Print Report

CEGL005924 Pinus contorta / Vaccinium scoparium / Xerophyllum tenax Forest

Type Concept Sentence: No Data Available


Common (Translated Scientific) Name: Lodgepole Pine / Grouse Whortleberry / Common Beargrass Forest

Colloquial Name: No Data Available

Hierarchy Level:  Association

Type Concept: This large-patch to matrix lodgepole pine forest occurs in upper subalpine habitats from central Idaho to northern Idaho and west as far as the eastern slope of the Cascade Range in Washington and east to just beyond the Continental Divide in central Montana and southwestern Alberta. In the northern Rockies of Idaho and Montana, this association ranges from 1585 to 2530 m (5200-8300 feet); just east of the Cascades its elevation range is considerably depressed, from 1310 to 1730 m (4300-5680 feet). Sites are generally cool to cold and have deep snow accumulations. It is found on gentle to moderate slopes of all aspects, mostly associated with shedding terrain, such as backslopes and slope shoulders; it also occurs on well-drained benches associated with ridgetops. Parent materials include volcanics, metamorphic rock, and sedimentary. Soils are uniformly well- to excessively drained with textures predominantly sandy loams and loams. Gravel content is usually at least 10% for the upper profile and increases markedly with depth, averaging about twice that of the surface soil. Soils are also very to moderately acidic. The tree canopy is highly variable with regard to structure, ranging from closed at 60-80% canopy cover in many of the northern Idaho stands to extremely to moderately open (25-50% canopy cover) along the Continental Divide in Montana. Compositionally the canopy is strongly dominated by Pinus contorta; usually Abies lasiocarpa, Tsuga mertensiana, or Picea engelmannii are present. A greater diversity of tree species occur on Washington''s examples of this association. The undergrowth may be quite dense with a layer of Vaccinium scoparium (or Vaccinium myrtillus) surrounding tussocks of Xerophyllum tenax over which a sparse cover of Vaccinium membranaceum is superimposed. Tall shrubs, if present, could be characterized as accidentals. Spiraea betulifolia and Lonicera utahensis are consistently scattered in the short-shrub layer. Carex geyeri and Calamagrostis rubescens are the only graminoids regularly present and their cover seldom exceeds 10%. Other than the abundance of the diagnostic Xerophyllum tenax, the forb layer has low cover and is comparatively depauperate with only Arnica latifolia, Anemone piperi, Goodyera oblongifolia, Orthilia secunda, and Viola orbiculata having a consistent presence.

Diagnostic Characteristics: No Data Available

Rationale for Nominal Species or Physiognomic Features: No Data Available

Classification Comments: With the comparatively recent emphasis on developing descriptions of (and keys to) existing vegetation (Grossman et al. 1998), it has yet to be determined at what cover values forest vegetation types will be distinguished one from another when the canopy tree species are mostly seral in nature and have a broad environmental range (broad niche). This contrasts with the approach of Pfister et al. (1977) who recognize Pinus contorta community types only when virtually no representation of other, more shade-tolerant tree species could be found on site. With Pinus contorta, which is exclusively seral except with respect to some subspecies on unusual substrates or atypical environments, researchers in Glacier National Park have taken the position that this very shade-intolerant, stand-replacing fire-adapted species should have several times the cover of the next most abundant canopy species for a Pinus contorta type to be recognized. What is really being indicated by this approach are areas having experienced stand-replacing fires (or similar catastrophic disturbance, e.g., clearcutting). Another approach would be to simply recognize the tree species with the plurality of canopy cover in assigning stands to particular associations treating existing vegetation. The stands representing this type are climax in two different tree series, Abies lasiocarpa and Tsuga mertensiana. That this association in fact occurs in the states and USFS Sections listed derives from this writer''s interpretation of constancy/cover tables of various authors/publications; where Pinus contorta is listed as a major seral species, the inference has been made that stands quite likely occur on the landscape where this species strongly dominates the upper canopy (often observed throughout northern Idaho and western Montana) (S. Cooper pers. comm.).

