Print Report

M511 North Pacific Coastal Ruderal Grassland & Shrubland Macrogroup

Type Concept Sentence: This macrogroup includes coastal sand dunes dominated by non-native beachgrasses Ammophila arenaria, Ammophila breviligulata, forbs and succulents such as Cakile edentula, Cakile maritima, Carpobrotus spp., Mesembryanthemum spp., and/or introduced shrubs such as Lupinus arboreus. It occurs along the coast of California north into British Columbia.


Common (Translated Scientific) Name: North Pacific Coastal Ruderal Grassland & Shrubland Macrogroup

Colloquial Name: North Pacific Coastal Ruderal Grassland & Shrubland

Hierarchy Level:  Macrogroup

Type Concept: This macrogroup occurs on coastal dunes and other sandy areas at sea level from California into British Columbia. Substrates are eolian sands without horizon development or accumulated organic material. These areas are dominated by non-native beachgrasses Ammophila arenaria, Ammophila breviligulata, and/or forbs and succulents such as Cakile edentula, Cakile maritima, Carpobrotus spp., and Mesembryanthemum spp. Dunes may also have introduced shrubs such as Lupinus arboreus.

Diagnostic Characteristics: No Data Available

Rationale for Nominal Species or Physiognomic Features: No Data Available

Classification Comments: Cakile spp. are more typical of beaches, and perhaps should not be included here. Pickart and Barbour (2007) state that naturalized (what we term ruderal) vegetation now exceeds native vegetation for most of California''s dune systems. In Oregon, this type includes restoration areas of planted Leymus mollis ssp. mollis that have spread beyond where the natural type would have occurred.

Similar NVC Types: No Data Available
note: No Data Available

Physiognomy and Structure: These areas can be dense shrublands totally comprised of one species or they can be more open mosaics of grassland and shrubland, or pure grassland.

Floristics: This macrogroup is dominated by non-native or introduced species such as Ammophila arenaria or Ammophila breviligulata, Cakile edentula, Cakile maritima, Carpobrotus chilensis, Carpobrotus edulis, Mesembryanthemum spp., and/or Lupinus arboreus. Other non-native species may also be present, including Bromus diandrus, Cirsium vulgare, Medicago polymorpha, Poa pratensis, and Rumex acetosella. Native species may be scattered about but these are at very low cover. Some restoration areas of planted Leymus mollis ssp. mollis have spread beyond where the natural type would have occurred, and are thus more invasive than "restored" and fit this concept, although this may be difficult to determine in the field and makes knowledge of site history imperative.

Dynamics:  According to Pickart and Barbour (2007, and references therein), "invasion by Carpobrotus edulis results in displacement of native herbaceous and even shrub species by direct overgrowth or indirectly by competition for resources…ecosystem impacts include changes to soil pH...the buildup of organic matter, and loss of sand movement...High Carpobrotus edulis cover is detrimental to burrowing dune insects...." They also note that Ammophila arenaria, which was introduced to stabilize the dunes, "has built a steep, continuous foredune over much of its range, replacing the original low, hummocky and mobile foredunes."

Environmental Description:  This macrogroup occupies coastal dunes and other sandy areas at sea level. Substrates are entirely eolian sands without horizon development or accumulated organic material. These dunes tend to be higher and more stabilized and therefore more abundant than historically due to the introduction of non-native dunegrass. A striking pair of photographs illustrate this point in Pickart and Barbour (2007, Figure 6.11).

Geographic Range: This macrogroup ranges from California into British Columbia, Canada, and possibly southeastern Alaska. It may reach south into Mexico. No data or descriptive information are available from Mexico or Alaska.

Nations: CA,MX?,US

States/Provinces:  BC, CA, OR, WA




Confidence Level: Moderate

Confidence Level Comments: No Data Available

Grank: GNA

Greasons: No Data Available


Concept Lineage: No Data Available

Predecessors: No Data Available

Obsolete Names: No Data Available

Obsolete Parents: No Data Available

Synonomy: No Data Available

Concept Author(s): M.G. Barbour and J. Major (1977)

Author of Description: G. Kittel

Acknowledgements: No Data Available

Version Date: 03-29-17

  • Barbour, M. G., and J. Major, editors. 1977. Terrestrial vegetation of California. John Wiley and Sons, New York. 1002 pp.
  • Faber-Langendoen, D., J. Drake, S. Gawler, M. Hall, C. Josse, G. Kittel, S. Menard, C. Nordman, M. Pyne, M. Reid, L. Sneddon, K. Schulz, J. Teague, M. Russo, K. Snow, and P. Comer, editors. 2010-2019a. Divisions, Macrogroups and Groups for the Revised U.S. National Vegetation Classification. NatureServe, Arlington, VA. plus appendices. [in preparation]
  • Pickart, A., and M. Barbour 2007. Beach and dune. Chapter 6, pages 155-179 in: M. G. Barbour, M. G., T. Keeler-Wolf, and A. A. Schoenherr, editors. Terrestrial vegetation of California. Third edition. University of California Press, Berkeley.
  • Wiedemann, A. M. 1966. Contributions to the plant ecology of the Oregon coastal sand dunes. Ph.D. dissertation, Oregon State University, Corvallis. 255 pp.