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G493 Morella pensylvanica - Ammophila breviligulata - Schizachyrium littorale Dune & Grassland Group
Type Concept Sentence: This group encompasses northeastern maritime dune vegetation comprising grasses, low shrubs, dwarf-shrubs, and forbs ranging from sparse to patchy in cover. It occupies sandy coastlines and barrier islands, ranging from northernmost North Carolina northward to southern Maine where extensive sandy coastlines are replaced by rocky coasts.
Common (Translated Scientific) Name: Northern Bayberry - American Beachgrass - Shore Little Bluestem Dune & Grassland Group
Colloquial Name: North Atlantic Coastal Dune & Grassland
Hierarchy Level: Group
Type Concept: This group encompasses vegetation of sandy coastlines and barrier islands, ranging from northernmost North Carolina northward to southern Maine where extensive sandy coastlines are replaced by rocky coasts. A range of physiognomies may be present, from somewhat sparse herbaceous dune vegetation with Ammophila breviligulata diagnostic, to backdune shrublands, and shrub-herb patchworks on nearshore sandplains. Characteristic species (in different associations within the group) include Ammophila breviligulata, Andropogon virginicus, Cenchrus tribuloides, Diodia teres, Hudsonia tomentosa, Morella pensylvanica, Panicum amarum, Panicum virgatum, Polygonella articulata, Schizachyrium littorale, Schizachyrium scoparium, and Solidago sempervirens, among others. Dominant ecological processes are those associated with the maritime environment, including frequent salt spray, wind exposure, saltwater overwash, and wind-transported sand movement that are severe enough to limit tree growth. Vegetation in this group includes grasslands and shrublands of foredunes and more stabilized backdunes, as well as sandplain grasslands and heathlands of the southern New England / New York coast. In Atlantic Canada, coastal sandplain grasslands can also occur with 10s to 100s of meters from the coast. Most of the dwarf dune heath is Empetrum spp.-dominated, but some stands feature Corema conradii and Hudsonia tomentosa.
Diagnostic Characteristics: Restricted to maritime sand dunes and coastal sandplains; dominance by low shrubs and grasses, ranging from sparse to patchy in cover. This group does not include maritime forests also occurring on dunes.
Rationale for Nominal Species or Physiognomic Features: No Data Available
Classification Comments: This group shares physiognomic character and some species with the Carolinian and Gulf coasts analog ~South Atlantic & Gulf Coastal Dune & Grassland Group (G494)$$, but treating the two as one group results in one that ranges from Maine to Texas with almost no species in common at the extremes. The separation of the groups parallels broad-scale biogeographic and climatic differences believed to be important in this environment, and occurs more-or-less in the southern part of the Virginian Province (as labeled by Cowardin et al. 1979) as it transitions to the Carolinian Province around Cape Hatteras. A useful vegetation indicator of this transition is the shift in herbaceous dominance on the dunes from Ammophila breviligulata in the present group to Uniola paniculata in the south. Although the location of this shift itself is somewhat imprecise because of widespread planting of both species on artificially enhanced dunes, this boundary appears to be well approximated by Omernik Ecoregion 63g vs. 63d (EPA 2004). The group extends north to the end of extensive sandy coastlines and the beginning of rocky coasts.
Similar NVC Types: No Data Available
note: No Data Available
Physiognomy and Structure: This vegetation is variable in structure, ranging from sparsely distributed grasses on foredunes, sometimes intermixed with low deciduous shrubs, to dwarf-shrublands of interdunes and low shrublands of coastal sandplains.
Floristics: Morella pensylvanica, Ammophila breviligulata, and Hudsonia tomentosa are species that distinguish this group from vegetation in similar habitats farther south. The foredune areas tend to be low in plant species richness but have a characteristic set of forbs and occasional low shrubs associated with them. Backdune areas may be dominated by a variety of herbs and sometimes have fairly high species richness. Characteristic species (in different associations within the group) include Ammophila breviligulata, Andropogon virginicus, Cenchrus tribuloides, Diodia teres, Hudsonia tomentosa, Morella pensylvanica, Panicum amarum, Panicum virgatum, Polygonella articulata, Schizachyrium littorale, Schizachyrium scoparium, and Solidago sempervirens, among others. Schoenoplectus pungens sometimes occurs in overwash areas where sand has buried interdunal wetlands. Heathland and sandplain grassland settings are characterized by Amelanchier nantucketensis, Arctostaphylos uva-ursi, Carex pensylvanica, Corema conradii, Deschampsia flexuosa, Gaylussacia baccata, Hudsonia ericoides, Hudsonia tomentosa, Schizachyrium scoparium, and Vaccinium angustifolium. They provide habitat for several rare or uncommon forbs, including Liatris scariosa var. novae-angliae and Agalinis acuta.
