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CEGL006597 Carex scabrata - Viola cucullata / Plagiomnium ciliare Seepage Meadow

Type Concept Sentence: No Data Available


Common (Translated Scientific) Name: Eastern Rough Sedge - Marsh Blue Violet / Toothed Leafy Moss Seepage Meadow

Colloquial Name: Allegheny Mountains Eastern Rough Sedge / Wavy-leaf Moss Seep

Hierarchy Level:  Association

Type Concept: This linear, sloping forested seep occurs on saturated, temporarily flooded, and semipermanently flooded soils in headwater basins of the Allegheny Mountains region of West Virginia, at elevations between 780 and 1300 m. It occupies gently to steeply sloping drainages (1.5-20°) in upland forest, typically occurring as small, discontinuous patches that repeat across a mountainside. It is fed primarily by seepage and springs from the adjacent upland forest. Microtopography is characterized by a complex of tip-up mounds, buttressed roots, and downed wood over large and small rocks. Bedrock is typically shale or sometimes sandstone. Soils are moderately to poorly drained and stony, with variable texture ranging from muck to silt loam to sandy loam. The canopy is open and dominated by Betula alleghaniensis var. alleghaniensis with an herbaceous layer dominated by Carex scabrata, Viola cucullata, and the characteristic bryophyte Plagiomnium ciliare. The canopy often includes low cover by Acer saccharum var. saccharum, Picea rubens, and Fagus grandifolia. Most trees are not rooted in the seep, but rather overhang and shade the seep from the edges. The subcanopy and shrub layers are sparse. The herbaceous layer is dominated by Carex scabrata, Viola cucullata, Laportea canadensis, Tiarella cordifolia, and Glyceria melicaria. Other herbaceous species with high constancy include Dryopteris intermedia, Saxifraga micranthidifolia, Cardamine diphylla, Chrysosplenium americanum, Poa alsodes, Symphyotrichum prenanthoides, Packera aurea, Thelypteris noveboracensis, Cardamine pensylvanica, Arisaema triphyllum, Monarda didyma, and Oxalis montana. Nonvascular plants are dominated by Plagiomnium ciliare with lesser amounts of Brachythecium rivulare, Thuidium delicatulum, and Rhizomnium appalachianum. The indicator species that help to distinguish this community from others within the forested seep physiognomy in West Virginia are Carex scabrata, Laportea canadensis, Tiarella cordifolia, Acer saccharum var. saccharum, Brachythecium rivulare, Cardamine diphylla, Carex leptonervia, and Plagiomnium ciliare. Mean species richness of all vascular plants and any nonvascular plants with cover >1% is 32 taxa per 400 m2, with about 15% of the diversity in the bryophyte layer.

Diagnostic Characteristics: No Data Available

Rationale for Nominal Species or Physiognomic Features: No Data Available

Classification Comments: Seven plots represent this type, which was classified as part of a 2004-2006 study of high-elevation wetlands in West Virginia''s Allegheny Mountains region (Byers et al. 2007). The type clusters and ordinates well and has been sampled throughout its range in West Virginia. Similar vegetation is very likely in the High Allegheny region of Virginia, but may be better interpreted there as an inclusion in a small stream alluvial forest.