Similar NVC Types: No Data Available
note: No Data Available

Physiognomy and Structure: No Data Available

Floristics: The tree canopy is highly variable with regard to structure, ranging from closed at 60-80% canopy cover in many of the northern Idaho stands to extremely to moderately open (25-50% canopy cover) along the Continental Divide in Montana (other Montana areas report 60-70% average cover). Compositionally the canopy is strongly dominated by Pinus contorta; it has several times the cover of the next most abundant species, which is usually Abies lasiocarpa, Tsuga mertensiana, or Picea engelmannii. These sites are generally at the cold limits of Pseudotsuga menziesii (which can be frost-stunted) and at the cold limits or largely beyond the range limits of Larix occidentalis (as it occurs in Montana and Idaho). A greater diversity of tree species occur on Washington''s examples of this association. The undergrowth may be quite dense with a layer of Vaccinium scoparium (or Vaccinium myrtillus) surrounding tussocks of Xerophyllum tenax over which a sparse cover of Vaccinium membranaceum is superimposed. Tall shrubs, if present, could be characterized as accidentals. Spiraea betulifolia and Lonicera utahensis are consistently scattered in the short-shrub layer. Carex geyeri and Calamagrostis rubescens are the only graminoids regularly present and their cover seldom exceeds 10%. Other than the abundance of the diagnostic Xerophyllum tenax, the forb layer has low cover and is comparatively depauperate with only Arnica latifolia, Anemone piperi, Goodyera oblongifolia, Orthilia secunda (= Pyrola secunda), and Viola orbiculata having a consistent presence. In terms of potential vegetation, this association is transitional throughout much of its range between ~Abies lasiocarpa - Picea engelmannii / Vaccinium membranaceum / Xerophyllum tenax Forest (CEGL005917)$$ of lower elevations and warmer exposures and ~Abies lasiocarpa - Picea engelmannii / Xerophyllum tenax - Luzula glabrata var. hitchcockii Woodland (CEGL005898)$$ that characterizes some of the highest and snowiest subalpine sites.

Dynamics:  Data from the subalpine zone of northern Idaho indicate that because of natural mortality patterns Pinus contorta canopy domination is not expected to last more than 160-180 years following stand initiation and that many Pinus contorta-dominated stands commence to break-up at around 100 to 120 years (Cooper et al. 1987). Stands of this association are predominantly early-, mid- and late-seral stages succeeding to ~Abies lasiocarpa - Picea engelmannii / Vaccinium scoparium / Xerophyllum tenax Forest (CEGL005914)$$. It has been noted in northern Idaho and western Montana that following disturbance in this type Pinus contorta often succeeds itself; this response is thought to reflect the primary mode of stand initiation which is stand-replacing fire (or clearcutting). Higher incidences of cone serotiny (and thus increased opportunities to reseed burns) are found in regions/ecosystems experiencing a higher degree of stand-replacing fire (Lotan and Perry 1983). This plant association (because of the requirement of Pinus contorta dominance) is expected to be more prevalent in high montane and subalpine environments, where stand-replacing fire is the primary mode of stand initiation. In the more mesic montane environments stand-replacing fire is a less frequent mode of stand initiation and, being very shade-intolerant, Pinus contorta does poorly at establishing in these less-than-full-sunlight conditions; this association is expected to be an uncommon type in the Tsuga heterophylla, Thuja plicata, and Abies grandis series.

