ to split from higher elevation stands dominated by Artemisia tridentata ssp. vaseyana placed in this alliance (A3571).

Similar NVC Types: No Data Available
note: No Data Available

Physiognomy and Structure: Vegetation included in this alliance has a moderately sparse to moderately dense tree canopy that is typically 3-10 m tall. Stands are either solely dominated by evergreen needle-leaved trees or may be codominated by broad-leaved or scale-leaved evergreen trees. A sparse to moderately dense shrub layer (0.5-3 m tall) is present. If present, the shrub layer ranges from a single species to a diverse mixture of broad-leaved and microphyllous deciduous or evergreen shrubs that are usually less than 3 m tall. A sparse to moderate ground layer dominated by perennial graminoids may be present. Perennial forbs and cacti are often scattered throughout the stands. Annual forbs and grasses may be seasonally present.

Floristics: This woodland alliance is characterized by diagnostic tree species Pinus edulis that forms an open to dense tree layer often with Juniperus osteosperma or, less frequently, Juniperus monosperma within the range of Juniperus osteosperma in northern Arizona. Juniperus osteosperma may also dominate stands as long as there is significant presence of Pinus edulis (not accidental) to characterize the stand as a pinyon-juniper stand and not the more xeric, typically lower elevation Juniperus osteosperma woodland. At higher elevations and relatively mesic sites, such as along drainages, Juniperus scopulorum may be present and sometimes dominant. Other conifers are absent or accidental with very low cover. The understory is characterized by relatively mesic shrubs that typically form an open to moderately dense layer. Diagnostic shrubs include Amelanchier utahensis, Arctostaphylos patula, Arctostaphylos pungens, Artemisia tridentata ssp. tridentata, Artemisia tridentata ssp. vaseyana, Artemisia tridentata ssp. wyomingensis, Cercocarpus ledifolius, Quercus gambelii, and Symphoricarpos oreophilus. Many other shrubs may be present, including Ericameria spp., Gutierrezia sarothrae, Holodiscus dumosus, Opuntia spp., Peraphyllum ramosissimum, Purshia stansburiana, and Shepherdia rotundifolia. The herbaceous layer is sparse to moderate and composed of grasses often with diverse, but low cover of forbs. Common species may include Achnatherum hymenoides, Bouteloua gracilis, Hesperostipa comata, Hesperostipa neomexicana, Poa fendleriana, Poa secunda, Pleuraphis jamesii, and Pseudoroegneria spicata. Commonly present forbs include species of Arenaria, Artemisia, Balsamorhiza, Chenopodium, Cryptantha, Eriogonum, Heterotheca, Geranium, Machaeranthera, Mirabilis, Packera, Penstemon, Phlox, Senecio, Stenotus, Tetraneuris, Thalictrum, Vicia, and Zinnia. Annual grasses and forbs are seasonally present. Seedling Pinus edulis and Juniperus spp. are often present. The non-native, invasive annual grass Bromus tectorum may become abundant in disturbed stands and dominate the herbaceous layer of highly disturbed stands.

Dynamics:  Pinus edulis is extremely drought-tolerant and slow-growing (Little 1987, Powell 1988b, Muldavin et al. 1998c). It is also non-sprouting and may be killed by fire (Wright et al. 1979). The effect of fire on a stand is largely dependent on the tree height and density, fine-fuel load on the ground, weather conditions, and season (Dwyer and Pieper 1967, Wright et al. 1979). Trees are more vulnerable in open stands where fires frequently occur in the spring, when the relative humidity is low, wind speeds are over 10-20 mph, and there are adequate fine fuels to carry fire (Wright et al. 1979). Under other conditions, burns tend to be spotty with low tree mortality. Large trees are generally not killed unless fine fuels, such as tumbleweeds, have accumulated beneath the tree to provide ladder fuels for the fire to reach the crown (Jameson 1962). Closed-canopy stands rarely burn because they typically do not have enough understory or wind to carry a fire (Wright et al. 1979).

Although Pinus edulis is drought-tolerant, prolonged droughts will weaken trees and promote mortality by secondary agents. Periodic die-offs of pinyon pine caused by insects, such as the pinyon Ips beetle (Ips confusus), or fungal agents, such as blackstain root-rot (Leptographium wageneri), tend to be correlated with droughts (Anhold 2005). These mortality events may be localized or widespread but can result in 50 to 90% mortality of Pinus edulis (Harrington and Cobb 1988).

Climatic and other factors have resulted in denser and expanded pinyon-juniper stands throughout the Colorado Plateau and Great Basin. Denser stands are more susceptible to attack by insects and disease (Anhold 2005). In addition, altered fire regimes, cutting trees for fencing or firewood, and improper grazing by livestock have significant impacts on the quality of sites. Grazing by livestock can modify the fire regime by removing the fine fuels that carry fire. Fire, livestock grazing, and trampling by recreationalists and vehicles disturb cryptogamic soil crusts that help maintain soil structure, reduce soil erosion, provide habitat for plants and preserve biological diversity (Ladyman and Muldavin 1996). More study is needed to understand and manage these woodlands ecologically.

