Print Report
A3392 Pseudotsuga menziesii - Pinus ponderosa / Shrub Understory Central Rocky Mountain Forest & Woodland Alliance
Type Concept Sentence: This alliance represents montane coniferous forests found in the interior Pacific Northwest that are dominated by a mix of Pseudotsuga menziesii and Pinus ponderosa (but there can be stands without the latter) with a shrub-dominated understory.
Common (Translated Scientific) Name: Douglas-fir - Ponderosa Pine / Shrub Understory Central Rocky Mountain Forest & Woodland Alliance
Colloquial Name: Central Rocky Mountain Douglas-fir / Shrub Forest & Woodland
Hierarchy Level: Alliance
Type Concept: Most occurrences of this montane coniferous forest alliance are dominated by a mix of Pseudotsuga menziesii and Pinus ponderosa (but there can be stands without the latter) and sometimes with other species, including Pinus contorta, Pinus monticola, and Larix occidentalis. This understory is characterized by open to dense layer of shrubs and dwarf-shrubs. Characteristic species include Arctostaphylos uva-ursi, Holodiscus discolor, Juniperus communis, Paxistima myrsinites, Penstemon fruticosus, Physocarpus malvaceus, Purshia tridentata, Spiraea betulifolia, Symphoricarpos albus, or Vaccinium membranaceum. There herbaceous layer is variable. If herbaceous cover exceeds the shrub cover, then there is a distinct shrub layer present with at least 10% cover. This alliance is found in the interior Pacific Northwest, from southernmost interior British Columbia, eastern Washington, eastern Oregon, northern Idaho, western and northern Montana, and south along the east slope of the Cascades in Washington and Oregon. It is associated with a submesic climate regime with annual precipitation ranging from 40 to >150 cm, with a maximum in winter or late spring. Winter snowpacks typically melt off in early spring at lower elevations. Elevations range from 460 to 2400 m (1500-7900 feet). Stands occur on cool, dry sites on mid to upper slopes, steep slopes, ridgelines, rocky sites and benches on all aspects. Slopes can be moderate to steep (3-60%), and soils are derived from pumice ash, lava, tuff, and granitics. Surface rocks can be abundant. It can be associated with glacial outwash or till in cold-air drainages.
Diagnostic Characteristics: Woodlands characterized by dominance of Pseudotsuga menziesii or codominated by Pinus ponderosa with understories characterized by shrubs and dwarf-shrubs, such as Arctostaphylos uva-ursi, Juniperus communis, Physocarpus malvaceus, Purshia tridentata, Symphoricarpos albus, or Vaccinium membranaceum.
Rationale for Nominal Species or Physiognomic Features: No Data Available
Classification Comments: Included in this alliance are some broadly defined associations such as ~Pinus ponderosa - Pseudotsuga menziesii / Purshia tridentata Woodland (CEGL000214)$$ that is reported to occur in the southern Rocky Mountains (Utah) and will need to be reviewed and concept clarified.
Similar NVC Types: No Data Available
note: No Data Available
Physiognomy and Structure: Vegetation included in this alliance is characterized by open to closed stands of evergreen needle-leaved trees 15-50 m tall. There is a sparse cover of understory trees (5-15 m tall). Shrubs (0.5-2 m tall) generally form a layer (>10% cover) but may be sparse on rocky sites and still characterize the understory. The herbaceous layer is sparse under denser canopies and on rock substrates, but is typically moderately dense. If herbaceous cover exceeds the shrub cover, then there is a distinct shrub layer present with at least 10% cover. The herbaceous layer is usually dominated by perennial graminoids with forbs, but occasionally perennial forbs are dominant. Annual grasses and forbs are seasonally present.
