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CEGL006592 Abies balsamea - Picea rubens / Danthonia compressa - Lycopodium spp. / Sphagnum spp. Swamp Forest

Type Concept Sentence: No Data Available


Common (Translated Scientific) Name: Balsam Fir - Red Spruce / Flattened Oatgrass - Clubmoss species / Peatmoss species Swamp Forest

Colloquial Name: Balsam Fir - Red Spruce / Oatgrass / Peatmoss Swamp Forest

Hierarchy Level:  Association

Type Concept: This acidic conifer woodland or forested swamp occurs on moist to saturated soils in headwater basins in the Allegheny Mountains region of West Virginia, at elevations between 960 and 1130 m. It is a small-patch community fed by slow seepage and rainfall. It occupies gently sloping land (1-10° slopes) along small headwater streams and in mixed wetland mosaics, often at the base of upland slopes. Microtopography is characterized by irregular hummocks formed over tree roots, tip-up mounds, nurse logs, and decaying wood. Soils are moderately to poorly drained loamy soils with mottling in the upper 20 cm and occasional gleyed horizons. The canopy is open to closed and dominated by Abies balsamea and Picea rubens, with occasional presence of Prunus serotina var. serotina. The subcanopy is dominated by Abies balsamea with smaller amounts of Picea rubens, Acer rubrum, and Betula alleghaniensis var. alleghaniensis. The tall-shrub layer is also dominated by Abies balsamea and Picea rubens. The short-shrub layer contains regenerating canopy species and locally abundant Vaccinium myrtilloides and Hypericum densiflorum. The herbaceous ground layer is fairly diverse, typically including 25-50 species. The most abundant species are Danthonia compressa, Lycopodium obscurum, and Rubus hispidus. Other herbaceous species with high constancy include the regenerating canopy species, Lycopodium clavatum, Carex folliculata, Dennstaedtia punctilobula, Pteridium aquilinum, and Oclemena acuminata. Nonvascular plants are dominated by Sphagnum and Polytrichum species; Leucobryum glaucum is also common. The community is characterized by a number of diagnostic species that highlight the slightly drier habitat of this swamp type and include Carex debilis, Crataegus spp., Lycopodium clavatum, Lycopodium digitatum, Mitchella repens, Polytrichum ssp., Prunus serotina var. serotina, and Pteridium aquilinum. Mean species richness of all vascular plants and any nonvascular plants with cover >1% is 41 taxa per 400 m2.

Diagnostic Characteristics: No Data Available

Rationale for Nominal Species or Physiognomic Features: No Data Available

Classification Comments: Twelve plots represent this type, which was classified as part of a 2004-2006 study of high-elevation wetlands in West Virginia''s Allegheny Mountains region (Byers et al. 2007). The type clusters and ordinates well and has been sampled throughout its range in West Virginia. Eleven of the plots were sampled by Ceperley (2002) and classified into two types which are included in this broader type.

Similar NVC Types: No Data Available
note: No Data Available

Physiognomy and Structure: No Data Available

Floristics: This evergreen swamp occurs in scattered high-elevation sites in the Allegheny Mountains of West Virginia. The canopy is open to closed and dominated by Abies balsamea and Picea rubens, with occasional presence of Prunus serotina var. serotina. Mean canopy cover is 30%. Canopy height rarely exceeds 20 m and is generally less than 15 m. The subcanopy averages 19% cover and is dominated by Abies balsamea with smaller amounts of Picea rubens, Acer rubrum, and Betula alleghaniensis var. alleghaniensis. Tsuga canadensis, Amelanchier arborea var. arborea, and Crataegus spp. may be present with very low cover in the subcanopy. The tall-shrub layer averages 31% cover and is also dominated by Abies balsamea and Picea rubens. The short-shrub layer averages 18% cover, with locally abundant Vaccinium myrtilloides and Hypericum densiflorum. Regeneration of canopy species is present in this stratum. The herbaceous ground layer is fairly diverse, with mean 48% cover and typically including 25-50 species. The most abundant species are Danthonia compressa, Lycopodium obscurum, and Rubus hispidus. Other herbaceous species with high constancy include the canopy species, Lycopodium clavatum, Carex folliculata, Dennstaedtia punctilobula, Pteridium aquilinum, and Oclemena acuminata. Nonvascular plants average 17% cover in this community, dominated by Sphagnum and Polytrichum species; Leucobryum glaucum is also common. Indicator species that help to distinguish this community from others within the forest/woodland physiognomy for high-elevation wetlands of the Allegheny Mountains region include Carex debilis, Crataegus spp., Lycopodium clavatum, Lycopodium digitatum, Mitchella repens, Polytrichum ssp., Prunus serotina var. serotina, and Pteridium aquilinum. These indicator species highlight the slightly drier habitat of this swamp community. Mean species richness of all vascular plants and any nonvascular plants with cover >1% is 41 taxa per 400 m2 (Ceperley 2002, Byers et al. 2007).

