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G033 Taxodium distichum - Nyssa aquatica Floodplain Forest Group

Type Concept Sentence: Dominant and characteristic trees are Taxodium distichum (occasionally Taxodium ascendens) and Nyssa aquatica (and occasionally Nyssa biflora). This vegetation is primarily found in the Atlantic and Gulf coastal plains, the adjacent Piedmont and interior regions, and the Mississippi River Alluvial Plain and adjacent Upper East Gulf Coastal Plain, and west into the coastal plains of Louisiana and Texas.


Common (Translated Scientific) Name: Bald-cypress - Water Tupelo Floodplain Forest Group

Colloquial Name: Bald-cypress - Tupelo Floodplain Forest

Hierarchy Level:  Group

Type Concept: This wetland forest group is primarily affiliated with the Atlantic and Gulf coastal plains from southeastern Virginia (rarely north into the Chesapeake Bay) to Texas, portions of the adjacent Piedmont and interior regions, and the Mississippi River Alluvial Plain and adjacent Upper East Gulf Coastal Plain from southern Illinois, western Kentucky and southeastern Missouri, south to the Gulf of Mexico, and west in the coastal plains of Louisiana and Texas. Dominant and characteristic trees are Taxodium distichum (and occasionally Taxodium ascendens) and Nyssa aquatica (and occasionally Nyssa biflora). These are generally known as "Bald-Cypress - Tupelo Forests." Both Taxodium distichum and Nyssa aquatica are primarily, if not entirely, found in the coastal plains. Their ranges are not entirely congruent, with Nyssa aquatica being more limited. So, vegetation may be dominated by either or both of these characteristic species (and occasionally including Taxodium ascendens and/or Nyssa biflora as noted above). Also included here are embedded patches of smaller trees, including Forestiera acuminata, Itea virginica, and Planera aquatica, as well as Cephalanthus occidentalis, which can range in lifeform from small tree to shrub. These are usually smaller-patch features embedded within larger-patch (and taller stature) bald-cypress - tupelo forests. The basis of this group is floristic rather than purely hydrologic. This group primarily encompasses communities of streams and rivers of all orders and sizes, including some (or most) tidal forests. Stands are typically flooded for periods of up to 3 months, but not during the growing season. Stands are mainly from low bottomlands, depressions, sloughs and abandoned channel segments, and are not typically affiliated with elevated features such as riverfronts, point bars, natural levees, high bottomlands, levees, ridges, and upper terraces. This group includes forests known as "blackwater" as well as "brownwater" examples, as well as several tidally flooded associations. Isolated wetlands such as depression ponds dominated by Taxodium are excluded.

Diagnostic Characteristics: These are riverine forests (and more rarely woodlands) dominated by a combination of broad-leaved deciduous trees and a needle-leaved deciduous tree (Taxodium). Examples may be tidally or non-tidally flooded, in "blackwater" and "brownwater" situations. Characteristic trees include Acer rubrum var. drummondii, Carya aquatica, Fraxinus profunda, Gleditsia aquatica, Nyssa aquatica, Nyssa biflora, Planera aquatica, and Taxodium distichum. Some shrubs include Cephalanthus occidentalis, Decodon verticillatus, Forestiera acuminata, Itea virginica, and Planera aquatica. Characteristic forbs and graminoids include Boehmeria cylindrica, Carex decomposita, Carex grayi, Carex intumescens, Carex joorii, Carex lupulina, other Carex spp., and Saururus cernuus. Aquatic and floating forbs include Ceratophyllum spp., Elodea spp., Lemna minor, Ludwigia peploides, Potamogeton spp., and Sagittaria lancifolia.