Environmental Description:  This large-patch to matrix type occurs in upper subalpine habitats from central Idaho to northern Idaho and west as far as the eastern slope of the Cascade Range in Washington and east to just beyond the Continental Divide in central Montana and southwestern Alberta. In central Idaho this association ranges from 1980 to 2530 m (6500-8300 feet); at increasing latitudes and with less Inland Maritime climatic influence (in west-central Montana) the range is 1830 to 2285 m (6000-7500 feet); within Inland Maritime-influenced northern Idaho the type exhibits a range of 1585 to 2105 m (5200-6900 feet); just east of the Cascades its elevation range is considerably depressed, from 1310 to 1730 m (4300-5680 feet). For the stated regional elevation ranges the majority of the type occurs in the upper one-half to one-third of the distribution. Sites are generally cool to cold and have deep snow accumulations. It is found on gentle to moderate slopes of all aspects, mostly associated with shedding terrain, such as backslopes and slope shoulders; it also occurs on well-drained benches associated with ridgetops. It is infrequently associated with what ostensibly are frost-pocket conditions (at lower elevations within a region). Parent materials include volcanics (andesites, basalts, rhyolite, granite, granodiorite), metamorphic rock (schist, mica schist, quartzite, argillite) and sedimentary (siltstone, sandstone). Soils are uniformly well- to excessively drained with textures ranging widely, but they are mostly coarser, predominantly sandy loams and loams. Gravel content is usually at least 10% for the upper profile (ranging to 40%) and increases markedly with depth, averaging about twice that of the surface soil. Soils are also very to moderately acidic, mean pH values from various studies ranging from 4.5 to 5.2.

Geographic Range: This association is found from the northern portion of the middle Rocky Mountains of Idaho to the northern Rocky Mountains and documented well into the southeastern portion of the Canadian Rockies; it may extend as far west as the west slope of the Cascade Range in Washington, where possibly it is a seral community of ~Abies lasiocarpa - Tsuga mertensiana / Vaccinium scoparium Woodland (CEGL005522)$$ or ~Tsuga mertensiana - (Abies amabilis) / Vaccinium scoparium Woodland (CEGL005582)$$ and north to Jasper and Banff national parks of Canada (Xerophyllum tenax is apparently scarce in this region).

Nations: CA,US

States/Provinces:  AB, ID, MT, WA, WY?




Confidence Level: Low

Confidence Level Comments: No Data Available

Grank: G3G4

Greasons: No Data Available


Concept Lineage: No Data Available

Predecessors: No Data Available

Obsolete Names: No Data Available

Obsolete Parents: No Data Available

Synonomy: >< Abies lasiocarpa / Xerophyllum tenax Habitat Type (Daubenmire and Daubenmire 1968)
>< Abies lasiocarpa / Xerophyllum tenax Habitat Type, Vaccinium membranaceum Phase (Steele et al. 1981)
>< Abies lasiocarpa / Xerophyllum tenax Habitat Type, Vaccinium membranaceum Phase (Pfister et al. 1977)
>< Abies lasiocarpa / Xerophyllum tenax Habitat Type, Vaccinium scoparium Phase (Pfister et al. 1977)
>< Abies lasiocarpa / Xerophyllum tenax Habitat Type, Vaccinium scoparium Phase (Steele et al. 1981)
>< Abies lasiocarpa / Xerophyllum tenax Habitat Type, Vaccinium scoparium Phase (Cooper et al. 1987)
>< Abies lasiocarpa / Xerophyllum tenax Plant Association (Williams et al. 1995) [(small fraction)]
< Picea engelmannii - Abies lasiocarpa / Xerophyllum tenax Habitat Type (Ogilvie 1962)
= Pinus contorta / Vaccinium scoparium / Xerophyllum tenax Forest (Hop et al. 2007)
>< Tsuga mertensiana / Xerophyllum tenax - Vaccinium myrtillus (Lillybridge et al. 1995) [TSME/XETE-VAMY ASSOCIATION CMF131]
>< Tsuga mertensiana / Xerophyllum tenax Habitat Type, Vaccinium scoparium Phase (Cooper et al. 1987)

Concept Author(s): Hop et al. (2007)

Author of Description: S.V. Cooper

Acknowledgements: No Data Available

Version Date: 03-03-04

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