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CEGL005916 Pinus contorta / Clintonia uniflora Forest
Type Concept Sentence: No Data Available
Common (Translated Scientific) Name: Lodgepole Pine / Bride''s Bonnet Forest
Colloquial Name: No Data Available
Hierarchy Level: Association
Type Concept: Broadly distributed throughout the northern Rocky Mountains and adjacent terrain, this large-patch to matrix seral lodgepole pine forest association occupies relatively moist (mesic) and warm to cool sites having free air drainage and lacking frost-pocket conditions. It occurs on slopes of all degrees of steepness and aspect orientation, though it is more likely to occur in predominantly collecting positions. At the dry extreme of its distribution it is more strongly associated with protected positions such as concave slopes, moist depressions in gently sloping plateau areas, stringers along perennial stream bottoms, toeslopes and northeastern aspects. In the north it ranges from 760 to 1585 m (450-5200 feet), whereas to the south it ranges from 1060 to 1710 m (3500-5600 feet). A wide variety of parent materials are represented including those as disparate as granite, limestone, and all manner of glacio-fluvial material. It is also routinely found on ash caps, ranging from 3 to 60 cm in depth. The soil textures are predominantly loams and silt loams; soils typically have less than 15% coarse-fragment content and are well-drained. This mesic, wholly seral association is characterized by Pinus contorta dominating the upper canopy. Other tree species do occur in the overstory but with much less cover, including the seral Larix occidentalis and Pinus monticola as well as those from warmer environments: Pinus ponderosa, Pseudotsuga menziesii, Thuja plicata, and Tsuga heterophylla, and those of colder environments: Abies lasiocarpa, Abies grandis, and Picea engelmannii. The shrub layer may be highly diverse with tall shrubs (e.g., Acer glabrum, Taxus brevifolia, Amelanchier alnifolia), short shrubs (Symphoricarpos albus, Paxistima myrsinites, Rubus parviflorus, Spiraea betulifolia), and dwarf-shrubs (e.g., Chimaphila umbellata, Linnaea borealis, Mahonia repens) abundantly represented. The graminoid component is inconspicuous. The cover of the diagnostic forbs Clintonia uniflora and Tiarella trifoliata is greatest when this type occurs in warmer environments, up to 30% canopy cover. In the colder environments cover of these diagnostics and all forbs is generally less. Other forbs of high constancy are Aralia nudicaulis, Adenocaulon bicolor, Coptis occidentalis, Cornus canadensis, Galium triflorum, Goodyera oblongifolia, Maianthemum stellatum, Osmorhiza berteroi, Orthilia secunda, Thalictrum occidentale, Trillium ovatum, Viola glabella (or Viola canadensis), and Viola orbiculata.
Diagnostic Characteristics: No Data Available
Rationale for Nominal Species or Physiognomic Features: No Data Available
Classification Comments: With the comparatively recent emphasis on developing descriptions of (and keys to) existing vegetation (Grossman et al. 1998), it has yet to be determined at what cover values forest vegetation types will be distinguished one from another when the canopy tree species are mostly, or exclusively, seral in nature and have a broad environmental range (broad niche). Pfister et al. (1977) recognized Pinus contorta community types only when no other, more shade-tolerant tree species could be found on site. With Pinus contorta, which is exclusively seral, except with respect to some subspecies on unusual substrates or atypical environments, researchers in Glacier National Park took the position that this very shade-intolerant, stand-replacing fire-adapted species should have several times the cover of the next most abundant canopy species for a Pinus contorta type to be recognized. What is really being indicated by this second approach are areas that have experienced stand-replacing fires (or similar catastrophic disturbance, e.g., clearcutting). Another approach when treating existing vegetation could simply be to recognize the plurality of cover in assigning stands to particular associations. The stands representing this type are climax in a number of different tree series, including Abies lasiocarpa, Abies grandis, Tsuga heterophylla, Tsuga mertensiana, and Thuja plicata. That this association in fact occurs in the states and USFS Sections listed derives from this writer''s interpretation of constancy/cover tables of various authors/publications; where Pinus contorta is listed as a major seral species, the inference has been made that stands quite likely occur on the landscape where this species strongly dominates the upper canopy (often observed throughout northern Idaho and western Montana) (S. Cooper pers. comm.). It should also be noted that this type is probably less common in zones where Thuja plicata, Tsuga heterophylla, and Abies amabilis are the climax dominants. This is because following disturbance in these zones the climax trees are quick to reclaim the site, i.e., they comprise a significant cover on the earliest forested successional stages (Pinus contorta may seldom be a major seral component on these sites and thus the association is rare as well), and Pinus contorta is favored by stand-replacing fire which is uncommon in these mesic forests. Note that the Aralia nudicaulis Phase of the Abies lasiocarpa - Picea engelmannii / Clintonia uniflora Habitat Type (Pfister et al. 1977, Steele et al. 1981, Cooper et al. 1987) has been combined with the Clintonia uniflora Phase because it could not be established that it consistently defined unique environments. Thus, all of the recently defined, seral tree-dominated associations that had been subsumed in the Abies lasiocarpa - Picea engelmannii / Clintonia uniflora Habitat Type are somewhat more broadly defined.
