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G285 Carex cusickii - Carex aquatilis - Myrica gale Alkaline Fen Group
Type Concept Sentence: This group consists of neutral to alkaline fens where brown mosses such as Campylium, Drepanocladus, Scorpidium, Tomentypnum, and Warnstorfia and sedges (Carex) are characteristic, and other indicator species include Betula nana, Carex aquatilis, Carex cusickii, Carex limosa, Carex livida, Carex utriculata, Comarum palustre, Menyanthes trifoliata, Myrica gale, and Spiraea douglasii. It is known from the Pacific Coast from Alaska south to northern California, in and west of the coastal mountain summits but including the Puget Sound lowlands.
Common (Translated Scientific) Name: Cusick''s Sedge - Water Sedge - Sweetgale Alkaline Fen Group
Colloquial Name: North Pacific Alkaline Open Fen
Hierarchy Level: Group
Type Concept: This wetland group occurs in peatlands along the Pacific Coast from Alaska south to northern California, in and west of the coastal mountain summits but including the Puget Sound lowlands. This group consists of neutral to alkaline fens (pH base-rich, neutral or alkaline); the extreme end is found often on calcareous substrates, but this is more the exception. Brown mosses such as Campylium, Drepanocladus, Scorpidium, Tomentypnum, and Warnstorfia and sedges (Carex) characterize rich fens (pH >5.5). Some sites may have high cover of minerotrophic non-hummock-forming Sphagnum species. Low shrub or graminoid physiognomy is typical. Groundwater input provides for the high mineral content, and soil saturation in general precludes tall vegetative growth. Shrubs are usually deciduous willows and birch, but evergreen dwarf-shrubs are not uncommon. Indicator species include Betula nana, Carex aquatilis, Carex cusickii, Carex limosa, Carex livida, Carex utriculata, Comarum palustre, Menyanthes trifoliata, Myrica gale, and Spiraea douglasii. Trees may be present at the edges or on raised hummocks where soils are not anoxic and include Callitropsis nootkatensis, Picea sitchensis, Pinus contorta, Thuja plicata, and/or Tsuga heterophylla. Elevations range from under 457 m (1500 feet) to over 2100 m (7000 feet), and annual precipitation ranges from 890-3050 mm (35-120 inches). These wetlands are relatively abundant in Alaska and British Columbia but diminish rapidly in size and number farther south. They occur in river valleys, around lakes and marshes or on slopes. The organic soils can be characterized by an abundance of sodium cations from oceanic precipitation when they reside within the hypermaritime zone.
Diagnostic Characteristics: Wetlands with deep organic soils (>40 cm) with strong groundwater influence that are >5.5 pH soil water.
Rationale for Nominal Species or Physiognomic Features: No Data Available
Classification Comments: These herbaceous and dwarf-shrub bogs and fens are distinguished from North Pacific treed bogs and fens by having (if present) stunted trees <5 m in height. These peatlands are distinguished from boreal continental bogs and fens by the presence of Pacific coastal species, including Carex aquatilis var. dives, Carex obnupta, Carex pluriflora, Darlingtonia californica, Gaultheria shallon, Sphagnum austinii, Sphagnum henryense, Sphagnum mendocinum, Sphagnum pacificum, Sphagnum papillosum, Sphagnum rubellum, and Spiraea douglasii. Stunted tree species, if present, include Callitropsis nootkatensis, Picea sitchensis, Pinus contorta var. contorta, Thuja plicata, and Tsuga heterophylla.
Pacific Northwest coastal peatlands occur across a gradient of pH depending on the amount of groundwater influence and the nutrient status (geology) of the water. The two ends of the spectrum are very acidic ("poor") fens and bogs to highly alkaline ("rich") fens. However, many wetlands occur in the mid-range of this gradient, and change overtime, such that it may prove difficult to keep "acidic" and "neutral-alkaline" separated at the group level. Indeed, many of the floristic dominants of described associations in British Columbia, Oregon and Washington occur across the pH gradient. These wetlands are not acidic and have plenty of contact with groundwater inflows.
Pacific Northwest coastal peatlands occur across a gradient of pH depending on the amount of groundwater influence and the nutrient status (geology) of the water. The two ends of the spectrum are very acidic ("poor") fens and bogs to highly alkaline ("rich") fens. However, many wetlands occur in the mid-range of this gradient, and change overtime, such that it may prove difficult to keep "acidic" and "neutral-alkaline" separated at the group level. Indeed, many of the floristic dominants of described associations in British Columbia, Oregon and Washington occur across the pH gradient. These wetlands are not acidic and have plenty of contact with groundwater inflows.
Similar NVC Types: No Data Available
note: No Data Available
Physiognomy and Structure: Brown mosses and sedges characterize alkaline and rich fens (pH >5.5). Some sites may have high cover of minerotrophic non-hummock-forming Sphagnum species. Low shrub or graminoid physiognomy is typical with some dwarf-shrubs.