Floristics: Most occurrences of this alliance are dominated by a mix of Pseudotsuga menziesii and Pinus ponderosa (but there can be stands without the latter) and sometimes with other species, including Pinus contorta, Pinus monticola, and Larix occidentalis. At the southern end of the alliance''s range, scattered Pinus lambertiana, Abies concolor, and Calocedrus decurrens can be found in the stands, but Abies grandis will not be found, although trace amounts of Abies grandis may occur in northern portion of its range. This understory is characterized by open or patchy to dense layer of shrubs and dwarf-shrubs. Characteristic species include Amelanchier alnifolia, Arctostaphylos uva-ursi, Arctostaphylos nevadensis, Arctostaphylos patula, Chrysolepis chrysophylla (= Castanopsis chrysophylla), Ceanothus velutinus, Holodiscus discolor, Juniperus communis, Mahonia nervosa, Paxistima myrsinites, Penstemon fruticosus, Physocarpus malvaceus, Purshia tridentata, Spiraea betulifolia, Symphoricarpos albus, or Vaccinium membranaceum may occur. The herbaceous layer is variable depending on density of shrub layer, but is usually dominated by graminoids. Important or dominant graminoids are Achnatherum occidentale (= Stipa occidentalis), Bromus ciliatus, Calamagrostis rubescens, Carex geyeri, Carex rossii, Festuca idahoensis, Festuca occidentalis, Koeleria macrantha, Luzula parviflora, and Pseudoroegneria spicata. Characteristic forbs include Apocynum androsaemifolium, Arnica cordifolia, Balsamorhiza sagittata, Eriophyllum lanatum, Moehringia macrophylla (= Arenaria macrophylla), Osmorhiza berteroi, Pteridium aquilinum, Thalictrum occidentale, Viola adunca, and species of many other genera, including Arenaria, Erigeron, Eriogonum, Fragaria, Galium, Lathyrus, Lupinus, Penstemon, and Vicia. If herbaceous cover exceeds the shrub cover, then there is a distinct shrub layer present with at least 10% cover.
Dynamics: Vegetation within this alliance is often comprised of relatively young stands, which were historically subject to recurrent fires. Both Pinus ponderosa and Pseudotsuga menziesii are tolerant of surface fires, and many mature trees have fire scars. The fire frequency is 5-30 years. Fire suppression has favored Pseudotsuga menziesii, but canopy closure is rare because of the harshness of these sites (Williams et al. 1990b). In addition, Pseudotsuga menziesii is more tolerant than Pinus ponderosa. Following fire, regeneration of Pinus ponderosa is aggressive. With increasing stand age, Pseudotsuga menziesii comes to dominate regeneration and eventually attains canopy dominance. Some stands require fire or other disturbance at moderate return intervals to maintain a mixed canopy. For example, in northern stands, Abies grandis will eventually become the canopy dominant with fire suppression. This alliance also occurs as an edaphic climax on moderate to steep slopes, in canyons, and in places along the ecotone between the Pseudotsuga menziesii and Pinus ponderosa zones. The exotic grass Bromus tectorum often invades the herbaceous layer of disturbed stands.
Environmental Description: This alliance is composed of montane coniferous forests found in the interior Pacific Northwest, from southernmost interior British Columbia, eastern Washington, eastern Oregon, northern Idaho, western and northern Montana, and south along the east slope of the Cascades in Washington and Oregon. Elevations range from 460 to 2400 m (1500-7900 feet). Climate is mostly temperate, continental with maritime influences and is strongly influenced by the rainshadow effect of the Cascade Range or Coast Ranges of southern Oregon. Winter snowpacks typically melt off in early spring at lower elevations. Annual precipitation ranges from 40 cm in the Blue Mountains, up to 150-230 cm in the Cascades. Sites are typically on hot, dry, middle to upper slopes near grassy ridgetops. Slopes are moderate to steep with typically southern to western aspects, but stands may occur on northern slopes at low elevations. Substrates are typically shallow, lithic, course-textured soils derived from colluvium and residuum. Soil texture ranges from gravelly sand to sandy loam. There is often high surface cover of rock (to 50%), and coarse fragments average 40%. Parent materials may include granitic till or outwash, basalt, lava, pumice ash, tuff, granitics and sandstone. These rocky, shallow soils have so little moisture-holding capacity that these sites are effectively very dry for plant growth despite the substantial input of precipitation on most sites. These sites are exposed to high winds. Wind detracts from soil moisture status by blowing off snow, directly desiccating plants, and eroding soil.