Similar NVC Types: No Data Available
note: No Data Available
Physiognomy and Structure: No Data Available
Floristics: This mesic, wholly seral association is characterized by Pinus contorta dominating the upper canopy, by definition having three times the cover of other canopy tree species; other tree species do occur in the overstory but with much less cover, including both other species considered almost exclusively seral (Larix occidentalis and Pinus monticola) and those capable of functioning as both seral and climax species, including those from warmer environments, Pinus ponderosa, Pseudotsuga menziesii, Thuja plicata, and Tsuga heterophylla and those of colder environments, Abies lasiocarpa, Abies grandis, and Picea engelmannii. The shrub layer may be highly diverse with tall shrubs (e.g., Acer glabrum, Taxus brevifolia, Amelanchier alnifolia), short shrubs (Symphoricarpos albus, Paxistima myrsinites, Rubus parviflorus, Spiraea betulifolia), and dwarf-shrubs (e.g., Chimaphila umbellata, Linnaea borealis, Mahonia repens) abundantly represented. Often one of the forenamed short shrubs will be dominant; historical accident suffices for an explanation of this shifting dominance until such time as a thorough analysis is undertaken. The graminoid component is inconspicuous with no one species exhibiting high constancy, though Bromus vulgaris, Bromus ciliatus, and Calamagrostis rubescens are more consistently present and with greater cover than other graminoids. The cover of the diagnostic forbs Clintonia uniflora and Tiarella trifoliata is greatest when this type occurs in the zones potentially dominated by Thuja plicata and Tsuga heterophylla, up to 30% canopy cover (can even be dominant forbs), whereas in the colder environments characterized by Abies lasiocarpa, Abies grandis and Picea engelmannii, cover of these diagnostics and all forbs is generally less. Other forbs of high constancy, at least in some portion of this association''s considerable range, are Aralia nudicaulis, Adenocaulon bicolor, Coptis occidentalis, Cornus canadensis, Galium triflorum, Goodyera oblongifolia, Maianthemum stellatum, Osmorhiza berteroi (= Osmorhiza chilensis), Orthilia secunda (= Pyrola secunda), Thalictrum occidentale, Trillium ovatum, Viola glabella (or Viola canadensis), and Viola orbiculata.
Dynamics: This association is sufficiently mesic to support a host of tree species more shade-tolerant than Pinus contorta and, therefore, the association is purely a seral community type. Pinus contorta is a relatively short-lived species in these mesic forests; seldom does it exceed 200 years in the northern Rocky Mountains and considerably less in the Inland Northwest, where many Pinus contorta stands are found naturally breaking-up at around 120 years since initiation. It has been noted in northern Idaho that following disturbance in this type Pinus contorta often does not succeed itself, the first tree-dominated successional stages being dominated by Larix occidentalis, Pseudotsuga menziesii, or less frequently by more shade-tolerant species; some of this variation in fire succession can be related to the varying degrees of cone serotiny in Pinus contorta. Higher incidences of cone serotiny are found in regions/ecosystems experiencing a higher degree of stand-replacing fire (Lotan and Perry 1983). This plant association (because of the requirement of Pinus contorta dominance) is expected to be more prevalent in high montane and subalpine environments, where stand-replacing fire is the primary mode of stand initiation. In the more mesic montane environments stand-replacing fire is a less frequent mode of stand initiation and, being very shade-intolerant, Pinus contorta does poorly at establishing in these less-than-full-sunlight conditions; this association is expected to be an uncommon type in the Tsuga heterophylla, Thuja plicata, and Abies grandis series.
