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G127 Scaevola plumieri - Suriana maritima - Ipomoea pes-caprae Caribbean Coastal Beach & Dune Group
Type Concept Sentence: This group consists of Caribbean, south Florida and Bahamas sparse vegetation of the beach strand line, transitional dune, and stabilized coastal dunes with grasses, herbs and low shrubs.
Common (Translated Scientific) Name: Gull-feed - Bay-cedar - Bayhops Caribbean Coastal Beach & Dune Group
Colloquial Name: Caribbean Coastal Beach & Dune
Hierarchy Level: Group
Type Concept: This group consists of Caribbean, south Florida and Bahamas sparse vegetation of the beach strand line, transitional dune, and stabilized coastal dunes with grasses, herbs and low shrubs. In the Caribbean islands, it occurs also in hard-packed sand areas behind sand beaches. Dunes are uncommon on Caribbean islands; open beaches are common. The process of sand movement due to the forces of wind and water is part of the natural dynamics of beach ecosystems. This includes transport of sand along the coast, and movement of sand by wind or water between the dunes, beach and subtidal areas. The following list of species is diagnostic for this group: Ambrosia hispida, Argusia gnaphalodes, Borrichia arborescens, Cakile lanceolata, Calotropis procera, Canavalia rosea, Cassia lineata, Cenchrus echinatus, Chamaesyce blodgettii, Chamaesyce dioica, Chamaesyce mesembrianthemifolia, Chrysobalanus icaco, Coccoloba uvifera, Croton punctatus, Cyperus planifolius, Egletes prostrata, Heliotropium nanum, Heterostachys ritteriana, Hippomane mancinella, Ipomoea pes-caprae, Iva imbricata, Lycium tweedianum, Opuntia caracassana, Piscidia piscipula, Pithecellobium keyense, Portulaca pilosa, Scaevola plumieri, Sesuvium portulacastrum, Spartina patens, Sporobolus virginicus, Tournefortia volubilis, Tribulus zeyheri, and Turnera diffusa.
Diagnostic Characteristics: The beach environment of this group is primarily upland above the normal high tide line, with some wet patches fed by groundwater. The beach vegetation may be very sparse, but slightly higher areas and dunes may be sparse or patchy and grass-dominated in its cover. The shifting substrate (of sand, small pebbles or shell fragments) largely limits the vegetation to pioneering, salt-tolerant, succulent annuals, grasses, perennial vines, and a few shrubs. Perennial vines such as Ipomoea pes-caprae are generally more characteristic of tropical beaches than of temperate ones.
Rationale for Nominal Species or Physiognomic Features: No Data Available
Classification Comments: This group includes vegetation of carbonate sand beaches of the Florida Keys and south Florida mangrove islands (after Johnson and Barbour 1990). Should the beaches of southern Texas and southern Florida be placed with other neotropical beaches? Are there species that reliably distinguish neotropical beaches from tropical beaches in Hawai''i and the Pacific? Should south Texas beach systems be placed here with those of southern Florida? There are presumably a number of vegetation types which would be components of this group that are as yet undescribed. It may not be possible (nor necessary) to distinguish these tropical beaches of the United States from beaches of the Caribbean. If this group is to be maintained, should it be "Tropical Mainland North America" Beach? Where to draw the line among beaches of the mainland Gulf and Caribbean coasts from Texas to Yucatan and beyond?
Similar NVC Types: No Data Available
note: No Data Available
Physiognomy and Structure: The dynamic disturbance regimes largely limit the vegetation to pioneering, salt-tolerant, succulent annuals and perennial vines, which are more prevalent in these tropical examples in contrast to more northerly and temperate ones. The stands of vegetation are generally low in stature (with some clumps of taller grasses) and sparse to patchy in their cover.
