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CEGL007054 Nyssa biflora - (Taxodium distichum) / Clethra alnifolia - Viburnum nudum / Woodwardia areolata Floodplain Forest
Type Concept Sentence: No Data Available
Common (Translated Scientific) Name: Swamp Tupelo - (Bald-cypress) / Coastal Sweet-pepperbush - Possumhaw / Netted Chainfern Floodplain Forest
Colloquial Name: Central Atlantic Blackwater Floodplain Forest
Hierarchy Level: Association
Type Concept: This association includes seasonally flooded forests in acidic Coastal Plain bottomlands of southeastern and east-central Virginia, the Maryland Eastern Shore, and likely Delaware. It occurs along headwater swamps, and occasionally larger river backswamps, in sandy, nutrient-poor portions of the Coastal Plain. Soils are often organic or contain high organic matter content. Forest vegetation is generally dominated by Nyssa biflora, often with associated hardwoods such as Acer rubrum, Liquidambar styraciflua, Quercus laurifolia, Quercus nigra, and Magnolia virginiana. Although absent from many stands, Taxodium distichum is occasionally an important overstory associate. The understory is characterized by sparse to often dense cover of acidophilic, ericaceous and non-ericaceous shrubs such as Eubotrys racemosa, Vaccinium formosum, Vaccinium fuscatum, Clethra alnifolia, Viburnum nudum, Itea virginica, Ilex verticillata, and Arundinaria tecta. The herb layer varies from sparse to well-developed, with characteristic species such as Woodwardia areolata, Saururus cernuus, and others.
Diagnostic Characteristics: No Data Available
Rationale for Nominal Species or Physiognomic Features: No Data Available
Classification Comments: Classification of this type is based primarily on quantitative analysis of a regional dataset of bottomland plots collected across the entire Virginia Coastal Plain. The group of 19 plots representing this type was also analyzed with the addition of one Maryland plot, and compared analytically against a group of 12 plots representing the analogous but more southern blackwater swamp forest of North Carolina and South Carolina, ~Taxodium distichum - Nyssa biflora / Fraxinus caroliniana / Lyonia lucida Floodplain Forest (CEGL004733)$$ (M. Schafale pers. comm.). In the latter analysis, the two types showed a convincing separation, and a number of differential species were evident.
Similar NVC Types: No Data Available
note: No Data Available
Physiognomy and Structure: No Data Available
Floristics: Vegetation is a closed-canopy forest usually dominated by Nyssa biflora, but often with significant associates of Liquidambar styraciflua and Acer rubrum. Although absent from many stands, Taxodium distichum is occasionally important or codominant in the canopy. Other inconstant overstory trees that can be important in some stands include Fraxinus pennsylvanica, Quercus laurifolia, Quercus nigra, and Quercus phellos. Nyssa biflora, Acer rubrum, Ilex opaca var. opaca, and Magnolia virginiana are the most typical subcanopy trees. The shrub layer varies from sparse to more often dense and consists of acidophilic, ericaceous and non-ericaceous species such as Eubotrys racemosa (= Leucothoe racemosa), Vaccinium formosum, Vaccinium fuscatum, Rhododendron viscosum, Clethra alnifolia, Viburnum nudum, Itea virginica, Ilex verticillata, and Arundinaria tecta (= Arundinaria gigantea ssp. tecta). Smilax rotundifolia and Smilax walteri are common scrambling vines. The herb layer varies from sparse to well-developed. The most constant herbs in 20 Virginia and Maryland plots are Woodwardia areolata, Mitchella repens, Saururus cernuus, Carex seorsa, Osmunda cinnamomea, Triadenum walteri, Boehmeria cylindrica, and Carex crinita. Less constant herbs that attain moderately high cover in some stands include Osmunda regalis var. spectabilis, Carex intumescens, Eutrochium dubium (= Eupatorium dubium), Carex lonchocarpa, and Bidens discoidea. Mean species richness was 38 taxa per 400 m2. Stands on the Delmarva Peninsula of Virginia and Maryland lack some of the southern species (e.g., Arundinaria tecta, Quercus laurifolia) that are frequent in southeastern Virginia. In Tiner and Burke''s (1995) study of Maryland wetlands, this community was described as the non-tidal swamp blackgum dominance type, with associated trees of Acer rubrum, Magnolia virginiana, Fraxinus pennsylvanica, and Liquidambar styraciflua; characteristic shrubs and vines of Clethra alnifolia, Eubotrys racemosa, Viburnum nudum, and Smilax rotundifolia; and characteristic herbs of Carex intumescens, Boehmeria cylindrica, Woodwardia areolata, Glyceria striata, Bidens sp., Lycopus sp., Carex folliculata, Cinna arundinacea, Saururus cernuus, and Eutrochium dubium. More recent field investigations in Maryland have found Ilex opaca var. opaca, Smilax laurifolia, and Phoradendron leucarpum (= Phoradendron serotinum) to be frequent or common in the type.
