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CEGL003746 (Cephalanthus occidentalis) / Dulichium arundinaceum - (Polygonum hydropiperoides, Glyceria acutiflora, Proserpinaca palustris) Marsh

Type Concept Sentence: No Data Available


Common (Translated Scientific) Name: (Common Buttonbush) / Threeway Sedge - (Swamp Smartweed, Creeping Mannagrass, Marsh Mermaidweed) Marsh

Colloquial Name: Central Appalachian Montane Depression Pond (Threeway Sedge - Buttonbush Type)

Hierarchy Level:  Association

Type Concept: This community occurs in mostly seasonally flooded (rarely semipermanently flooded) depression ponds in the Central Appalachian, Ridge and Valley, and Northern Blue Ridge provinces. Habitats are thought to have formed by solution or collapse of underlying bedrock strata. Occurrences range from 470 m (1540 feet) to over 900 m (3000 feet) elevation. Topographic position varies from ridge crests to mountain-foot alluvial. Ridge and Valley ponds developed on sideslope benches resulting from landsliding and catastrophic slope failure are often referred to as "sag ponds." Ponds developed by solution and collapse of carbonate rocks underlying acidic alluvial and colluvial materials eroded from the Blue Ridge and deposited along the eastern edge of the Great Valley of Virginia in massive alluvial fans are often referred to as "Shenandoah Valley sinkhole ponds." These topographically variable wetlands range from seasonally to semipermanently flooded, with water levels maintained by continually perched groundwater or seepage inputs. Overall hydrologic regime is best characterized as seasonally flooded, but two or three of the ponds are almost always flooded year-round. Substrates are generally organic, and the surficial peat in which emergent plants are rooted is extremely acidic. Vegetation varies from entirely herbaceous to a fairly dense shrubland dominated by Cephalanthus occidentalis. Many of the ponds are primarily herbaceous but have outer zones, discrete patches, or islands of shrubs within them. Where present, Cephalanthus occurs in variable abundance, often being limited to marginal zones or specific areas of the pond. In the typical, seasonally flooded ponds, the herb layer is typically dense and dominated by Dulichium arundinaceum. The most frequent and characteristic herbaceous associates are Glyceria acutiflora, Polygonum hydropiperoides, and Proserpinaca palustris. Some ponds have a deep center that is dominated by a zone of Nuphar advena. Several other forbs and graminoids, including Bidens discoidea, Carex canescens ssp. disjuncta, Carex lasiocarpa var. americana, Carex lupulina, Carex stricta, Carex vesicaria, Glyceria septentrionalis, Glyceria canadensis, Glyceria obtusa, Scirpus ancistrochaetus, Schoenoplectus torreyi, Triadenum virginicum, Triadenum fraseri, and Utricularia spp. are inconstant but locally patch-dominant. Semipermanently flooded ponds tend to be floristically depauperate and contain a high proportion of floating or submersed aquatic species, including Torreyochloa pallida, Eleocharis palustris, Glyceria acutiflora, Potamogeton spp., Proserpinaca palustris, and Utricularia spp.

Diagnostic Characteristics: No Data Available

Rationale for Nominal Species or Physiognomic Features: No Data Available

Classification Comments: This type was based on quantitative analysis of all montane depression wetlands in Virginia (Fleming and Patterson 2009b). Excellent examples occur in the Maple Flats pond complex in the Shenandoah Valley (Augusta County, Virginia), and the type also appears to be the major type of ridge crest pond in the Ridge and Valley province (Fleming 1998). Other examples include Browns Pond (Warm Springs District, George Washington National Forest), Potts Mountain Pond (New River Ranger District, Jefferson National Forest), and Peters Mountain Pond (James River District, George Washington National Forest).