Environmental Description:  This woodland alliance occurs on warm, dry-mesic sites on mountain slopes and foothills and in the higher plateaus and mountains in the Colorado Plateau and the western slope of the southern Rocky Mountains. Climate is semi-arid. Summers are generally hot, and winters are cold with occasional snows and extended periods of freezing temperatures. The seasonality of precipitation varies from east to west with summer rain more common in the southern and eastern portion of the alliance''s range and winter precipitation more common in the western portion of the range. Mean annual precipitation ranges from 30-46 cm. Sites are variable but generally are relatively mesic. Stands occur on flat to moderate slopes along drainages and on mesatops, and on moderate to steep, sometimes rocky slopes of foothills, mountains and canyons, especially in draws where soil moisture is concentrated, or on northern aspects or where shaded by upper canyon walls. Aspects are variable and elevations range from 1500-2770 m (4920-9090 feet). Stands in the northern part of the range and at higher elevations (>2400 m) tend to occupy southerly aspects; whereas in the southern part of the range and at lower elevations (<2000 m), stands on northerly aspects are common unless they are located in a moisture-concentrating gully. The soils are variable but tend to be shallow and rapidly drained, ranging from sandy loam, to sandy clay loam, silt loam, and silt clay loam soil textures. They are often gravelly or rocky. Litter from Quercus gambelii and other shrubs may be extensive (over 50% cover). Parent materials include alluvial, colluvial or eolian deposits derived from sandstone, shale, limestone, granite quartzite and rhyolite.

Geographic Range: The core distribution of this woodland alliance is in high plateaus and mountains in the Colorado Plateau extending east into the western slope of the southern Rocky Mountains.

Nations: US

States/Provinces:  AZ, CA, CO, NM, NV, UT, WY




Confidence Level: Moderate

Confidence Level Comments: No Data Available

Grank: GNR

Greasons: No Data Available

Type Name Database Code Classification Code
Class 1 Forest & Woodland Class C01 1
Subclass 1.B Temperate & Boreal Forest & Woodland Subclass S15 1.B
Formation 1.B.2 Cool Temperate Forest & Woodland Formation F008 1.B.2
Division 1.B.2.Nc Western North American Pinyon - Juniper Woodland & Scrub Division D010 1.B.2.Nc
Macrogroup 1.B.2.Nc.1 Singleleaf Pinyon - Utah Juniper - Western Juniper Intermountain Woodland Macrogroup M896 1.B.2.Nc.1
Group 1.B.2.Nc.1.a Two-needle Pinyon - Utah Juniper Woodland Group G900 1.B.2.Nc.1.a
Alliance A3571 Two-needle Pinyon - Utah Juniper / Shrub Understory Foothill & Lower Montane Dry-Mesic Woodland Alliance A3571 1.B.2.Nc.1.a
Association CEGL000775 Two-needle Pinyon - Utah Juniper / Pointleaf Manzanita Woodland CEGL000775 1.B.2.Nc.1.a
Association CEGL000789 Two-needle Pinyon - Utah Juniper / Antelope Bitterbrush Woodland CEGL000789 1.B.2.Nc.1.a
Association CEGL002329 Two-needle Pinyon - Utah Juniper / Utah Serviceberry Woodland CEGL002329 1.B.2.Nc.1.a
Association CEGL002939 Two-needle Pinyon - Utah Juniper / Greenleaf Manzanita Woodland CEGL002939 1.B.2.Nc.1.a
Association CEGL002940 Two-needle Pinyon - Utah Juniper / Curl-leaf Mountain-mahogany Woodland CEGL002940 1.B.2.Nc.1.a
Association CEGL004008 Two-needle Pinyon - Juniper species / Little Sagebrush Woodland CEGL004008 1.B.2.Nc.1.a
Association CEGL005651 Two-needle Pinyon - (One-seed Juniper, Alligator Juniper) / Gambel Oak Woodland CEGL005651 1.B.2.Nc.1.a
Association CEGL005658 Two-needle Pinyon - Utah Juniper / (Wyoming Big Sagebrush, Mountain Big Sagebrush) Woodland CEGL005658 1.B.2.Nc.1.a
Association CEGL005659 Two-needle Pinyon - Utah Juniper / Gambel Oak Woodland CEGL005659 1.B.2.Nc.1.a

Concept Lineage: This alliance is composed of 8 shrub understory associations from Old Alliance II.A.4.N.a. Pinus edulis - (Juniperus spp.) Woodland Alliance (A.516). A.516 (in part)

Predecessors: No Data Available

Obsolete Names: No Data Available

Obsolete Parents: No Data Available

Synonomy: ? Pinus edulis (Two-needle pinyon stands) Special Stands (Sawyer et al. 2009) [87.050.00]
< Pinus edulis Series (Francis 1986)
>< Pinus edulis Woodland Alliance (Evens et al. 2014)
< Colorado Pinyon-Utah Juniper Series (Dick-Peddie 1993)

Concept Author(s): K.A. Schulz, in Faber-Langendoen et al. (2013)

Author of Description: K.A. Schulz

Acknowledgements: We have incorporated significant descriptive information previously compiled by J.J. Coles.

Version Date: 03-14-14

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