Dynamics: The streams along which this community type occurs carry limited clay and silt sediments. Consequently, nutrient inputs by flooding are limited. Tree growth appears to be notably slow in the wet, infertile soils and only very old trees attain large size.
Environmental Description: This association occurs in seasonally flooded, acidic bottomlands of the central Atlantic Coastal Plain. It occurs along headwater swamps, and occasionally larger river backswamps, in nutrient-poor, usually sandy portions of the region. In Virginia, it often occurs along small streams and river tributaries through inner Coastal Plain sandhills. Soils are often organic or contain high organic matter content. Sites typically have complex microtopography, with numerous hummocks, hollows, and braided channels. Occasional sites in larger-river sloughs may receive acidic groundwater inputs from the base of adjoining slopes, in addition to overland flooding. Among 19 Virginia plots, hummocks and hollows averaged about 45% and 55%, respectively, of the ground surface. Soil samples collected from the A-horizon (if not entirely organic) had a mean organic matter content of 15%, >50% sand content, and a mean pH of 3.9, with low calcium, magnesium, and total base saturation, and high iron and aluminum.
Geographic Range: This community type is frequent in scattered patches throughout southeastern and east-central Virginia Coastal Plain (Meherrin, Nottoway, Blackwater, Chickahominy, and Piankatank river drainages). It is also known from the Pocomoke River drainage on the Maryland Eastern Shore. Similar vegetation has been reported from Delaware (W. McAvoy pers. comm. 2011), but details are currently lacking. Additional occurrences are likely throughout this region.
Nations: US
States/Provinces: DE?, MD, VA
Plot Analysis Summary:
http://vegbank.org/natureserve/ELEMENT_GLOBAL.2.868705
Confidence Level: Low - Poorly Documented
Confidence Level Comments: No Data Available
Grank: G3
Greasons: No Data Available
Type | Name | Database Code | Classification Code |
---|---|---|---|
Class | 1 Forest & Woodland Class | C01 | 1 |
Subclass | 1.B Temperate & Boreal Forest & Woodland Subclass | S15 | 1.B |
Formation | 1.B.3 Temperate Flooded & Swamp Forest Formation | F026 | 1.B.3 |
Division | 1.B.3.Nb Southeastern North American Flooded & Swamp Forest Division | D062 | 1.B.3.Nb |
Macrogroup | 1.B.3.Nb.4 Swamp Chestnut Oak - Bald-cypress - Pecan Southern Floodplain Forest Macrogroup | M031 | 1.B.3.Nb.4 |
Group | 1.B.3.Nb.4.c Bald-cypress - Water Tupelo Floodplain Forest Group | G033 | 1.B.3.Nb.4.c |
Alliance | A0323 Swamp Tupelo - Water Tupelo Floodplain Forest Alliance | A0323 | 1.B.3.Nb.4.c |
Association | CEGL007054 Swamp Tupelo - (Bald-cypress) / Coastal Sweet-pepperbush - Possumhaw / Netted Chainfern Floodplain Forest | CEGL007054 | 1.B.3.Nb.4.c |
Concept Lineage: CEGL007743 merged partly into CEGL004651 and partly into CEGL007054.
Predecessors: No Data Available
Obsolete Names: No Data Available
Obsolete Parents: No Data Available
Synonomy: ? Swamp black gum dominance type (Tiner and Burke 1995)
- Eastern Ecology Working Group of NatureServe. No date. International Ecological Classification Standard: International Vegetation Classification. Terrestrial Vegetation. NatureServe, Boston, MA.
- Fleming, G. P., K. D. Patterson, and K. Taverna. 2017. The natural communities of Virginia: A classification of ecological community groups and community types. Third approximation. Version 3.0. Virginia Department of Conservation and Recreation, Division of Natural Heritage, Richmond, VA. [http://www.dcr.virginia.gov/natural-heritage/natural-communities/]
- Harrison, J. W. 2011. The natural communities of Maryland: 2011 working list of ecological community groups and community types. Unpublished report. Maryland Department of Natural Resources, Wildlife and Heritage Service, Natural Heritage Program, Annapolis. 33 pp.
- McAvoy, William. Personal communication. Botanist, Delaware Natural Heritage Program, Delaware Division of Fish & Wildlife, Department of Natural Resources and Environmental Control, Smyrna, DE.
- Schafale, Mike P. Personal communication. Ecologist, North Carolina Department of Environment, Health, and Natural Resources, Division of Parks and Recreation, Natural Heritage Program, Raleigh.
- Tiner, R. W., and D. G. Burke. 1995. Wetlands of Maryland. USDI Fish & Wildlife Service, Ecological Services, Region 5, Hadley, MA, and Maryland Department of Natural Resources, Annapolis, MD. Cooperative publication. 193 pp. plus appendices. [http://library.fws.gov/Wetlands/MD_wetlands85.pdf]