Print Report

A3208 Artemisia tridentata ssp. vaseyana - Mixed Steppe & Shrubland Alliance

Type Concept Sentence: This alliance is widespread in mountainous areas across the western U.S. The vegetation is characterized by a moderate to dense shrub layer in which Artemisia tridentata ssp. vaseyana is codominant with non-sagebrush shrub species Amelanchier utahensis, Holodiscus dumosus, Purshia tridentata, or Symphoricarpos oreophilus. Perennial graminoids typically dominate the open to moderately dense herbaceous layer. This alliance forms large, continuous stands on mid-elevation mountain slopes and foothills, and can extend above the lower treeline as patches within montane or subalpine coniferous forests.


Common (Translated Scientific) Name: Mountain Big Sagebrush - Mixed Steppe & Shrubland Alliance

Colloquial Name: Mountain Big Sagebrush - Mixed Steppe & Shrubland

Hierarchy Level:  Alliance

Type Concept: This alliance is widespread in mountainous areas across the western U.S. The vegetation included in this alliance is characterized by an open to dense (10-70% cover) shrub layer in which Artemisia tridentata ssp. vaseyana is codominant, usually with 40-60% relative cover with non-sagebrush shrub species such as Amelanchier utahensis, Holodiscus dumosus, Purshia tridentata, or Symphoricarpos oreophilus. Perennial graminoids typically dominate the open to moderately dense herbaceous layer. The most widespread species are Pseudoroegneria spicata and Festuca idahoensis, which occur from the Columbia Basin to the northern Rockies, although they may not be the most abundant species in individual stands. Other locally important species may include Achnatherum occidentale, Bouteloua gracilis, Bromus carinatus, Elymus trachycaulus, Festuca thurberi, Festuca viridula, Koeleria macrantha, Leucopoa kingii, Leymus cinereus, Pascopyrum smithii, Poa fendleriana, and Poa secunda. The forb layer is variable and can be very diverse. Species of Castilleja, Potentilla, Erigeron, Phlox, Astragalus, Geum, Lupinus, and Eriogonum are characteristic. The alliance forms large, continuous stands on mid-elevation mountain slopes and foothills, and can extend above the lower treeline as patches within montane or subalpine coniferous forests. Sites are variable and range from flats to steep slopes to ridgetops with deep to shallow rocky soil.

Diagnostic Characteristics: This sagebrush mixed shrub alliance is widespread in mountainous areas across the western U.S. The vegetation included in this alliance is characterized by a moderate to dense shrub layer in which Artemisia tridentata ssp. vaseyana is codominant with non-sagebrush shrub species Amelanchier utahensis, Holodiscus dumosus, Purshia tridentata, or Symphoricarpos oreophilus. Perennial graminoids typically dominate the open to moderately dense herbaceous layer. Characteristic grasses are Bromus carinatus, Elymus trachycaulus ssp. trachycaulus, Festuca idahoensis, Hesperostipa comata, Muhlenbergia montana, Poa secunda, and Pseudoroegneria spicata.

Rationale for Nominal Species or Physiognomic Features: No Data Available

Classification Comments: Although the dominant species is shared with the different ~Artemisia tridentata ssp. spiciformis - Artemisia tridentata ssp. vaseyana Steppe & Shrubland Alliance (A3207)$$, Young et al. (2007) show that there are substantial compositional and environmental differences which warrant separation into two different alliances. Much of the area occupied by this widespread alliance has been moderately to severely degraded by livestock grazing and subsequent invasion by Bromus tectorum with a consequent alteration of its fire regime. Individuals of the dominant species do not usually survive fire and usually take a decade or more to reestablish (Tirmenstein 1999c).

Similar NVC Types: No Data Available
note: No Data Available

Physiognomy and Structure: The shrublands in this alliance are characterized by an open to dense (10-75%) cover of microphyllous evergreen shrubs, usually 1-2 m in height. Cespitose graminoids are scattered in the shrub matrix and can occasionally match the shrubs in height. A sparse, emergent layer of scale-leaved or needle-leaved evergreen trees may also be present, but in many areas no trees are present. Forb species may be frequent, but are usually of low cover. With increasing summer rain in the southeastern edge of the range, there is an increase in the importance of sod-forming grasses in comparison to bunchgrasses. In many areas, undisturbed stands in this alliance will have a cryptogamic soil crust composed of a mix of lichens, tiny mosses, and bacteria.

