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A3382 Carex gynandra - Glyceria melicaria - Glyceria striata Seep Alliance

Type Concept Sentence: These are Southern Appalachian, primarily high-elevation seeps and other non-alluvial wetlands found near streams, in flats away from the immediate streambed, along wetland margins, and in seepage-fed oxbow fens. Examples are generally graminoid-dominated but may have significant coverage by trees or shrubs, especially around their edges.


Common (Translated Scientific) Name: Nodding Sedge - Melic Mannagrass - Fowl Mannagrass Seep Alliance

Colloquial Name: Southern Appalachian Herbaceous Seep

Hierarchy Level:  Alliance

Type Concept: These are Southern Appalachian, primarily high-elevation seeps and other non-alluvial wetlands found on a variety of sites associated with groundwater seepage. Some characteristic herbaceous species include Calopogon sp., Carex atlantica, Carex gynandra, Carex leptalea ssp. harperi, Carex lurida, Carex scoparia var. scoparia, Carex stipata, Chelone glabra, Doellingeria umbellata, Dryopteris cristata, Eriophorum virginicum, Glyceria melicaria, Glyceria striata, Impatiens capensis, Lycopus virginicus, Lygodium palmatum, Platanthera clavellata, Platanthera flava var. flava, Pogonia ophioglossoides, Osmunda cinnamomea, Osmunda regalis var. spectabilis, Oxypolis rigidior, Polygonum sagittatum, Solidago patula var. patula, Solidago rugosa, and Solidago uliginosa. Sphagnum spp. may occur in mats and include Sphagnum palustre and Sphagnum recurvum. Shrubs can occur as scattered clumps or zones and include Hypericum densiflorum, Ilex opaca, Kalmia latifolia, Aronia arbutifolia, Rhododendron maximum, Salix sericea, and Viburnum nudum var. cassinoides. At higher elevations, component species may include Calamagrostis cainii, Carex debilis, Carex misera, Carex ruthii, Dennstaedtia punctilobula, Gentiana linearis, Glyceria nubigena, Hypericum graveolens, Krigia montana, Parnassia asarifolia, and Solidago glomerata. Woody species in these examples include Abies fraseri, Picea rubens, Prunus pensylvanica, Rhododendron catawbiense, Rubus canadensis, and Vaccinium erythrocarpum. Sites are influenced by high rainfall and low evaporation rates in the mountainous landscapes. Examples may be small-scale patches. These non-alluvial wetlands are generally graminoid-dominated but may have significant coverage by trees or shrubs, especially around their edges. Particularly small examples may be completely shaded by trees within the community occurrence and in the adjacent forests. Examples of this alliance may occur on a variety of sites associated with groundwater seepage. These include areas of flat to very gently sloping topography (0-2°) located near streams, in flats away from the immediate streambed, along wetland margins, and in seepage-fed oxbow fens. These small palustrine, open seeps generally occur at higher elevations, greater than 1200 m [4000 feet]), on upper slopes and ridgetops. Some associations occur on pronounced slopes at elevations over 1525 m (5000 feet).

Diagnostic Characteristics: These are Southern Appalachian herbaceous seeps. This group of associations is not very "boggy" (i.e., they do not have a particularly organic or mucky soil, and are not "rich" or mafic in their soil and water chemistry). They are primarily graminoid-dominated but one distinctive association (CEGL004293) is forb-dominated.

Rationale for Nominal Species or Physiognomic Features: No Data Available

Classification Comments: No Data Available

Similar NVC Types: No Data Available
note: No Data Available

Physiognomy and Structure: Examples may be small-scale patches. Examples are generally graminoid-dominated but may have significant coverage by trees or shrubs, especially around their edges. Shrubs can occur as scattered clumps or zones. One distinctive association (CEGL004293) is forb-dominated. Particularly small examples may be completely shaded by trees within the community occurrence and in the adjacent forests. Some component associations have well-developed Sphagnum mats.

Floristics: Some characteristic herbaceous species include Calopogon sp., Carex atlantica, Carex gynandra, Carex leptalea ssp. harperi, Carex lurida, Carex scoparia var. scoparia, Carex stipata, Chelone glabra, Doellingeria umbellata (= Aster umbellatus), Dryopteris cristata, Eriophorum virginicum, Glyceria melicaria, Glyceria striata, Impatiens capensis, Lycopus virginicus, Lygodium palmatum, Platanthera clavellata, Platanthera flava var. flava, Pogonia ophioglossoides, Osmunda cinnamomea, Osmunda regalis var. spectabilis, Oxypolis rigidior, Polygonum sagittatum, Solidago patula var. patula, Solidago rugosa, and Solidago uliginosa. Sphagnum spp. may occur in mats and include Sphagnum palustre and Sphagnum recurvum. One distinctive association is forb-dominated. Its stands may lack extensive Sphagnum and include Aconitum reclinatum, Cardamine clematitis, Carex debilis var. rudgei (= Carex flexuosa), Carex leptonervia, Carex ruthii, Chelone lyonii, Cicuta maculata, Claytonia caroliniana, Conioselinum chinense, Euonymus obovatus, Geum geniculatum, Helenium autumnale, Houstonia serpyllifolia, Lilium superbum, Lilium grayi, Packera aurea (= Senecio aureus), Solidago patula, Thalictrum clavatum, Trautvetteria caroliniensis, Veratrum viride, Viola cucullata, and Viola macloskeyi ssp. pallens. Shrubs can occur as scattered clumps or zones and include Hypericum densiflorum, Ilex opaca, Kalmia latifolia, Aronia arbutifolia, Rhododendron maximum, Salix sericea, and Viburnum nudum var. cassinoides. At higher elevations, component species may include Calamagrostis cainii, Carex debilis, Carex misera, Carex ruthii, Dennstaedtia punctilobula, Gentiana linearis, Glyceria nubigena, Hypericum graveolens, Krigia montana, Parnassia asarifolia, and Solidago glomerata. Woody species in these examples include Abies fraseri, Picea rubens, Prunus pensylvanica, Rhododendron catawbiense, Rubus canadensis, and Vaccinium erythrocarpum.