Floristics: This group consists of the beach strand line, transitional dune, and stabilized coastal dunes with grasses and herbs. In the Caribbean islands, it occurs also in hard-packed sand areas behind sand beaches. Dunes are uncommon on Caribbean islands; open beaches are common. The following list of species is diagnostic for this group: Ambrosia hispida, Argusia gnaphalodes (= Mallotonia gnaphalodes, = Tournefortia gnaphalodes), Borrichia arborescens, Cakile lanceolata, Calotropis procera, Canavalia rosea (= Canavalia maritima), Cassia lineata, Cenchrus echinatus, Chamaesyce blodgettii (= Euphorbia blodgettii), Chamaesyce dioica, Chamaesyce mesembrianthemifolia (= Euphorbia buxifolia), Chrysobalanus icaco, Coccoloba uvifera, Croton punctatus, Cyperus planifolius, Egletes prostrata, Heliotropium nanum, Heterostachys ritteriana, Hippomane mancinella, Ipomoea pes-caprae, Iva imbricata, Lycium tweedianum (= Lycium nodosum), Opuntia caracassana, Piscidia piscipula, Pithecellobium keyense, Portulaca pilosa, Scaevola plumieri, Sesuvium portulacastrum, Spartina patens, Sporobolus virginicus, Tournefortia volubilis, Tribulus zeyheri, and Turnera diffusa. Along the Caribbean coast of Mexico and Central America, plants include Acoelorraphe wrightii, Bromus sp., Clitoria falcata (= Clitoria rubiginosa), Crotalaria retusa, Dactyloctenium aegyptium, Dodonaea sp., Hibiscus tiliaceus, Hymenocallis littoralis, Manicaria sp., Mimosa pudica, Morinda citrifolia, Phyllanthus acidus, Sphagneticola trilobata (= Wedelia trilobata), Stachytarpheta jamaicensis, and Tridax procumbens. On isolated dunes there are short trees or shrubs of Conocarpus erectus, Prosopis juliflora, and Acacia tortuosa. Cocos nucifera is a common introduced palm, and the introduced tree Casuarina equisetifolia has been planted along the coast in Florida, the Bahamas, and the Caribbean. Other invasive exotic plants in coastal strand in south Florida include Colubrina asiatica, Cupaniopsis anacardioides, Neyraudia reynaudiana, Scaevola sericea var. taccada (= Scaevola taccada), and Schinus terebinthifolius (FNAI 2010a).
Dynamics: The process of sand movement due to the forces of wind and water is part of the natural dynamics of beach ecosystems. This includes transport of sand along the coast, and movement of sand by wind or water between the dunes, beach and subtidal areas. If not restricted by infrastructure or engineered hard structures, beaches and dunes can migrate as coastlines change over time in response to the action of wind and water (Defeo et al. 2009). Coastal engineering such as seawalls and other hard structures are among the threats to the natural dynamics of the beach (Defeo et al. 2009). Dunes are best developed along the Atlantic Coast of Florida; many beaches in the Caribbean lack dunes. Plants on the foredune are regularly exposed to salt spray and sand burial from onshore winds. Plants on the upper beach are subject to these stresses plus occasional inundation by seasonal or storm tides and periodic destruction by waves. The plants of the beach dune community are adapted to either withstand these stresses or to rapidly re-colonize from seed or vegetative parts following destruction. Fertilization from piles of seaweed washed up by storms helps to speed plant growth and the re-colonization process. Once a new foredune ridge blocks salt spray and plant cover inhibits sand movement, inland herbaceous and eventually woody species can begin to replace the coastal pioneer species of the beach dune community in the backdune area. The coastal berm is deposited by storm waves along low-energy coasts. Their distance inland depends on the height of the storm surge. Tall berms may be the product of repeated storm deposition. Excavation of one berm in the Florida Keys revealed several layers of buried soils, evidence of burial by repeated storms at relatively long intervals (Kruer 1992, as cited in FNAI 2010a).
Environmental Description: Climate: The climate is tropical or subtropical, and these sites have onshore winds. The Caribbean shores are subject to hurricanes. Frosts are rare events in coastal areas of south Texas and south Florida.
Soil/substrate/hydrology: The communities within this group are found on reliefs constituted either by dunes, onshore wind-carried sand deposits arranged in cordons of ridges parallel to the coast, or by beach-ridges, wave and longshore drift-carried sand deposits, also often organized in successive parallel berms produced by the progradation of the beach. Dunes are best developed along the Atlantic Coast of Florida; many beaches in the Caribbean lack dunes. Beach dune may be distinguished from coastal grassland by its position above the immediate shoreline and by the dominance of burial-tolerant grasses such as Uniola paniculata and Panicum amarum in Florida, the Bahamas, and Cuba. It differs from coastal berm in its position facing the open ocean on a sandy coast rather than on a storm-deposited shell ridge on a mangrove-dominated shoreline.
Soil/substrate/hydrology: The communities within this group are found on reliefs constituted either by dunes, onshore wind-carried sand deposits arranged in cordons of ridges parallel to the coast, or by beach-ridges, wave and longshore drift-carried sand deposits, also often organized in successive parallel berms produced by the progradation of the beach. Dunes are best developed along the Atlantic Coast of Florida; many beaches in the Caribbean lack dunes. Beach dune may be distinguished from coastal grassland by its position above the immediate shoreline and by the dominance of burial-tolerant grasses such as Uniola paniculata and Panicum amarum in Florida, the Bahamas, and Cuba. It differs from coastal berm in its position facing the open ocean on a sandy coast rather than on a storm-deposited shell ridge on a mangrove-dominated shoreline.