Rationale for Nominal Species or Physiognomic Features: No Data Available

Classification Comments: This group includes associations and alliances dominated by Taxodium distichum, Taxodium ascendens, and Nyssa aquatica, typically in riverine settings. These are generally known as "Bald-Cypress - Tupelo Forests"; this is in contrast to bottomland hardwood forests (oak-sweetgum) which are accommodated in a different group. Both Taxodium distichum and Nyssa aquatica are primarily, if not entirely, found in the coastal plains. Their ranges are not entirely congruent, with Nyssa aquatica being more limited. So, associations accommodated here may be dominated by either or both of these characteristic species (and occasionally including Taxodium ascendens and/or Nyssa biflora as noted above). These are associations that are generally of a longer hydroperiod than those in ~Oak - Sweetgum Floodplain Forest Group (G034)$$, but some in that group are of a long hydroperiod as well. Also included here are some embedded patches of smaller trees including Cornus foemina, Forestiera acuminata, Itea virginica, and Planera aquatica, as well as Cephalanthus occidentalis, which can range in lifeform from small tree to shrub. These are usually smaller-patch features imbedded within larger-patch (and taller stature) bald-cypress - tupelo forests. The basis of the distinction between these two groups is floristic and broadly ecological rather than purely hydrologic. This group primarily encompasses communities of streams and rivers of all orders and sizes, including some (or most) tidal forests. Isolated wetlands such as depression ponds dominated by Taxodium are placed in nonriverine groups.

Similar NVC Types: No Data Available
note: No Data Available

Physiognomy and Structure: These are flooded riverine forests composed of tall trees, mostly broad-leaved deciduous trees and one needle-leaved deciduous tree (Taxodium).

Floristics: Stands of this group are primarily tall-statured forests dominated by Taxodium distichum (and occasionally Taxodium ascendens) and/or Nyssa aquatica (and occasionally Nyssa biflora). The ranges of Taxodium distichum and Nyssa aquatica are not entirely congruent, with Nyssa aquatica being more limited. Therefore, in some parts of the range of the group, one of these characteristic species may not be present. Some other trees which may be associated with stands of this group include Acer rubrum var. drummondii, Carya aquatica, Fraxinus pennsylvanica, Gleditsia aquatica, Salix nigra, and other trees tolerant of flooding. Associations dominated by Quercus spp. are included in a different group. Also included here are some associations dominated by smaller trees, including Forestiera acuminata, Itea virginica, and Planera aquatica, as well as Cephalanthus occidentalis, which can range in lifeform from small tree to shrub. Cornus foemina and Decodon verticillatus may also be present. Terrestrial herbs are usually limited due to long hydroperiods, and are often only found on elevated logs and stumps, but can include Boehmeria cylindrica, Carex decomposita, Carex grayi, Carex intumescens, Carex joorii, Carex lupulina, and Saururus cernuus. Aquatic and floating herbs include Ceratophyllum spp., Elodea spp., Lemna minor, Ludwigia peploides, Potamogeton spp., and Sagittaria lancifolia.

Dynamics:  Flooding is an important ecological factor in examples of this group, and the length of the flooding may be the most important factor separating this vegetation from that of other wetland groups. In addition to disturbance, floods bring nutrient input, deposit sediment, exclude non-flood-tolerant species and disperse plant seeds. Flooding is most common in the winter but may occur in other seasons. The small flows, low gradient, and binding of sediment by vegetation limit channel shifts and sediment movement, but floods may cause local disturbance by scouring. Flood waters may have significant energy in higher-gradient systems, but scouring and reworking of sediment rarely affect more than small patches. Most of these forests would exist naturally as multi-aged old-growth forests driven by gap-phase regeneration. Windthrow is probably the most important cause of gaps, as wind disturbance is perhaps more important than in uplands because of frequently wet soils, less dense soils, and more shallow-rooted trees.

Fire does not appear to be a dominant factor, and most floodplain vegetation in this group is not very flammable. Fire is probably more important in small stream examples than in larger river ones, because distances to uplands are short and stream channels and sloughs are smaller and less effective as firebreaks. Stands of this group often have limited herbaceous layers, making fire less of an important factor, even during extended dry periods.