Dynamics:  This is a small-patch woodland/forest swamp community. It is fed by slow seepage and rainfall. Nutrient cycling occurs from the decay of fallen trees, litter, and herbaceous vegetation.

Environmental Description:  This acidic conifer woodland swamp occurs on moist to saturated soils in headwater basins of the Allegheny Mountains region of West Virginia, at elevations of 960-1130 m above sea level. It occurs on Mississippian and Pennsylvanian bedrock, usually on Mauch Chunk shale or Greenbrier limestone, and occasionally on the more acidic Pottsville or Conemaugh formations. It occupies gently sloping land (1-10° slopes) along small headwater streams, in mixed wetland mosaics, often at the base of upland slopes. This type is at the drier end of the wetland range. Microtopography is characterized by irregular hummocks formed over tree roots, tip-up mounds, nurse logs, and decaying wood. Anthropogenic disturbance includes historic logging and fires around the year 1900. Current fluctuations in hydrologic regime occur as a result of beaver activity, but do not appear to immediately threaten this community. Most Abies balsamea show slight to severe damage from Adelges piceae (balsam woolly adelgid), including bark infestation and gout damage; however, this community type also includes the best multi-aged disease-free stand of Abies balsamea in West Virginia, where the species is at the southernmost edge of its range. Combined with excessive deer herbivory, the adelgid damage may be severe enough to eliminate balsam fir and change the dominant species composition of this community in the future. Soils are moderately to poorly drained loamy soils with mottling in the upper 20 cm and occasional gleyed horizons. Hydric soil indicators include sandy redox, stripped matrix, depleted matrix, and redox depressions. Soil chemistry is characterized by high Al, Fe, H, N, and total exchange capacity; moderate K, P, organic matter; and low B, Ca, Cu, Mg, Mn, Na, S, Zn (n=10). The unvegetated surface is predominantly litter, with an average of 3% downed wood and 1% standing water (Ceperley 2002, Byers et al. 2007).

Geographic Range: The known distribution of this community is restricted to the Allegheny Mountains region of West Virginia, at elevations above 960 m. Within this region, the community is known only from Canaan Valley, Blister Run Swamp, and Dolly Sods.

Nations: US

States/Provinces:  WV




Confidence Level: High

Confidence Level Comments: No Data Available

Grank: G2

Greasons: No Data Available


Concept Lineage: No Data Available

Predecessors: No Data Available

Obsolete Names: No Data Available

Obsolete Parents: No Data Available

Synonomy: = Abies balsamea - Picea rubens / Danthonia compressa - Lycopodium spp. / Sphagnum spp. swamp (Byers et al. 2007)
> Abies balsamea - Picea rubens / Vaccinium / Lycopodium hummock wetland (Ceperley 2002)
> Abies balsamea / Pteridium aquilinum hummock wetland (Ceperley 2002)
< Picea rubens - Tsuga canadensis - Abies balsamea swamp forest (Fortney et al. 2005)

Concept Author(s): E.A. Byers et al. (2007)

Author of Description: E.A. Byers

Acknowledgements: No Data Available

Version Date: 03-30-07

  • Byers, E. A., J. P. Vanderhorst, and B. P. Streets. 2007. Classification and conservation assessment of high elevation wetland communities in the Allegheny Mountains of West Virginia. West Virginia Natural Heritage Program, West Virginia Division of Natural Resources, Elkins.
  • Ceperley, L. 2002. Classification of West Virginia and Virginia Abies balsemea communities affected by balsam woolly adelgid (Adelges piceae) infestation. M.S. thesis, Department of Environmental Studies, Antioch New England Graduate School, Antioch University, Keene, NH. 97 pp.
  • Eastern Ecology Working Group of NatureServe. No date. International Ecological Classification Standard: International Vegetation Classification. Terrestrial Vegetation. NatureServe, Boston, MA.
  • Fortney, R. H., S. L. Stephenson, and J. S. Rentch. 2005. Rare plant communities of Canaan Valley, West Virginia, USA. Unpublished manuscript.
  • WVNHP [West Virginia Natural Heritage Program]. No date. Unpublished data. West Virginia Natural Heritage Program, Elkins.