Dynamics:  Sites are influenced by the high rainfall and low evaporation rates in the mountainous landscapes. Some areas can be modified by beaver activity, though they do not appear to be formed by beavers.

Environmental Description:  Examples of this alliance may occur on a variety of sites associated with groundwater seepage. These include areas of flat to very gently sloping topography (0-2°) located near streams, in flats away from the immediate streambed, along wetland margins, and in seepage-fed oxbow fens. These small palustrine, open seeps generally occur at higher elevations, greater than 1200 m [4000 feet]), on upper slopes and ridgetops. Some associations occur on pronounced slopes at elevations over 1525 m (5000 feet).

Geographic Range: This vegetation occurs at moderate to high elevations in the southern Blue Ridge Mountains and adjacent ecoregions of West Virginia, eastern Kentucky, southwestern Virginia, western North Carolina, northwestern South Carolina, eastern Tennessee, and northern Georgia. It may be found in northern Alabama as well.

Nations: US

States/Provinces:  AL?, GA, KY, NC, SC, TN, VA, WV




Confidence Level: Moderate

Confidence Level Comments: No Data Available

Grank: GNR

Greasons: No Data Available

Type Name Database Code Classification Code
Class 2 Shrub & Herb Vegetation Class C02 2
Subclass 2.C Shrub & Herb Wetland Subclass S44 2.C
Formation 2.C.4 Temperate to Polar Freshwater Marsh, Wet Meadow & Shrubland Formation F013 2.C.4
Division 2.C.4.Nd Eastern North American Temperate Freshwater Marsh, Wet Meadow & Shrubland Division D323 2.C.4.Nd
Macrogroup 2.C.4.Nd.1 Shrubby-cinquefoil - Canadian Burnet / Inland Sedge Seep Macrogroup M061 2.C.4.Nd.1
Group 2.C.4.Nd.1.a Catawba Rosebay / Tawny Cottongrass / Peatmoss species Shrub & Herb Seep Group G184 2.C.4.Nd.1.a
Alliance A3382 Nodding Sedge - Melic Mannagrass - Fowl Mannagrass Seep Alliance A3382 2.C.4.Nd.1.a
Association CEGL004293 (Orange Jewelweed, Pale Touch-me-not) - Scarlet Beebalm - Greenhead Coneflower Herbaceous Seep CEGL004293 2.C.4.Nd.1.a
Association CEGL004296 American Umbrella-leaf - Lettuceleaf Saxifrage - Canadian Woodnettle Forested Herbaceous Seep CEGL004296 2.C.4.Nd.1.a
Association CEGL007043 Stiff Cowbane - Roundleaf Sundew - Small Green Wood Orchid - Maryland Meadowbeauty Herbaceous Seep CEGL007043 2.C.4.Nd.1.a
Association CEGL007697 Nodding Sedge - Small Green Wood Orchid - Roundleaf Sundew - Ruth''s Sedge / Peatmoss species Herbaceous Seep CEGL007697 2.C.4.Nd.1.a
Association CEGL007771 Nodding Sedge - Woolgrass - Tawny Cottongrass - Cinnamon Fern Herbaceous Seep CEGL007771 2.C.4.Nd.1.a
Association CEGL007877 Cain''s Reedgrass - Ruth''s Sedge - Kidneyleaf Grass-of-Parnassus / Peatmoss species Herbaceous Seep CEGL007877 2.C.4.Nd.1.a
Association CEGL008438 Fowl Mannagrass - Nodding Sedge - White Turtlehead - Purple-stem Aster / Peatmoss species Herbaceous Seep CEGL008438 2.C.4.Nd.1.a

Concept Lineage: This alliance integrates members from the old alliances A.1451 (1/5), A.1690 (1/1), A.1898 (2/2), A.2016 (1/1); one association placed here (CEGL007043) was not part of any old alliance.

Predecessors: No Data Available

Obsolete Names: No Data Available

Obsolete Parents: No Data Available

Synonomy: ? Carex gynandra Wetland (Newell and Peet 1996a)
? Carex ruthii Wetland (Newell and Peet 1996a)
>< IID3a. Herbaceous High Elevation Seepage Slope (Allard 1990)

Concept Author(s): M. Pyne, in Faber-Langendoen et al. (2013)

Author of Description: M. Pyne

Acknowledgements: We have incorporated significant descriptive information previously compiled by A.S. Weakley, and K.D. Patterson.

Version Date: 09-26-14

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