Floristics: The vegetation included in this alliance is characterized by an open to dense (10-70% cover) short-shrub layer in which Artemisia tridentata ssp. vaseyana is codominant, usually with 40-60% relative cover, with non-sagebrush shrub species such as Amelanchier utahensis, Holodiscus dumosus, Purshia tridentata, or Symphoricarpos oreophilus. Perennial graminoids typically dominate the open to moderately dense herbaceous layer. The most widespread species are Pseudoroegneria spicata and Festuca idahoensis, which occur from the Great Basin to the Rocky Mountains, although they may not be the most abundant species in individual stands. Other locally important species may include Bromus carinatus, Elymus trachycaulus, Hesperostipa comata, Koeleria macrantha, Muhlenbergia montana, Poa fendleriana, and Poa secunda. The forb layer is variable and can be very diverse. Species of Castilleja, Potentilla, Erigeron, Phlox, Astragalus, Geum, Lupinus, and Eriogonum are characteristic.

Dynamics:  Complex ecological interactions between fire regimes, grazing history, and climate patterns result in equally complex patterns of species structure and composition in Artemisia tridentata. These present corresponding difficulties in the classification of these shrublands, which have been compounded by the influence of human settlement and agricultural patterns. What follows is a summary of some of the influences of altered fire regimes, and grazing history on Artemisia tridentata shrublands and shrub herbaceous vegetation.

Artemisia tridentata ssp. vaseyana shrublands may represent either drier or more disturbed examples of the Artemisia tridentata ssp. vaseyana shrubland complex. Shrub densities typically increase with overgrazing of the bunchgrass component or with increasing summer drought (West 1983c). There is considerable debate over whether present shrub-dominated stands are actually degraded "steppe" (e.g., shrub-herbaceous physiognomy), and if the stands will return to steppe with changes in grazing and fire management. Artemisia tridentata is inhibited by fire, and excessive grazing may decrease fire frequency due to consumption of herbaceous forage, resulting in increased shrub density. Conversely, invasion by non-native annual grasses (e.g., Bromus tectorum at lower elevations) may increase fire frequency sufficiently to eliminate the shrubs from the stands (Hironaka et al. 1983). With a change in fire frequency, species composition will be altered as well (West 1983c). With a high fire frequency (every 2-5 years), perennial grasses and shrubs are eliminated and non-native annual grasses dominate. At fire-return intervals of 10-30 years, short-lived resprouting shrubs such as Chrysothamnus or Tetradymia spp. dominate. At fire intervals of 30-70 years, a mixture of perennial bunchgrasses and shrubs is maintained. Finally, in the complete absence of fire, deep-rooted shrubs such as Artemisia tridentata become the theoretical dominants.

Environmental Description:  This alliance is widespread in mountainous areas across the western U.S. and forms large, continuous stands on mid-elevation mountain slopes and foothills, and can extend above the lower treeline as patches within montane or subalpine coniferous forests. The climate regime is cool, semi-arid to subhumid, with yearly precipitation ranging from 18-60 cm. Much of the yearly precipitation falls as snow, which may cover the ground for long periods in winter. Temperatures are continental with large annual and diurnal variation. The elevation range for this alliance is large, from about 1060 m in eastern Oregon and Washington, to well over 3500 m in the mountains of northern Nevada, Idaho, and Colorado. Landscape positions are variable as well, but primarily are deep-soiled to stony flats, ridges, nearly flat ridgetops, and mountain slopes. All aspects are represented, but the higher elevation occurrences may be mainly on south- or west-facing slopes. Soils generally are moderately deep to deep, well-drained, and of loam, sandy loam, clay loam, or gravelly loam textural classes; they often have a substantial volume of coarse fragments. The soils are derived from a variety of parent materials (although sandstones, limestones, and crystalline rocks are common). In some cases, soils supporting stands of this alliance are unstable and prone to mass movement (Bramble-Brodahl 1978, Hironaka et al. 1983). In subalpine environments, these shrublands are found on deeper soils than Artemisia arbuscula subalpine shrublands. Although the vegetation may grow in alkaline soils at the edge of internally drained basins, Artemisia tridentata is a non-halophyte and requires low salinity for optimum growth.

Geographic Range: This shrubland alliance occurs in mountainous regions from eastern California, Oregon, and Washington, across the Great Basin in Nevada, the northern Rocky Mountain foothills of Idaho, and in Colorado, Wyoming, and Montana. It has not been reported from Utah, Arizona, or New Mexico, but it is very likely to occur in these states at high elevations. In addition, the alliance probably extends north into Alberta, Canada.