Some examples are affected by diurnal tidal flooding or by storm tides. In these cases, infrequent intrusion of saltier water, which is stressful or fatal to many of the plant species, is an important periodic disturbance created by storms. Natural fire is not frequent in these tidal examples, but may sometimes be important in determining the boundary between tidal swamps and tidal marshes. The tidal forest examples generally appear to be in a shifting relationship with adjacent tidal freshwater marshes. Many of these tidal marshes have standing dead trees in them, suggesting they recently were swamps. But some marshes are being invaded with trees and may be turning into swamps. Rising sea level is causing more inland swamps to develop tidal characteristics and causing some stands to turn into marshes. In areas not too strongly affected by salt intrusion, drowning by rising sea level, or fire, the communities can be expected to exist as old-growth, multi-aged forests.

The associations in this group are generally of a longer hydroperiod than those in ~Oak - Sweetgum Floodplain Forest Group (G034)$$, but some in that group are of a long hydroperiod as well. Stands of this group, because of their longer hydroperiod, have been subject to more indirect disturbance effects than others, but they are still subject to a variety of indirect modern human influences. Many larger rivers have been dammed, and power generation and regulation of waterflow create unnatural flood regimes. Extensive erosion of uplands, caused by many years of poor agricultural practices, transported large amounts of sediment into floodplains. River bottoms were the focus of agriculture among Native Americans, so some members of this group have a long history of human clearing. A number of exotic plant species have invaded floodplains as well. Isolated wetlands such as depression ponds dominated by Taxodium are for the time being placed in nonriverine groups.

Environmental Description:  Stands of this group are found on streams and rivers of all orders and sizes, including some (or most) tidal forests. The elements included are mainly from low bottomlands, depressions, sloughs and abandoned channel segments, and are not typically affiliated with elevated features such as riverfronts, point bars, natural levees, high bottomlands, levees, ridges, and upper terraces. Wetness is a major influence, especially within channels and where water is ponded in local depressions. Tidally influenced examples occur in lower reaches of river floodplains and along estuary shorelines, in places regularly or irregularly flooded by lunar or wind tides. The water has little salt content, due to distance from the ocean and/or strong freshwater input. Soils may be mineral or organic. Soils are generally permanently saturated even when the tide is low. The transition of the hydrology to flood dominance rather than tidal dominance may be very gradual.

Geographic Range: This wetland forest group is primarily affiliated with the Atlantic and Gulf coastal plains from southeastern Virginia (rarely north into the Chesapeake Bay) to Texas, portions of the adjacent Piedmont and interior regions, and the Mississippi River Alluvial Plain and adjacent Upper East Gulf Coastal Plain from southern Illinois and Indiana, western Kentucky (including areas along the Ohio River (EPA Ecoregion 72a)) and southeastern Missouri, south to the Gulf of Mexico, and west to the coastal plains of Louisiana and Texas. There is also one association in the Ouachitas of Arkansas and Oklahoma.

Nations: US

States/Provinces:  AL, AR, DE, FL, GA, IL, IN, KY, LA, MD, MO, MS, NC, NY, OK, SC, TN, TX, VA




Confidence Level: Moderate

Confidence Level Comments: No Data Available

Grank: GNR

Greasons: No Data Available


Concept Lineage: No Data Available

Predecessors: No Data Available

Obsolete Names: No Data Available

Obsolete Parents: No Data Available

Synonomy: >< Bottomland Forest (FNAI 1990)
>< Floodplain Forest (FNAI 1990)
> Floodplain Hardwood Forest (Marks and Harcombe 1981)
>< Floodplain Swamp (FNAI 1990)
> Swamp Cypress Tupelo Forest (Marks and Harcombe 1981)

Concept Author(s): R.R. Sharitz and W.J. Mitsch (1993)

Author of Description: M. Pyne

Acknowledgements: No Data Available

Version Date: 05-13-15

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