Floristics: Brown mosses such as Campylium, Drepanocladus, Scorpidium, Tomentypnum, and Warnstorfia and sedges (Carex) characterize rich fens (pH >5.5). Some sites may have high cover of minerotrophic non-hummock-forming Sphagnum species. Low shrub or graminoid physiognomy is typical. Groundwater inputs provide for the high mineral content, and soil saturation in general precludes tall vegetative growth. Shrubs are usually deciduous willows and birch, but evergreen dwarf-shrubs are not uncommon. Indicator species include Betula nana, Carex aquatilis, Carex cusickii, Carex limosa, Carex livida, Carex utriculata, Comarum palustre, Menyanthes trifoliata, Myrica gale, and Spiraea douglasii. Trees may be present at the edges or on raised hummocks where soils are not anoxic and include Callitropsis nootkatensis (= Chamaecyparis nootkatensis), Picea sitchensis, Pinus contorta, Thuja plicata, and/or Tsuga heterophylla. Floristic information was summarized from the following sources: Shacklette et al. (1969), Eyre (1980), Banner et al. (1986, 1988, 1993), DeMeo et al. (1992), Viereck et al. (1992), Kunze (1994), Talbot and Talbot (1994), Martin et al. (1995), Shephard (1995), DeVelice et al. (1999), Boggs et al. (2003, 2008a, 2008b), MacKenzie and Moran (2004), Talbot et al. (2006), and Fleming and Spencer (2007).
Dynamics: Much literature documents the formation and succession of peatlands in Boreal, Temperate and Pacific Coastal settings (Banner et al. 1988, 2005, Mitsch and Gosselink 2000, and many others). These processes tend to take several hundreds to thousands of years, are well-documented and will not be repeated here. The purpose of this section is to discuss dynamics from disturbance. Natural disturbance events along north coast peatlands play an important role in slowing and reducing organic matter accumulation on specific types of sites. The main types of natural disturbance on the north Pacific coast are landslides, windthrow, and fluvial activity (Banner et al. 2005). Peatlands on steep slopes experience landslides and windthrow events which tend to mix soil and slow the buildup of organic layers. On more gentle terrain, disturbance is less frequent which allows for a deep accumulation of organic matter. Disturbance here is of small and localized scale. Large-scale disturbance such as major blowdown or severe fire are infrequent (>1000 years).
Fire, when it does occur, stimulates growth of mosses that prefer bare substrates, and "re-starts" a successional sequence of different mosses and vascular plants. Fires also increase nutrient availability and may temporarily create a carbon sink. Fire scars on stumps within herbaceous bogs and fens have been observed, pointing to a need for more study (Banner et al. 2005, J. Rocchio pers. comm. 2013). It has been documented that Native Americans burned herbaceous wetlands of the Olympic Peninsula annually to keep trees from invading wetland openings (Anderson 2009). Logging has also been shown to increase Sphagnum growth, through increased sunlight availability (Banner et al. 2005).
Landslides, fire and changes to water table depth and water source will alter the amount, composition and competition of peatland moss and higher vascular plant species. Fire can remove surface vegetation in dry seasons and stimulate the growth of Sphagnum spp. such that hummocks develop where once a depression occurred (Banner et al. 1988), shifting the small-scale local mosaic of swale and hummock within a peatland occurrence. Lowering the water table can allow trees such as Pinus contorta, Tsuga heterophylla, and Betula papyrifera to expand into peatland areas. Disturbance may bring the influx of mineral-rich waters from surrounding uplands (particularly along drainage channels), and flora characteristic of bogs is replaced by fen plants such as Spiraea spp., Salix spp., Pyrus spp., Juncus spp. and Carex spp. (Banner et al. 1988). Succession of mined peatlands depends on the degree of humification of the remaining substrate and hydrology. On unhumified peatmoss Sphagnum spp., Drosera rotundifolia, Rubus chamaemorus, and Ledum groenlandicum can regenerate and new Sphagnum peat can accumulate in pits and pools after mining (Banner et al 1988).
In addition to disturbance, there are natural cycles and interplay between the advancement and retreat of peat levels (the buildup of peat that alters depth of rooting zone relative to groundwater, or decrease in in peat depth from an increase decomposition due to drought) and encroachment peat into surrounding uplands (paludification) or the reverse, surrounding upland species that may shade out moss species. Generally, the greatest (annually measurable) change occurs on the edges of fens and bogs and at the tops of hummocks. These small-scale dynamics depend on the type of wetland and specific local site characteristics that cannot be generalized in a group description.
Monitoring peat wetlands is an important source of information as to whether observed changes are due to natural climatic or successional cycles or direct changes of water sources by human hands, or other indirect effects of human activity that may cause sedimentation, draining, flooding, or other changes to peatlands.
Fire, when it does occur, stimulates growth of mosses that prefer bare substrates, and "re-starts" a successional sequence of different mosses and vascular plants. Fires also increase nutrient availability and may temporarily create a carbon sink. Fire scars on stumps within herbaceous bogs and fens have been observed, pointing to a need for more study (Banner et al. 2005, J. Rocchio pers. comm. 2013). It has been documented that Native Americans burned herbaceous wetlands of the Olympic Peninsula annually to keep trees from invading wetland openings (Anderson 2009). Logging has also been shown to increase Sphagnum growth, through increased sunlight availability (Banner et al. 2005).