Geographic Range: This alliance is composed of montane coniferous forests found in the interior Pacific Northwest, from southernmost interior British Columbia, eastern Washington, eastern Oregon, northern Idaho, western and northern Montana, and south along the east slope of the Cascades in Washington and Oregon. Peripheral stands are found on relatively dry sites in the northern Cascades such as Mount Rainier.
Nations: CA,US
States/Provinces: AB, BC, CA?, ID, MT, OR, UT, WA
Plot Analysis Summary:
http://vegbank.org/natureserve/ELEMENT_GLOBAL.2.899516
Confidence Level: Low
Confidence Level Comments: No Data Available
Grank: GNR
Greasons: No Data Available
Concept Lineage: This alliance is based on A.NCCN-036 Pseudotsuga menziesii - (Pinus ponderosa) Forest Alliance. I have split this alliance further into shrub- and grass-dominated understory alliances. This new alliance is based in part on three old NVC alliances including Old Alliance I.A.8.N.b. Pinus ponderosa - Pseudotsuga menziesii Forest Alliance (A.134), Old Alliance I.A.8.N.c. Pseudotsuga menziesii Forest Alliance (A.157), and Old Alliance II.A.4.N.a. Pinus ponderosa - Pseudotsuga menziesii Woodland Alliance (A.533).
Predecessors: No Data Available
Obsolete Names: No Data Available
Obsolete Parents: No Data Available
Synonomy: >< Pinus ponderosa - Pseudotsuga menziesii Woodlands (Chappell et al. 1997)
> Pinus ponderosa-Pseudotsuga menziesii/Purshia tridentata Woodland (Crawford et al. 2009)
> Pseudotsuga menziesii - Pinus ponderosa / Purshia tridentata Plant Association (Williams and Smith 1990)
> Pseudotsuga menziesii / Arctostaphylos uva-ursi - Purshia tridentata Plant Association (Williams and Lillybridge 1983)
> Pseudotsuga menziesii / Arctostaphylos uva-ursi - Purshia tridentata Plant Association (Williams and Smith 1990)
> Pseudotsuga menziesii / Arctostaphylos uva-ursi (Henderson et al. 1989)
> Pseudotsuga menziesii / Holodiscus discolor / Carex geyeriAssociation (Topik et al. 1988)
> Pseudotsuga menziesii-(Pinus contorta)/Arctostaphylos uva-ursi/Racomitrium canescens Woodland (Crawford et al. 2009)
>< Interior Douglas-fir: 210 (Eyre 1980)
>< Interior Ponderosa Pine: 237 (Eyre 1980)
> Pinus ponderosa-Pseudotsuga menziesii/Purshia tridentata Woodland (Crawford et al. 2009)
> Pseudotsuga menziesii - Pinus ponderosa / Purshia tridentata Plant Association (Williams and Smith 1990)
> Pseudotsuga menziesii / Arctostaphylos uva-ursi - Purshia tridentata Plant Association (Williams and Lillybridge 1983)
> Pseudotsuga menziesii / Arctostaphylos uva-ursi - Purshia tridentata Plant Association (Williams and Smith 1990)
> Pseudotsuga menziesii / Arctostaphylos uva-ursi (Henderson et al. 1989)
> Pseudotsuga menziesii / Holodiscus discolor / Carex geyeriAssociation (Topik et al. 1988)
> Pseudotsuga menziesii-(Pinus contorta)/Arctostaphylos uva-ursi/Racomitrium canescens Woodland (Crawford et al. 2009)
>< Interior Douglas-fir: 210 (Eyre 1980)
>< Interior Ponderosa Pine: 237 (Eyre 1980)
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