Print Report

M402 Leymus mollis - Honckenya peploides - Lupinus nootkatensis Coastal Shore Macrogroup

Type Concept Sentence: This type is found on North American Arctic coastline beaches, beach dunes, stabilized sand or cobble deposits, on sea cliffs, rocky headlands, and cobble beaches. Vegetation consists of mostly grasslands, dwarf-shrublands, stunted trees, and other herbaceous species. Some of the common dominant species include Leymus mollis, Honckenya peploides, and/or Lupinus nootkatensis.


Common (Translated Scientific) Name: American Dunegrass - Seaside Sandplant - Nootka Lupine Coastal Shore Macrogroup

Colloquial Name: North American Arctic Coastal Shore

Hierarchy Level:  Macrogroup

Type Concept: This type occurs along the North American Arctic coastline and consists of stabilized coastal beaches and beach dunes. It also occurs on Arctic coastline sea cliffs, rocky headlands, and cobble beaches. Patch size is small to moderate and often linear. Two different physiognomic structures are found in the group: grasslands and dwarf-shrublands, as well as areas of bare sand or cobbles. Vegetation typically includes herbaceous species with varying degrees of tolerance for salt spray and wind abrasion. Salt-tolerant forb communities occur just above mean high tide and are dominated or codominated by Cochlearia groenlandica, Achillea millefolium var. borealis, Honckenya peploides, and/or Mertensia maritima. As dune height and distance from the ocean increase, sites are dominated by Leymus mollis communities that may include near-monocultures of Leymus mollis to more species-rich associations including Leymus mollis, Lathyrus japonicus var. maritimus, and Poa eminens. Older dunes support dwarf-shrubs (primarily Empetrum nigrum) mixed with herbaceous species which often grow in narrow stringers on the older beach ridges behind the Leymus mollis zone. On rocky cliff and cobble shores, a variety of species may occur depending on the level of salt exposure, steepness, aspect, and available microsites. Shrubs such as Alnus viridis ssp. sinuata or Rubus spectabilis may be present but usually account for less than 5% of the total vascular plant cover. Herbaceous cover is diverse and may include many of the following species: Aruncus dioicus var. acuminatus, Campanula spp., Carex macrochaeta, Chamerion latifolium, Deschampsia spp., Heuchera glabra, Lupinus nootkatensis, Phegopteris connectilis, Potentilla villosa, Prenanthes alata, and Rhodiola rosea. Picea sitchensis may also occupy these rocky headlands, and is characterized by somewhat stunted growth, usually with branches from top to bottom of bole. Epiphytic lichens are abundant in this macrogroup. Beaches are dry to mesic and typically sandy. Cobble beaches are associated with cliff and bluff systems or coarse-textured glacial deposits (i.e., coastal moraines). Beaches are often steep and feature distinct storm berms. These are typically high-energy environments exposed to wave action, wind, salt spray and storm swell. Cobble beaches may have a mixture of silts and sands below the surface (particularly in outwash plains), but the fine material is buried and not subjected to wind and water transport. Forbs, grasses, shrubs, and stunted trees establish on ledges and in cracks.

Diagnostic Characteristics: This type is characterized by grasslands and dwarf-shrublands which occupy beaches and dunes. Leymus mollis is the most consistent species, but salt-tolerant forbs are also present. Diagnostic species of sea cliffs, rocky headlands, and cobble beaches of the Arctic region needs further review.

Rationale for Nominal Species or Physiognomic Features: No Data Available

Classification Comments: Storm surge coastal vegetation may require inclusion in this type or perhaps a new type with salt spray influence. There are no associations currently described for coastal cliff vegetation. Type description is largely based on information from Alaska, and rangewide input is needed.

Similar NVC Types: No Data Available
note: No Data Available

Physiognomy and Structure: Beach vegetation is characterized by graminoid- or dwarf evergreen shrub-dominated vegetation, and cliffs are defined by a highly variable mixture of forbs, grasses, shrubs, and stunted trees.

Floristics: Sandy or cobble beaches contain salt-tolerant forb communities that occur just above mean high tide and are dominated or codominated by Achillea millefolium var. borealis, Cochlearia groenlandica, Honckenya peploides, and/or Mertensia maritima. As dune height and distance from the ocean increase, sites are dominated by Leymus mollis communities that may include near-monocultures of Leymus mollis to more species-rich associations including Leymus mollis, Lathyrus japonicus var. maritimus (= Lathyrus maritimus), and Poa eminens. Older dunes support dwarf-shrubs (primarily Empetrum nigrum) mixed with herbaceous species which often grow in narrow stringers on the older beach ridges behind the Leymus mollis zone. In Alaska, Lathyrus japonicus var. maritimus, Conioselinum chinense, and Cnidium cnidiifolium are uncommon east of Cape Lisburne.