Landslides, fire and changes to water table depth and water source will alter the amount, composition and competition of peatland moss and higher vascular plant species. Fire can remove surface vegetation in dry seasons and stimulate the growth of Sphagnum spp. such that hummocks develop where once a depression occurred (Banner et al. 1988), shifting the small-scale local mosaic of swale and hummock within a peatland occurrence. Lowering the water table can allow trees such as Pinus contorta, Tsuga heterophylla, and Betula papyrifera to expand into peatland areas. Disturbance may bring the influx of mineral-rich waters from surrounding uplands (particularly along drainage channels), and flora characteristic of bogs is replaced by fen plants such as Spiraea spp., Salix spp., Pyrus spp., Juncus spp. and Carex spp. (Banner et al. 1988). Succession of mined peatlands depends on the degree of humification of the remaining substrate and hydrology. On unhumified peatmoss Sphagnum spp., Drosera rotundifolia, Rubus chamaemorus, and Ledum groenlandicum can regenerate and new Sphagnum peat can accumulate in pits and pools after mining (Banner et al 1988).
In addition to disturbance, there are natural cycles and interplay between the advancement and retreat of peat levels (the buildup of peat that alters depth of rooting zone relative to groundwater, or decrease in in peat depth from an increase decomposition due to drought) and encroachment peat into surrounding uplands (paludification) or the reverse, surrounding upland species that may shade out moss species. Generally, the greatest (annually measurable) change occurs on the edges of fens and bogs and at the tops of hummocks. These small-scale dynamics depend on the type of wetland and specific local site characteristics that cannot be generalized in a group description.
Monitoring peat wetlands is an important source of information as to whether observed changes are due to natural climatic or successional cycles or direct changes of water sources by human hands, or other indirect effects of human activity that may cause sedimentation, draining, flooding, or other changes to peatlands.
Environmental Description: Elevations are mostly under 457 m (1500 feet) but range up into the subalpine to 2100 m (7000 feet)), and annual precipitation ranges from 890-3050 mm (35-120 inches). These wetlands are relatively abundant in Alaska and British Columbia but diminish rapidly in size and number farther south. They occur in river valleys, around lakes and marshes, or on slopes. In Hypermaritime settings, the organic soils are characterized by an abundance of sodium cations from oceanic precipitation. This group consists of mostly circumneutral (ranging from slightly acidic to basic) fens. Some are found often on calcareous substrates, which represent the extreme alkaline range within the group. Environmental information was summarized from the following sources: Shacklette et al. (1969), Eyre (1980), Banner et al. (1986, 1988, 1993), DeMeo et al. (1992), Viereck et al. (1992), Kunze (1994), Talbot and Talbot (1994), Martin et al. (1995), Shephard (1995), DeVelice et al. (1999), Boggs et al. (2003, 2008a, 2008b), MacKenzie and Moran (2004), Talbot et al. (2006), and Fleming and Spencer (2007).
Geographic Range: This group is found along the Pacific Coast from central coast and southeastern Alaska, British Columbia, Washington, Oregon to northern California, in and west of the coastal mountain summits but including the Puget Sound lowlands.
Nations: CA,US
States/Provinces: AK, BC, CA, OR, WA
Plot Analysis Summary:
http://vegbank.org/natureserve/ELEMENT_GLOBAL.2.856669
Confidence Level: Moderate
Confidence Level Comments: No Data Available
Grank: GNR
Greasons: No Data Available
Type | Name | Database Code | Classification Code |
---|---|---|---|
Class | 2 Shrub & Herb Vegetation Class | C02 | 2 |
Subclass | 2.C Shrub & Herb Wetland Subclass | S44 | 2.C |
Formation | 2.C.2 Temperate to Polar Bog & Fen Formation | F016 | 2.C.2 |
Division | 2.C.2.Na North American Bog & Fen Division | D029 | 2.C.2.Na |
Macrogroup | 2.C.2.Na.3 Beach Pine / Bog Labrador-tea / Cusick''s Sedge North Pacific Bog & Fen Macrogroup | M063 | 2.C.2.Na.3 |
Group | 2.C.2.Na.3.a Cusick''s Sedge - Water Sedge - Sweetgale Alkaline Fen Group | G285 | 2.C.2.Na.3.a |
Alliance | A3432 Sweetgale - Rose Spirea Alkaline Fen Alliance | A3432 | 2.C.2.Na.3.a |
Alliance | A3433 Water Sedge - Livid Sedge - Cusick''s Sedge Alkaline Fen Alliance | A3433 | 2.C.2.Na.3.a |
Alliance | A3599 Woolly-fruit Sedge - Bellardi Bog Sedge - Northern Single-spike Sedge Alkaline Graminoid Fen Alliance | A3599 | 2.C.2.Na.3.a |
Concept Lineage: No Data Available
Predecessors: No Data Available
Obsolete Names: No Data Available
Obsolete Parents: No Data Available
Synonomy: No Data Available
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