Dynamics: The environment of this group is one of the most dynamic in existence for terrestrial vegetation. Reworking of sand by storms or by slower eolian processes may completely change the local environment in a short time, altering the substrate that effectively removes existing vegetation and provides new habitat for establishment of different vegetation. Some plants, such as Ammophila breviligulata and Hudsonia tomentosa, are well-adapted to sand burial and can send new shoots above the sand surface. Many of these sites are fairly early in the process of primary succession on recent surfaces. Chronic salt spray is an ongoing stress. Overwash and extreme salt spray from storms are frequent disturbances. Vegetation interacts strongly with geologic processes; the presence of grass is an important factor in the development of new dunes. Alteration of dynamic processes, such as artificial enhancement of dunes by planting or sand fencing, can have drastic effects on this vegetation, causing large areas to succeed to woody dominance. Limited areas along the New York and New England coastline support maritime sandplains, where sand movement is more limited but exposure to the elements is still high. The sandplain grasslands and heathlands that develop in these habitats have been affected by a combination of exposure, fire, and land use. Prior to European settlement, these habitats are believed to have occurred in limited areas near the coast, where the effects of wind and salt spray prevented tree growth; there may also have been patches in the vicinity of Native American settlements, based on the prevalence of charcoal in some palynological cores. Some heathlands may have developed on severely disturbed soils following the abandonment of agriculture and grazing.
Environmental Description: Climate: Vegetation in this group is subject to a north-temperate maritime climate. Wind and salt spray are important factors in vegetation development. Soil/substrate/hydrology: This vegetation occurs on coastal strands and barrier islands, on sand dunes and sand flats. Strong salt spray is an important influence on vegetation in many parts. Overwash by seawater during storms is important on sand flats not protected by continuous dunes. Overwash caused by coastal storms can profoundly change the character of the vegetation where it overwhelms dunes and deposits new sand over existing vegetation. On dunes, present or recent sand movement is an important factor. On sandplains away from the immediate shore, but still subject to maritime influence, sand movement is less of a factor but wind and salt spray remain important. The combination of these factors prevents the dominance of woody vegetation. Most sites may be dry, but some areas are temporarily wet by freshwater from rainfall and the local water table. Soils are sandy, with little organic matter and little or no horizon development. Soils may be excessively drained on the higher dunes. Soils are low in nutrient-holding capacity, but aerosol input of sea salt provides a continuous source of nutrients.
Geographic Range: This group ranges from northernmost North Carolina (EPA ecoregion 63d) and southeastern Virginia to southern Maine. The southern portion is a transition zone from around Kitty Hawk, North Carolina, to the Virginia-North Carolina border. The northern limit is around Merrymeeting Bay, Maine.
Nations: CA,US
States/Provinces: CT, DE, MA, MD, ME, NB, NC, NH, NJ, NS, NY, PE, RI, VA
Plot Analysis Summary:
http://vegbank.org/natureserve/ELEMENT_GLOBAL.2.837100
Confidence Level: Moderate
Confidence Level Comments: No Data Available
Grank: GNR
Greasons: No Data Available
Type | Name | Database Code | Classification Code |
---|---|---|---|
Class | 2 Shrub & Herb Vegetation Class | C02 | 2 |
Subclass | 2.B Temperate & Boreal Grassland & Shrubland Subclass | S18 | 2.B |
Formation | 2.B.4 Temperate to Polar Scrub & Herb Coastal Vegetation Formation | F005 | 2.B.4 |
Division | 2.B.4.Na Eastern North American Coastal Scrub & Herb Vegetation Division | D026 | 2.B.4.Na |
Macrogroup | 2.B.4.Na.2 American Beachgrass - Sea-oats - Seaside Goldenrod Dune & Grassland Macrogroup | M057 | 2.B.4.Na.2 |
Group | 2.B.4.Na.2.c Northern Bayberry - American Beachgrass - Shore Little Bluestem Dune & Grassland Group | G493 | 2.B.4.Na.2.c |
Alliance | A0902 Northern Bayberry Dune Shrubland Alliance | A0902 | 2.B.4.Na.2.c |
Alliance | A1062 Woolly Beach-heather Dune Dwarf-shrubland Alliance | A1062 | 2.B.4.Na.2.c |
Alliance | A4470 <i>Ammophila breviligulata - Schizachyrium littorale - Spartina patens</i> Dune Grassland | A4470 | 2.B.4.Na.2.c |
Concept Lineage: No Data Available
Predecessors: No Data Available
Obsolete Names: No Data Available
Obsolete Parents: No Data Available
Synonomy: > Andropogon Hempsteadi (Cain et al. 1937)
> Hudsonietum tomentosi (Conard 1935)
> Beach heather community (Robichaud and Buell 1973)
> Coastal Heathland and Sandplain Grassland (Dunwiddie et al. 1996)
> Coastal sand dune (Dunlop and Crow 1983)
> Dunegrass community (Robichaud and Buell 1973)
> Middle beach (Chrysler 1930)
> Myrica thicket (Chrysler 1930)
> Upper beach (Chrysler 1930)
> Hudsonietum tomentosi (Conard 1935)
> Beach heather community (Robichaud and Buell 1973)
> Coastal Heathland and Sandplain Grassland (Dunwiddie et al. 1996)
> Coastal sand dune (Dunlop and Crow 1983)
> Dunegrass community (Robichaud and Buell 1973)
> Middle beach (Chrysler 1930)
> Myrica thicket (Chrysler 1930)
> Upper beach (Chrysler 1930)
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