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G130 Quercus laurifolia - Quercus phellos - Pinus taeda Wet Flatwoods Group

Type Concept Sentence: These are nonriverine wetland hardwood-pine flatwood forests of the Atlantic and Gulf coastal plains and the Mississippi River Alluvial Plain, primarily dominated by Quercus spp., including primarily Quercus laurifolia, Quercus michauxii, Quercus nigra, Quercus pagoda, and Quercus phellos, and also with a variety of other hardwoods. In addition, Pinus glabra and/or Pinus taeda may be codominant or dominant in some examples, particularly in the Gulf Coastal Plain.


Common (Translated Scientific) Name: Laurel Oak - Willow Oak - Loblolly Pine Wet Flatwoods Group

Colloquial Name: Hardwood - Loblolly Pine Nonriverine Wet Flatwoods

Hierarchy Level:  Group

Type Concept: These are nonriverine wetland hardwood or pine-hardwood forests and woodlands of the Atlantic and Gulf coastal plains and the Mississippi River Alluvial Plain, ranging from southeastern Virginia to Texas. Stands occur in a variety of flat, wet habitats and are most frequently dominated by Quercus spp. (including Quercus laurifolia, Quercus phellos, Quercus pagoda, Quercus similis, and Quercus stellata). In the Atlantic Coastal Plain, Quercus michauxii, Quercus nigra, and Quercus virginiana are important components, In addition, particularly in the Gulf Coastal Plain, Pinus taeda is dominant or codominant. Pinus glabra may be present within its range. Other hardwoods, including Fraxinus pennsylvanica, Nyssa biflora, and Ulmus americana, may be present as well. Acer rubrum and Liquidambar styraciflua, which are often increased by timber removal or other soil disturbance, may be prominent in some examples. Sabal minor is abundant in the lower strata of some stands within its range. In the Mississippi River Alluvial Plain, which is outside of the typical range of pine, hardwoods are the sole dominants. These communities are generally known as "flatwoods," and are found on a variety of sites which are generally flat to very gently sloping, primarily Pleistocene terraces. These sites typically have poor internal drainage, often from strata in the soil that limit permeability (claypans, hardpans, etc.). This limited permeability of the soil contributes to shallowly perched water tables during portions of the year when precipitation is greatest and evapotranspiration is lowest. Conversely, during the dry season, there is very little available water, leading to seasonal droughtiness. The hydrologic regime is primarily influenced by rainwater (and groundwater in included depressions), rather than overbank flooding. In some cases the flatwoods are within the current floodplain, but the hydrology, soils and vegetation are dominated by flatwoods, rather than floodplain, characteristics. Soil moisture fluctuates widely throughout the growing season, partly as a result of included impermeable layers. Hydrologic conditions vary from saturated to very dry, a condition sometimes referred to as xerohydric or hydroxeric. Soils are primarily mineral but may have some organic matter or muck. In some areas (e.g., in the youngest Pleistocene terraces in the Upper West Gulf Coastal Plain of Arkansas), the local topography is a complex of ridges and swales, often in close proximity to one another. Ridges are typically drier than swales, which may hold water for varying periods of time. Within both ridges and swales, there is vegetation variability relating to soil texture and moisture and disturbance history. As a result of dry (xeric) conditions during the dry season (summer-fall), most sites with these communities burned frequently, and the characteristic physiognomy was woodland. In today''s landscapes, with reduced fire, most occurrences are forested.

Diagnostic Characteristics: The hydrology is nonriverine, with water coming from rainwater and (in depressions) groundwater rather than from overbank flooding. The environmental setting is wet flats with included depressions. Frequently burned sites have a woodland physiognomy, while most examples today are forested from lack of fire.