Similar NVC Types: No Data Available
note: No Data Available

Physiognomy and Structure: No Data Available

Floristics: Vegetation varies from entirely herbaceous to a fairly dense shrubland dominated by Cephalanthus occidentalis. Many of the ponds are primarily herbaceous but have outer zones, discrete patches, or islands of shrubs within them. Cephalanthus occurs in variable abundance, often being limited to marginal zones or specific areas of the pond. Other woody species occurring infrequently are Ilex verticillata, Decodon verticillatus, and Heteromeles arbutifolia (= Photinia arbutifolia). In the typical, seasonally flooded ponds, the herb layer is typically dense and dominated by Dulichium arundinaceum. The most frequent and characteristic herbaceous associates are Glyceria acutiflora, Polygonum hydropiperoides, and Proserpinaca palustris. Some ponds have a deep center that is dominated by a zone of Nuphar advena (= Nuphar lutea ssp. advena). Several other forbs and graminoids, including Bidens discoidea, Carex canescens ssp. disjuncta, Carex lasiocarpa var. americana, Carex lupulina, Carex stricta, Carex vesicaria, Glyceria septentrionalis, Glyceria canadensis, Glyceria obtusa, Scirpus ancistrochaetus, Schoenoplectus torreyi (= Scirpus torreyi), Triadenum virginicum, Triadenum fraseri, and Utricularia spp., are inconstant but locally patch-dominant. Additional low-cover herbaceous associations include Juncus canadensis, Leersia oryzoides, Potamogeton spp., Eleocharis acicularis, Panicum verrucosum, Boehmeria cylindrica, Ludwigia palustris, and many others. Semipermanently flooded ponds tend to be floristically depauperate and contain a high proportion of floating or submersed aquatic species, including Torreyochloa pallida, Eleocharis palustris (= Eleocharis smallii), Glyceria acutiflora, Potamogeton spp., Proserpinaca palustris, and Utricularia spp. Species richness of 14 plot-sampled stands ranges from 7 to 17 taxa per 100 m2 (mean = 11).

Dynamics:  No Data Available

Environmental Description:  Habitats are depression ponds thought to have formed by solution or collapse of underlying bedrock strata. Occurrences range from 470 m (1540 feet) to over 900 m (3000 feet) elevation. Topographic position varies from ridge crests to mountain-foot alluvial fans (Fleming and Van Alstine 1999). Ridge and Valley ponds developed on sideslope benches resulting from landsliding and catastrophic slope failure are often referred to as "sag ponds" (Frampton 1995, Harbor 1996). Ponds developed by solution and collapse of carbonate rocks underlying acidic alluvial and colluvial materials eroded from the Blue Ridge and deposited along the eastern edge of the Great Valley of Virginia in massive alluvial fans are often referred to as "Shenandoah Valley sinkhole ponds." These topographically variable wetlands range from seasonally to semipermanently flooded, with water levels maintained by continually perched groundwater or seepage inputs. Overall hydrologic regime is best characterized as seasonally flooded, but two or three of the ponds are almost always flooded year-round. Substrates are generally organic, and the surficial peat in which emergent plants are rooted is extremely acidic. C-14 dating of material extracted from a semipermanently flooded, peat-filled depression on the southeastern flank of Peters Mountain (Alleghany County) indicated a late Pleistocene origin for this wetland (Frampton 1995).

Geographic Range: This community type is known from a small number of natural mountain ponds in western Virginia, mostly in the Ridge and Valley province. It may also occur in southern Pennsylvania.

Nations: US

States/Provinces:  PA?, VA, WV




Confidence Level: Low

Confidence Level Comments: No Data Available

Grank: G1

Greasons: No Data Available


Concept Lineage: This type was created by the merger of the former CEGL007854 - Cephalanthus occidentalis / Dulichium arundinaceum Shrubland, the former CEGL007855 - Cephalanthus occidentalis / Torreyochloa pallida Shrubland (Provisional), and the former Virginia Ridge and Valley portion of CEGL006242 - Cephalanthus occidentalis / Polygonum hydropiperoides - Panicum verrucosum Shrubland.

