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CEGL004800 Nyssa biflora - Alnus serrulata / Carex (mitchelliana, atlantica ssp. capillacea) - Glyceria obtusa Floodplain Scrub

Type Concept Sentence: No Data Available


Common (Translated Scientific) Name: Swamp Tupelo - Hazel Alder / (Mitchell''s Sedge, Prickly Bog Sedge) - Atlantic Mannagrass Floodplain Scrub

Colloquial Name: Sandhill Beaver Pond Successional Scrub

Hierarchy Level:  Association

Type Concept: This community occurs as a long-lasting successional stage in drained beaver ponds in sandy areas with little sediment input, where deep organic deposits accumulate. They are presently known only from the Fall-line Sandhills region of North Carolina and possibly South Carolina. The abundant seepage and limited overbank flooding of floodplains in this region may be crucial to their development. The vegetation is a fine mosaic of shrub-dominated hummocks and herb-dominated hollows. It may include a sparse canopy of Nyssa biflora and Acer rubrum, consisting of young and stressed trees that have survived the impoundment. The shrub layer is most often dominated by Alnus serrulata and tree saplings. The herb layer is moderate to dense. Dominants in patches or in some sites include Carex mitchelliana, Carex atlantica ssp. capillacea, Carex stricta, Glyceria obtusa, Leersia oryzoides, Dichanthelium scabriusculum, Peltandra virginica, and Orontium aquaticum. Other frequent herbs include Dichanthelium dichotomum, Dulichium arundinaceum, Triadenum virginicum, Lycopus cokeri, Eupatorium resinosum, Carex atlantica, and Woodwardia areolata. Sphagnum is often abundant and extensive.

Diagnostic Characteristics: No Data Available

Rationale for Nominal Species or Physiognomic Features: No Data Available

Classification Comments: The classification of southern beaver pond communities remains tentative. This community appears to be distinct from those of typical brownwater or blackwater floodplains of the Coastal Plain and to depend on the unusual hydrological or soil characteristics of the Sandhills region. A study of these communities at Fort Bragg (Hall 2005) identified a number of types of successional mires. This association represents a coarser interpretation, covering all of the groups called mires. Additional splitting may be warranted after further study to determine whether these patterns are applicable across the range of these communities. It is not presently known how these types relate to beaver pond communities outside of Fort Bragg. In Hall''s (2005) analysis, the most marked variant was communities with herb layers strongly dominated by Carex stricta, with all other plants relegated to low abundance. Those dominated by Dichanthelium scabriusculum and Dichanthelium dichotomum are also distinct from the more typical examples.

Similar NVC Types: No Data Available
note: No Data Available

Physiognomy and Structure: No Data Available

Floristics: The vegetation is a fine mosaic of shrub-dominated hummocks and herb-dominated hollows. It may include a sparse canopy of Nyssa biflora and Acer rubrum, consisting of young and stressed trees that have survived the impoundment. The shrub layer generally has 30-60% cover and is most often dominated by Alnus serrulata and tree saplings. Other common shrubs include Viburnum nudum, Arundinaria tecta, Vaccinium formosum, and less abundantly, Cyrilla racemiflora, Lyonia lucida, or Clethra alnifolia. The herb layer is moderate to dense. Sphagnum is often abundant and extensive. Dominants in patches or in some sites include Carex mitchelliana, Carex atlantica ssp. capillacea (= Carex howei), Carex stricta, Glyceria obtusa, Leersia oryzoides, Dichanthelium scabriusculum, Peltandra virginica, and Orontium aquaticum. Other frequent herbs include Dichanthelium dichotomum, Dulichium arundinaceum, Triadenum virginicum, Lycopus cokeri, Eupatorium resinosum, Carex atlantica, and Woodwardia areolata.

Dynamics:  This is a successional community of moderate to long duration, occurring after the drainage of beaver ponds or similar small artificial impoundments on small streams. Stands probably start as open water (~Nuphar advena - Nymphaea odorata Aquatic Vegetation (CEGL002386)$$) or marsh (~Orontium aquaticum - Schoenoplectus (etuberculatus, subterminalis) - Eriocaulon decangulare - Juncus trigonocarpus Marsh (CEGL007860)$$). If a pond drains abruptly, a brief stage of dominance by relict pond species, such as Sparganium americanum, and short-lived opportunistic species, such as Rhynchospora spp. and Erechtites hieraciifolius, may be followed by establishment of this vegetation. If a pond is abandoned and the dam deteriorates gradually, this community may develop slowly from the open-water pond. Trees gradually invade and grow in this community, gradually forming a closed canopy that eliminates much of the distinctive herb layer. These communities eventually succeed to ~Nyssa biflora - Liriodendron tulipifera - Pinus (serotina, taeda) / Lyonia lucida Swamp Forest (CEGL004734)$$. The length of their persistence is not known, but it appears to be long enough to warrant recognition as an association and as a target for conservation action. While all beaver ponds must go through some form of herbaceous successional vegetation, the distinctive vegetation and perhaps the persistence of this community appears to be confined to the Sandhills region and to depend on the distinctive conditions there.

Environmental Description:  Stands occur in drained beaver ponds along creeks with very little sediment transport, where deep muck deposits develop. They are presently known only from the Fall-line Sandhills region, and the abundant seepage and limited overbank flooding of floodplains in this region may be crucial to their development. These sites begin with the drainage of an open-water beaver pond. Pre-existing organic-rich soils combined with limited sediment movement and wetness promote development of deep, soft muck in pond beds and slow succession back to swamp forest. These sites are permanently saturated, often shallowly flooded, and extremely acidic and nutrient-poor.

Geographic Range: This association is known from the Fall-line Sandhills region of North Carolina and possibly South Carolina.

Nations: US

States/Provinces:  NC, SC?




Confidence Level: Low - Poorly Documented

Confidence Level Comments: No Data Available

Grank: G2?

Greasons: No Data Available


Concept Lineage: No Data Available

Predecessors: No Data Available

Obsolete Names: No Data Available

Obsolete Parents: No Data Available

Synonomy: No Data Available

Concept Author(s): S.P. Hall (2005b)

Author of Description: M.P. Schafale

Acknowledgements: No Data Available

Version Date: 03-18-09

  • Hall, S.P. 2005b. A quantitative analysis and classification of the habitats of Neonympha mitchellii francisci at Fort Bragg and Camp MacKall. Report to the Endangered Species Branch, Fort Bragg, NC. North Carolina Natural Heritage Program, Office of Planning and Conservation, Department of Environment and Natural Resources, Raleigh.
  • Nelson, J. B. 1986. The natural communities of South Carolina: Initial classification and description. South Carolina Wildlife and Marine Resources Department, Division of Wildlife and Freshwater Fisheries, Columbia, SC. 55 pp.
  • Schafale, M. P. 2012. Classification of the natural communities of North Carolina, 4th Approximation. North Carolina Department of Environment, Health, and Natural Resources, Division of Parks and Recreation, Natural Heritage Program, Raleigh.
  • Southeastern Ecology Working Group of NatureServe. No date. International Ecological Classification Standard: International Vegetation Classification. Terrestrial Vegetation. NatureServe, Durham, NC.