Rationale for Nominal Species or Physiognomic Features: No Data Available

Classification Comments: This group encompasses a range of hydroperiods, as is traditional for the vegetation commonly called "flatwoods." It is drier than basin swamps, which can occur in similar topography, and wetter than mesic or upland groups. The placement and classification of vegetation commonly called "flatwoods" depend on the length of the hydroperiod. Forests in this group represent this hydrologic variation, which helps define the component alliances. Some other flatwoods included in this group are wetter than most flatwoods but not as wet as basin swamps. The relative abundance and ecological role of Pinus taeda in examples of this group is dependent on its importance in the particular region. In the West Gulf Coastal Plain, Pinus taeda (with some Pinus echinata) may be a primary dominant in stands of this group. The description of NVC associations in this group is probably incomplete, and more information is needed. On the lower terraces of the East Gulf Coastal Plain, the most western parts of the region contain flatwoods vegetation dominated primarily by Quercus spp., whereas in the more central part of the region (sometimes called the "Louisiana Florida Parishes Spruce Pine Flatwoods Forest"), the vegetation is dominated by a mixture of hardwoods with Pinus glabra and Pinus taeda. Characteristic hardwoods include Magnolia grandiflora, Quercus laurifolia, Quercus michauxii, Quercus nigra, Quercus pagoda, and Quercus virginiana. Some important understory trees and shrubs include Crataegus opaca and Sabal minor (which may often be very abundant or dominant), with Arundinaria tecta (Smith 1996b).

Similar NVC Types: No Data Available
note: No Data Available

Physiognomy and Structure: Stands have generally closed canopies, and are dominated by broad-leaved deciduous trees, particularly Quercus species, with Pinus taeda sometimes abundant or dominant in the West Gulf Coastal Plain and in altered or disturbed examples elsewhere. The lower strata may have a broad-leaved evergreen or deciduous shrub component. Characteristics include flat topography on coastal plain terraces or high Mississippi River Alluvial Plain terraces, a perched or seasonal high water table (sometimes also with dry periods), a lack of overland flooding, and dominance by wetland hardwoods and/or pines that are shared with river floodplain communities.

Floristics: Examples of this vegetation are most frequently dominated by Quercus spp., including Quercus laurifolia, Quercus phellos, Quercus pagoda, Quercus similis, and Quercus stellata. In the Atlantic Coastal Plain, Quercus michauxii, Quercus nigra, and Quercus virginiana are important components. In addition, particularly in the Gulf Coastal Plain, Pinus taeda is dominant or codominant. Pinus glabra may be present within its range. Other hardwoods, including Fraxinus pennsylvanica, Nyssa biflora, and Ulmus americana, may be present as well. Acer rubrum and Liquidambar styraciflua, which are often increased by timber removal or other soil disturbance, may be prominent in some examples. Sabal minor is abundant in the lower strata of some stands within its range. Shrub and understory species may include Clethra alnifolia, Crataegus aestivalis, Crataegus crus-galli, Crataegus marshallii, Diospyros virginiana, Hypericum hypericoides, Ilex opaca var. opaca, Ilex verticillata, Morella cerifera, Rhapidophyllum hystrix, Rhododendron viscosum, Sabal minor, Sabal palmetto, Serenoa repens, Vaccinium elliottii, Vaccinium formosum, Vaccinium fuscatum, Vaccinium virgatum, Viburnum dentatum, and Viburnum nudum var. nudum. Woody vines include Nekemias arborea (= Ampelopsis arborea), Berchemia scandens, Bignonia capreolata, Campsis radicans, Gelsemium sempervirens, Parthenocissus quinquefolia, Smilax bona-nox, Smilax glauca, Smilax rotundifolia, Toxicodendron radicans, and Vitis rotundifolia. The herbaceous layer may be dense with grass and sedge species such as Carex cherokeensis, Carex debilis var. debilis, Carex flaccosperma, Carex glaucescens, Carex intumescens, Carex seorsa, Chasmanthium laxum, Chasmanthium nitidum, Dichanthelium aciculare, Dichanthelium commutatum, Dichanthelium scoparium, Leersia spp., Rhynchospora caduca, Rhynchospora inexpansa, Scleria oligantha, and Scleria triglomerata. Ferns may also be prominent, with scattered colonies and individuals of Osmunda cinnamomea, Osmunda regalis var. spectabilis, Woodwardia areolata, and Woodwardia virginica.