Environmental Description: Broadly distributed throughout the northern Rocky Mountains and adjacent terrain, this large-patch to matrix seral community occupies relatively moist (mesic) and warm to cool sites having free air drainage and lacking frost-pocket conditions. It occurs on slopes of all degrees of steepness and aspect orientation, though it is more likely to occur from toeslope through midslope positions (predominantly collecting positions). At the dry extreme of its distribution it is more strongly associated with protected positions such as concave slopes, moist depressions in gently sloping plateau areas, stringers along perennial stream bottoms, toeslopes and northeastern aspects. In the north it ranges from 760 to 1585 m (450-5200 feet) (extreme outliers at 1710 m (5600 feet)), whereas to the south it ranges from 1060 to 1710 m (3500-5600 feet). A wide variety of parent materials are represented, including all major rock types (sedimentary, metamorphic and igneous) with examples as disparate as granite and limestone; all manner of glacio-fluvial material blankets stream and river terraces, and glacial till is common on the upland benches. In eastern Washington, northern Idaho and northwestern Montana it is routinely found on ash caps, ranging from 3 to 60 cm in depth. Soil textures are predominantly from the fine end of the spectrum with loams and silt loams common (reflecting in part a volcanic ash component, if not ash cap); soils typically have less than 15% coarse-fragment content and are well-drained.
Geographic Range: This association occurs from the southern portion of the Idaho Batholith of central Idaho northward to the eastern fringes of the Colville National Forest of northeastern Washington, across northern Idaho and southeastern British Columbia and eastward into western Montana, predominantly west of the Continental Divide to its northeast extremes in southwestern Alberta.
Nations: CA,US
States/Provinces: AB, ID, MT, OR, WA
Plot Analysis Summary:
http://vegbank.org/natureserve/ELEMENT_GLOBAL.2.731473
Confidence Level: Low
Confidence Level Comments: No Data Available
Grank: G5
Greasons: No Data Available
Type | Name | Database Code | Classification Code |
---|---|---|---|
Class | 1 Forest & Woodland Class | C01 | 1 |
Subclass | 1.B Temperate & Boreal Forest & Woodland Subclass | S15 | 1.B |
Formation | 1.B.2 Cool Temperate Forest & Woodland Formation | F008 | 1.B.2 |
Division | 1.B.2.Nb Rocky Mountain Forest & Woodland Division | D194 | 1.B.2.Nb |
Macrogroup | 1.B.2.Nb.5 Subalpine Fir - Engelmann Spruce - Whitebark Pine Rocky Mountain Forest Macrogroup | M020 | 1.B.2.Nb.5 |
Group | 1.B.2.Nb.5.a Lodgepole Pine Rocky Mountain Forest & Woodland Group | G220 | 1.B.2.Nb.5.a |
Alliance | A3366 Lodgepole Pine Rocky Mountain Forest Alliance | A3366 | 1.B.2.Nb.5.a |
Association | CEGL005916 Lodgepole Pine / Bride''s Bonnet Forest | CEGL005916 | 1.B.2.Nb.5.a |
Concept Lineage: No Data Available
Predecessors: No Data Available
Obsolete Names: No Data Available
Obsolete Parents: No Data Available
Synonomy: >< Abies grandis / Acer circinatum / Clintonia uniflora (Lillybridge et al. 1995) [ABGR/ACCI/CLUN COMMUNITY TYPE CWS553]
>< Abies grandis / Acer glabrum / Clintonia uniflora Plant Association (Williams et al. 1995)