Nations: CA?,US

States/Provinces:  AB?, AZ, CA, CO, ID, MT, NV, OR, UT, WY




Confidence Level: Low

Confidence Level Comments: No Data Available

Grank: GNR

Greasons: No Data Available

Type Name Database Code Classification Code
Class 3 Desert & Semi-Desert Class C03 3
Subclass 3.B Cool Semi-Desert Scrub & Grassland Subclass S11 3.B
Formation 3.B.1 Cool Semi-Desert Scrub & Grassland Formation F033 3.B.1
Division 3.B.1.Ne Western North American Cool Semi-Desert Scrub & Grassland Division D040 3.B.1.Ne
Macrogroup 3.B.1.Ne.3 Big Sagebrush - Threetip Sagebrush - Antelope Bitterbrush Steppe & Shrubland Macrogroup M169 3.B.1.Ne.3
Group 3.B.1.Ne.3.c Spiked Big Sagebrush - Mountain Big Sagebrush - Mountain Silver Sagebrush Steppe & Shrubland Group G304 3.B.1.Ne.3.c
Alliance A3208 Mountain Big Sagebrush - Mixed Steppe & Shrubland Alliance A3208 3.B.1.Ne.3.c
Association CEGL001003 Big Sagebrush - Joint-fir species Shrubland CEGL001003 3.B.1.Ne.3.c
Association CEGL001032 Mountain Big Sagebrush - Antelope Bitterbrush / Bluebunch Wheatgrass Shrubland CEGL001032 3.B.1.Ne.3.c
Association CEGL001034 Mountain Big Sagebrush - Mountain Snowberry / Slender Wheatgrass Shrubland CEGL001034 3.B.1.Ne.3.c
Association CEGL001035 Mountain Big Sagebrush - Mountain Snowberry / California Brome Shrubland CEGL001035 3.B.1.Ne.3.c
Association CEGL001036 Mountain Big Sagebrush - Mountain Snowberry / Idaho Fescue Shrubland CEGL001036 3.B.1.Ne.3.c
Association CEGL001037 Mountain Big Sagebrush - Mountain Snowberry / Sandberg Bluegrass Shrubland CEGL001037 3.B.1.Ne.3.c
Association CEGL001038 Mountain Big Sagebrush - Mountain Snowberry / Bluebunch Wheatgrass Shrubland CEGL001038 3.B.1.Ne.3.c
Association CEGL001039 Mountain Big Sagebrush - Mountain Snowberry / Needle-and-Thread Shrubland CEGL001039 3.B.1.Ne.3.c
Association CEGL002807 Mountain Big Sagebrush - Rockspirea Shrubland CEGL002807 3.B.1.Ne.3.c
Association CEGL002820 Utah Serviceberry - (Mountain Big Sagebrush, Wyoming Big Sagebrush) Shrubland CEGL002820 3.B.1.Ne.3.c
Association CEGL002951 Mountain Snowberry Shrubland CEGL002951 3.B.1.Ne.3.c
Association CEGL005474 Mountain Big Sagebrush - Desert-sweet / Scabland Penstemon Shrubland CEGL005474 3.B.1.Ne.3.c
Association CEGL005484 Desert-sweet / Scabland Penstemon Sparse Vegetation CEGL005484 3.B.1.Ne.3.c
Association CEGL005827 Mountain Big Sagebrush - (Antelope Bitterbrush) / Mountain Muhly - (Needle-and-Thread) Shrubland CEGL005827 3.B.1.Ne.3.c

Concept Lineage: This alliance includes stands where Artemisia tridentata ssp. vaseyana is codominant with non-sagebrush shrub species and is similar to proto-alliance Artemisia tridentata ssp. vaseyana - Mixed Shrubland Alliance (A2040) and to Old Alliance III.A.4.N.a. Artemisia tridentata ssp. vaseyana Shrubland Alliance (A.831) (Artemisia tridentata ssp. vaseyana shrub codominated stands)

Predecessors: No Data Available

Obsolete Names: No Data Available

Obsolete Parents: No Data Available

Synonomy: < Artemisia tridentata ssp. vaseyana (Mountain big sagebrush) Alliance (Sawyer et al. 2009) [35.111.00]
= Artemisia tridentata ssp. vaseyana Shrubland Alliance (Evens et al. 2014) [probably equivalent]
>< Big Sagebrush Scrub (#35210) (Holland 1986b)
>< Big Sagebrush Series (Sawyer and Keeler-Wolf 1995)
< SRM Cover Type #402 - Mountain Big Sagebrush (Shiflet 1994)
>< Western Shrub and Grasslands Combinations: 55: Sagebrush-Steppe (Artemisia-Agropyron) (Küchler 1964)
>< Western Shrub: 38: Great Basin Sagebrush (Artemisia) (Küchler 1964)

Concept Author(s): K.A. Schulz, in Faber-Langendoen et al. (2013)

Author of Description: K.A. Schulz

Acknowledgements: We have incorporated significant descriptive information previously compiled by D. Sarr.

Version Date: 12-18-14

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