Predecessors: No Data Available

Obsolete Names: No Data Available

Obsolete Parents: No Data Available

Synonomy: = (Cephalanthus occidentalis) / Dulichium arundinaceum - (Polygonum hydropiperoides, Glyceria acutiflora, Proserpinaca palustris) Herbaceous Vegetation (Fleming and Patterson 2009b)
> Cephalanthus occidentalis / Dulichium arundinaceum Shrubland (Fleming and Coulling 2001)
> Cephalanthus occidentalis / Dulichium arundinaceum Shrubland (Fleming and Moorhead 2000)
> Cephalanthus occidentalis / Polygonum hydropiperoides - Glyceria acutiflora - Proserpinaca palustris Shrubland (Fleming and Coulling 2001)
> Cephalanthus occidentalis / Proserpinaca palustris - Polygonum hydropiperoides community (Fleming and Van Alstine 1999)
> Cephalanthus occidentalis / Torreyochloa pallida Shrubland [Provisional] (Fleming and Coulling 2001)
> Cephalanthus occidentalis / Torreyochloa pallida community (Fleming and Van Alstine 1999)

Concept Author(s): G. Fleming and K. Patterson (2009b)

Author of Description: G.P. Fleming

Acknowledgements: No Data Available

Version Date: 02-17-12

  • Buhlmann, K. A., J. C. Mitchell, and L. R. Smith. 1999. Descriptive ecology of the Shenandoah Valley sinkhole pond system in Virginia. Banisteria 13:23-51.
  • Fleming, G. P., K. D. Patterson, and K. Taverna. 2017. The natural communities of Virginia: A classification of ecological community groups and community types. Third approximation. Version 3.0. Virginia Department of Conservation and Recreation, Division of Natural Heritage, Richmond, VA. [http://www.dcr.virginia.gov/natural-heritage/natural-communities/]
  • Fleming, G. P., and K. D. Patterson. 2009b. Classification of selected Virginia montane wetland groups. In-house analysis, December 2009. Virginia Department of Conservation and Recreation, Division of Natural Heritage, Richmond.
  • Fleming, G. P., and N. E. Van Alstine. 1999. Plant communities and floristic features of sinkhole ponds and seepage wetlands in southeastern Augusta County, Virginia. Banisteria 13:67-94.
  • Fleming, G. P., and P. P. Coulling. 2001. Ecological communities of the George Washington and Jefferson national forests, Virginia. Preliminary classification and description of vegetation types. Virginia Department of Conservation and Recreation, Division of Natural Heritage, Richmond, VA. 317 pp.
  • Fleming, G. P., and W. H. Moorhead, III. 2000. Plant communities and ecological land units of the Peter''s Mountain area, James River Ranger District, George Washington and Jefferson national forests, Virginia. Natural Heritage Technical Report 00-07. Virginia Department of Conservation and Recreation, Division of Natural Heritage, Richmond. Unpublished report submitted to the USDA Forest Service. 195 pp. plus appendices.
  • Frampton, J .B. 1995. Quaternary mass movements in Alleghany, Rockbridge, and Botetourt counties, Virginia. Unpublished senior research thesis, Washington and Lee University, Lexington, VA.
  • Harbor, D. J. 1996. Classification and mapping of geomorphology in the Peters Mountain, Laurel Fork, and Upper Piney River areas, George Washington National Forest. Unpublished report to the USDA Forest Service. Washington and Lee University, Lexington, VA. 25 pp.
  • Rawinski, T. J. 1997. Vegetation ecology of the Grafton Ponds, York County, Virginia, with notes on waterfowl use. Natural Heritage Technical Report 97-10. Virginia Department of Conservation and Recreation, Division of Natural Heritage, Richmond. 42 pp. plus appendix.
  • Southeastern Ecology Working Group of NatureServe. No date. International Ecological Classification Standard: International Vegetation Classification. Terrestrial Vegetation. NatureServe, Durham, NC.
  • WVNHP [West Virginia Natural Heritage Program]. No date. Unpublished data. West Virginia Natural Heritage Program, Elkins.
  • Zimmerman, E. A. 2011y. Pennsylvania Natural Heritage Program. Rice Cutgrass - Bulrush Marsh Factsheet. [http://www.naturalheritage.state.pa.us/Community.aspx?=30028] (accessed February 14, 2012)
  • Zimmerman, E. A., T. Davis, M. A. Furedi, B. Eichelberger, J. McPherson, S. Seymour, G. Podniesinski, N. Dewar, and J. Wagner, editors. 2012. Terrestrial and palustrine plant communities of Pennsylvania. Pennsylvania Natural Heritage Program, Pennsylvania Department of Conservation and Natural Resources, Harrisburg. [http://www.naturalheritage.state.pa.us/Communities.aspx]