Similar NVC Types: No Data Available
note: No Data Available

Physiognomy and Structure: No Data Available

Floristics: This linear forested seep occurs in the Allegheny Mountains region of West Virginia. It is characterized by an open canopy of Betula alleghaniensis var. alleghaniensis with an herbaceous layer dominated by Carex scabrata, Viola cucullata, and the characteristic bryophyte Plagiomnium ciliare. The canopy often includes low cover by Acer saccharum var. saccharum, Picea rubens, and Fagus grandifolia. Mean canopy cover is 40%. Most trees are not rooted in the seep, but rather overhang and shade the seep from the edges. Trees that are rooted within the seep tend to grow on moss-covered hummocks with buttressed roots. The subcanopy and shrub layers are sparse, totaling about 8% cover, and with a composition similar to that of the canopy. The herbaceous layer averages 60% cover and is dominated by Carex scabrata, Viola cucullata, Laportea canadensis, Tiarella cordifolia, and Glyceria melicaria. Other herbaceous species with high constancy include Dryopteris intermedia, Saxifraga micranthidifolia, Cardamine diphylla, Chrysosplenium americanum, Poa alsodes, Symphyotrichum prenanthoides, Packera aurea, Thelypteris noveboracensis, Cardamine pensylvanica, Arisaema triphyllum, Monarda didyma, and Oxalis montana. Nonvascular plants average 25% cover and are dominated by Plagiomnium ciliare with lesser amounts of Brachythecium rivulare, Thuidium delicatulum, and Rhizomnium appalachianum. The indicator species that help to distinguish this community from others within the forested seep physiognomy in West Virginia are Carex scabrata, Laportea canadensis, Tiarella cordifolia, Acer saccharum var. saccharum, Brachythecium rivulare, Cardamine diphylla, Carex leptonervia, and Plagiomnium ciliare. Mean species richness of all vascular plants and any nonvascular plants with cover >1% is 32 taxa per 400 m2, with about 15% of the diversity in the bryophyte layer (Byers et al. 2007).

Dynamics:  This is a small-patch, sloping forested seep community. It is fed primarily by seepage and springs from the adjacent upland forest. Nutrient cycling occurs from the decay of fallen trees, litter, and herbaceous vegetation.

Environmental Description:  This linear, sloping forested seep occurs on saturated, temporarily flooded, and semipermanently flooded soils in headwater basins of the Allegheny Mountains region of West Virginia, at elevations between 780 and 1300 m. It occupies gently to steeply sloping drainages (1.5-20°) in upland forest, typically occurring as small, discontinuous patches that repeat across a mountainside. Microtopography is characterized by a complex of tip-up mounds, buttressed roots, and downed wood over large and small rocks. Bedrock is typically shale or sometimes sandstone. Soils are moderately to poorly drained and stony, with variable texture ranging from muck to silt loam to sandy loam. Hydric soil indicators include histic epipedon, 2 cm muck, sandy mucky mineral, loamy gleyed matrix, and depleted matrix. Soil pH averages 4.8, but pore water pH is significantly higher, ranging from 5.8-7.0, with an average of 6.6 (n=4). Electrical conductivity of the pore water is low, averaging 48 micromhos/cm (n=4). Soil chemistry is characterized by high B, Ca, Cu, Fe, Mg, Mn, Na, P, S, Zn; moderate K and exchangeable nitrogen; and low Al, organic matter, and total exchange capacity (n=4). The unvegetated surface is predominantly litter, with an average of 10% rocks, 6% downed wood and 5% standing or flowing water (Byers et al. 2007).

Geographic Range: This community is known from the Allegheny Mountains region of West Virginia, at elevations between 780 and 1300 m.

Nations: US

States/Provinces:  VA, WV




Confidence Level: Low

Confidence Level Comments: No Data Available

Grank: G3

Greasons: No Data Available


Concept Lineage: No Data Available

Predecessors: No Data Available

Obsolete Names: No Data Available

Obsolete Parents: No Data Available

Synonomy: = Betula alleghaniensis var. alleghaniensis / Carex scabrata - Viola cucullata canadensis / Plagiomnium ciliare sloping linear forest seep (Byers et al. 2007)

Concept Author(s): E.A. Byers et al. (2007)

Author of Description: E.A. Byers

Acknowledgements: No Data Available

Version Date: 04-03-07

  • Byers, E. A., J. P. Vanderhorst, and B. P. Streets. 2007. Classification and conservation assessment of high elevation wetland communities in the Allegheny Mountains of West Virginia. West Virginia Natural Heritage Program, West Virginia Division of Natural Resources, Elkins.
  • Eastern Ecology Working Group of NatureServe. No date. International Ecological Classification Standard: International Vegetation Classification. Terrestrial Vegetation. NatureServe, Boston, MA.
  • Fleming, G. P., K. D. Patterson, and K. Taverna. 2017. The natural communities of Virginia: A classification of ecological community groups and community types. Third approximation. Version 3.0. Virginia Department of Conservation and Recreation, Division of Natural Heritage, Richmond, VA. [http://www.dcr.virginia.gov/natural-heritage/natural-communities/]
  • WVNHP [West Virginia Natural Heritage Program]. No date. Unpublished data. West Virginia Natural Heritage Program, Elkins.