Dynamics:  In these communities, because soils are seasonally very dry, fire was originally ecologically significant. In today''s landscapes, fire has been greatly reduced, leading to changes in community composition and structure. Canopy dynamics are usually driven by small gaps, but hurricanes may occasionally create large canopy gaps. Examples in drowned river valleys are subject to influence by rising sea level and can be expected to evolve into tidal swamp systems, sometimes fairly quickly. Low-intensity surface fires were ecologically important in inland examples. The fire regime in drier flatwoods may be characterized as short- to medium-interval, low-intensity, medium- to high-severity; and in wetter flatwoods as short-interval, low-intensity, low-severity (D. Zollner pers. comm. 2006). Some, such as the wet hardwood flats of North Carolina, apparently were non-flammable enough that fire had negligible influence. In Arkansas, there are two distinct landscapes in which these flatwoods play a prominent role. One of these is the Mississippi River Alluvial Plain, where there are Pleistocene valley train terraces formed by glacial outwash that were occupied principally by oak-dominated flatwoods. There are similar Pleistocene terraces in the Upper West Gulf Coastal Plain, but these were not formed from glacial outwash, by high waterflows at the same time. Those were dominated by Pinus taeda with hardwoods becoming more important in the wetter areas. Each of these distinctive areas occupies about a million acres, so they represented major portions of the pre-settlement landscape.

Environmental Description:  Vegetation of this group occurs on flat areas, typically terraces of Pleistocene age of the Atlantic and Gulf coastal plains and the Mississippi River Alluvial Plain, from southeastern Virginia to Texas, and in the interior north to Arkansas, Oklahoma, and the Missouri "bootheel," in areas where soils seasonally vary from saturated to extremely dry because of low relief, poor internal and external soil drainage, seasonally high water table, and impervious layers that result in limited soil moisture during the dry season. Soil moisture fluctuates widely throughout the growing season, from saturated to very dry, a condition sometimes referred to as xerohydric or hydroxeric. The hydrology is dominated by rainfall and sheetflow, with groundwater seepage in depressions and at the boundaries of terraces. Overbank flooding has little to no influence on the vegetation. The largest areas are on broad interfluvial flats. Some examples of this environment include sites above modern floodplains but with poor internal drainage, including nonriverine Pleistocene high terraces, as well as broad, low flats, small to large depressions, and areas along small, ill-defined drainages (locally known as "slashes" in Louisiana). In Louisiana, some soils on which this vegetation is found include the Gilbert and Calhoun series which support "Wet Hardwood Flatwoods" (LNHP 2004). In Arkansas, Calhoun soils are sometimes associated with flatwoods. Fire was originally a very important influence on vegetation composition and structure in these seasonally dry examples, but today is often of limited importance. Fire was a negligible influence in others, such as the hardwood flats of North Carolina.

Geographic Range: Vegetation of this group is found in the Atlantic and Gulf coastal plains from southeastern Virginia to Texas, extending down the Florida peninsula, and west to Arkansas, southeastern Oklahoma, and the Missouri "bootheel." It is abundant in the Embayed Region of northeastern North Carolina and southeastern Virginia (south of the James River), and widespread in the West Gulf Coastal Plain.

Nations: US

States/Provinces:  AL, AR, FL?, GA, LA, MD, MO?, MS, NC, OK, SC, TX, VA




Confidence Level: Moderate

Confidence Level Comments: No Data Available

Grank: GNR

Greasons: No Data Available


Concept Lineage: No Data Available

Predecessors: No Data Available

Obsolete Names: No Data Available

Obsolete Parents: No Data Available

Synonomy: ? Flatland Hardwood Forest (Marks and Harcombe 1981)
< IIA10e. Nonriverine Wet Hardwood Forest (Allard 1990)
< Nonriverine Wet Hardwood Forest (Schafale and Weakley 1990)
? Willow Oak - Water Oak - Diamondleaf (Laurel) Oak: 88 (Eyre 1980)

Concept Author(s): M.P. Schafale and A.S. Weakley (1990)

Author of Description: M. Pyne

Acknowledgements: We have incorporated information from T. Foti relative to the West Gulf Coastal Plain.

Version Date: 05-13-15

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