Cliffs and rocky headlands are dominated by forbs, grasses, shrubs, and stunted trees that establish on ledges and in cracks. A variety of species may occur depending on the level of salt exposure, steepness, aspect, and available microsites. Shrubs such as Alnus viridis ssp. sinuata or Rubus spectabilis may be present but usually account for less than 5% of the total vascular plant cover. Herbaceous cover is diverse and may include many of the following species: Aruncus dioicus var. acuminatus, Campanula spp., Carex macrochaeta, Chamerion latifolium, Deschampsia spp., Heuchera glabra, Lupinus nootkatensis, Phegopteris connectilis, Potentilla villosa, Prenanthes alata, and Rhodiola rosea (Boggs et al. 2008b). Picea sitchensis may also occupy these rocky headlands and often does. They are characterized by somewhat stunted growth, usually with branches from top to bottom of bole. Epiphytic lichens are abundant in this macrogroup.

Dynamics:  The primary disturbance is exposure to high wind, storm swells and salt spray.

Environmental Description:  Coastal beaches, beach dunes, and vegetation have stabilized sand or cobble deposits. Soils are dry to mesic and typically sandy. Sites are above the high tide line but still experience storm surges, high winds and salt spray. Sea cliffs, rocky headlands, and cobble beaches occur commonly along the Arctic coastline. Cobble beaches are associated with cliff and bluff systems or coarse-textured glacial deposits (i.e., coastal moraines). Beaches are often steep and feature distinct storm berms. These are typically high-energy environments exposed to wave action and storm swell. Cobble beaches may have a mixture of silts and sands below the surface (particularly in outwash plains), but the fine material is buried and not subjected to wind and water transport. Beach meadows may occupy well-drained stable portions of the upper beach. Permafrost is present at depth along the Arctic Ocean mainland coast and remnant barrier islands but becomes discontinuous along the Bering Sea coast and is absent from constructed barrier islands. Among the barrier islands, vegetated dunes are more common on the remnant barrier islands, which represent relict coastline, than they are on constructed types, which are highly dynamic systems representing comparatively recent depositions of sediment.

Geographic Range: This type occurs along North America''s Arctic coastline, from the Bristol Bay lowlands in southwestern Alaska to the North Slope on the Arctic Ocean, and northwestern Canada.

Nations: CA,GL,RU?,US

States/Provinces:  AK, NT, NU, YT




Confidence Level: Low

Confidence Level Comments: No Data Available

Grank: GNR

Greasons: No Data Available


Concept Lineage: No Data Available

Predecessors: No Data Available

Obsolete Names: No Data Available

Obsolete Parents: No Data Available

Synonomy: > III.A.1.a - Elymus (Viereck et al. 1992)
> III.B.2.d - Halophytic herb wet meadow (Viereck et al. 1992)

Concept Author(s): Faber-Langendoen et al. (2015)

Author of Description: K. Boggs, M.E. Hall, G. Kittel, D. Faber-Langendoen

Acknowledgements: No Data Available

Version Date: 03-29-17

  • Boggs, K., S. C. Klein, L. Flagstad, T. Boucher, J. Grunblatt, and B. Koltun. 2008b. Landcover classes, ecosystems and plant associations of Kenai Fjords National Park. Natural Resource Technical Report NPS/KEFJ/NRTR-2008/136. National Park Service, Fort Collins, CO.
  • Faber-Langendoen, D., J. Drake, S. Gawler, M. Hall, C. Josse, G. Kittel, S. Menard, C. Nordman, M. Pyne, M. Reid, L. Sneddon, K. Schulz, J. Teague, M. Russo, K. Snow, and P. Comer, editors. 2010-2019a. Divisions, Macrogroups and Groups for the Revised U.S. National Vegetation Classification. NatureServe, Arlington, VA. plus appendices. [in preparation]
  • Viereck, L. A., C. T. Dyrness, A. R. Batten, and K. J. Wenzlick. 1992. The Alaska vegetation classification. General Technical Report PNW-GTR286. USDA Forest Service, Pacific Northwest Research Station, Portland, OR. 278 pp.