Geographic Range: The range of this group includes the shorelines of the Caribbean, Bahamas, and including south Texas and south Florida.
Nations: BS?,BZ,CO,CR,CU,DO,GT,HN,HT,JM,MX,NI,PA,PR,US,VE,VI,XC
States/Provinces: FL, TX
Plot Analysis Summary:
http://vegbank.org/natureserve/ELEMENT_GLOBAL.2.837022
Confidence Level: Moderate
Confidence Level Comments: No Data Available
Grank: GNR
Greasons: No Data Available
Type | Name | Database Code | Classification Code |
---|---|---|---|
Class | 2 Shrub & Herb Vegetation Class | C02 | 2 |
Subclass | 2.A Tropical Grassland, Savanna & Shrubland Subclass | S01 | 2.A |
Formation | 2.A.3 Tropical Scrub & Herb Coastal Vegetation Formation | F024 | 2.A.3 |
Division | 2.A.3.Ee Caribbean-Mesoamerican Dune & Coastal Grassland & Shrubland Division | D254 | 2.A.3.Ee |
Macrogroup | 2.A.3.Ee.1 Seashore Dropseed - Bayhops - Bay-cedar Coastal Dune & Beach Vegetation Macrogroup | M700 | 2.A.3.Ee.1 |
Group | 2.A.3.Ee.1.a Gull-feed - Bay-cedar - Bayhops Caribbean Coastal Beach & Dune Group | G127 | 2.A.3.Ee.1.a |
Concept Lineage: G469 is redundant with G127 (DFL 5-17-13)
Predecessors: No Data Available
Obsolete Names: No Data Available
Obsolete Parents: No Data Available
Synonomy: < Dune (Johnson and Barbour 1990)
? Dunes with sparse vegetation Formation (Areces-Mallea et al. 1999)
? Littoral Subzone, dune ecosystem (Dansereau 1966)
? Puerto Rico land cover type 59, Fine to coarse sandy beaches, mixed sand and gravel beaches (Gould et al. 2008)
? Strand vegetation Formation (Areces-Mallea et al. 1999)
? Dunes with sparse vegetation Formation (Areces-Mallea et al. 1999)
? Littoral Subzone, dune ecosystem (Dansereau 1966)
? Puerto Rico land cover type 59, Fine to coarse sandy beaches, mixed sand and gravel beaches (Gould et al. 2008)
? Strand vegetation Formation (Areces-Mallea et al. 1999)
- Areces-Mallea, A. E., A. S. Weakley, X. Li, R. G. Sayre, J. D. Parrish, C. V. Tipton, and T. Boucher. 1999. A guide to Caribbean vegetation types: Preliminary classification system and descriptions. The Nature Conservancy, Arlington, VA. 166 pp.
- Barbour, M. G., M. Rejmanek, A. F. Johnson, and B. M. Pavlik. 1987. Beach vegetation and plant distribution patterns along the northern Gulf of Mexico. Phytocoenologia 15:201-234.
- Dansereau, P. 1966. Studies on the vegetation of Puerto Rico. Part I. Description and integration of the plant-communities. University of Puerto Rico, Institute of Caribbean Sciences. Special Publication No. 1. Mayagüez, Puerto Rico. 287 pp.
- FNAI [Florida Natural Areas Inventory]. 2010a. Guide to the natural communities of Florida: 2010 edition. Florida Natural Areas Inventory, Tallahassee, FL. 228 pp. [https://fnai.org/naturalcommguide.cfm]
- Faber-Langendoen, D., J. Drake, S. Gawler, M. Hall, C. Josse, G. Kittel, S. Menard, C. Nordman, M. Pyne, M. Reid, L. Sneddon, K. Schulz, J. Teague, M. Russo, K. Snow, and P. Comer, editors. 2010-2019a. Divisions, Macrogroups and Groups for the Revised U.S. National Vegetation Classification. NatureServe, Arlington, VA. plus appendices. [in preparation]
- Gould, W. A., C. Alarcón, B. Fevold, M. E. Jiménez, S. Martinuzzi, G. Potts, M. Quiñones, M. Solórzano, and E. Ventosa. 2008. The Puerto Rico Gap Analysis Project. Volume 1: Land cover, vertebrate species distributions, and land stewardship. General Technical Report IITF-GTR-39. USDA Forest Service, International Institute of Tropical Forestry, Río Piedras, PR. 165 pp.
- Johnson, A. F., and M. G. Barbour. 1990. Dunes and maritime forests. Pages 429-480 in: R. L. Myers and J. J. Ewel, editors. Ecosystems of Florida. University of Central Florida Press, Orlando.
- Tanner, W. F. 1960. Florida coastal classification. Gulf Coast Association of Geological Societies Transactions 10:259-266.