>< Abies grandis / Clintonia uniflora Habitat Type, Clintonia uniflora Phase (Pfister et al. 1977)
>< Abies grandis / Clintonia uniflora Habitat Type, Clintonia uniflora Phase (Cooper et al. 1987)
>< Abies grandis / Clintonia uniflora Plant Association (Johnson and Simon 1987)
>< Abies lasiocarpa / Clintonia uniflora Habitat Type, Aralia nudicaulis Phase (Steele et al. 1981)
>< Abies lasiocarpa / Clintonia uniflora Habitat Type, Aralia nudicaulis Phase (Pfister et al. 1977)
>< Abies lasiocarpa / Clintonia uniflora Habitat Type, Aralia nudicaulis Phase (Cooper et al. 1987)
>< Abies lasiocarpa / Clintonia uniflora Habitat Type, Clintonia uniflora Phase (Steele et al. 1981)
>< Abies lasiocarpa / Clintonia uniflora Habitat Type, Clintonia uniflora Phase (Cooper et al. 1987)
>< Abies lasiocarpa / Clintonia uniflora Habitat Type, Clintonia uniflora Phase (Pfister et al. 1977)
>< Abies lasiocarpa / Clintonia uniflora Plant Association (Johnson and Simon 1987)
>< Abies lasiocarpa / Clintonia uniflora Plant Association (Williams et al. 1995)
>< Abies lasiocarpa / Pachistima myrsinites Habitat Type (Daubenmire and Daubenmire 1968)
= Pinus contorta / Clintonia uniflora Forest (Hop et al. 2007)
>< Thuja plicata / Clintonia uniflora Habitat Type, Clintonia uniflora Phase (Pfister et al. 1977)
>< Thuja plicata / Clintonia uniflora Habitat Type, Clintonia uniflora Phase (Cooper et al. 1987)
>< Tsuga heterophylla / Acer circinatum / Clintonia uniflora (Lillybridge et al. 1995) [TSHE/ACCI/CLUN ASSOCIATION CHS227]
>< Tsuga heterophylla / Clintonia uniflora Habitat Type, Clintonia uniflora Phase (Cooper et al. 1987)
>< Tsuga heterophylla / Clintonia uniflora Habitat Type, Clintonia uniflora Phase (Pfister et al. 1977)
>< Tsuga heterophylla / Pachistima myrsinites / Clintonia uniflora (Lillybridge et al. 1995) [TSHE/PAMY/CLUN ASSOCIATION CHS143]
>< Abies grandis / Acer glabrum / Clintonia uniflora Plant Association (Williams et al. 1995)
>< Abies grandis / Clintonia uniflora Habitat Type, Clintonia uniflora Phase (Pfister et al. 1977)
>< Abies grandis / Clintonia uniflora Habitat Type, Clintonia uniflora Phase (Cooper et al. 1987)
>< Abies grandis / Clintonia uniflora Plant Association (Johnson and Simon 1987)
>< Abies lasiocarpa / Clintonia uniflora Habitat Type, Aralia nudicaulis Phase (Steele et al. 1981)
>< Abies lasiocarpa / Clintonia uniflora Habitat Type, Aralia nudicaulis Phase (Pfister et al. 1977)
>< Abies lasiocarpa / Clintonia uniflora Habitat Type, Aralia nudicaulis Phase (Cooper et al. 1987)
>< Abies lasiocarpa / Clintonia uniflora Habitat Type, Clintonia uniflora Phase (Steele et al. 1981)
>< Abies lasiocarpa / Clintonia uniflora Habitat Type, Clintonia uniflora Phase (Cooper et al. 1987)
>< Abies lasiocarpa / Clintonia uniflora Habitat Type, Clintonia uniflora Phase (Pfister et al. 1977)
>< Abies lasiocarpa / Clintonia uniflora Plant Association (Johnson and Simon 1987)
>< Abies lasiocarpa / Clintonia uniflora Plant Association (Williams et al. 1995)
>< Abies lasiocarpa / Pachistima myrsinites Habitat Type (Daubenmire and Daubenmire 1968)
= Pinus contorta / Clintonia uniflora Forest (Hop et al. 2007)
>< Thuja plicata / Clintonia uniflora Habitat Type, Clintonia uniflora Phase (Pfister et al. 1977)
>< Thuja plicata / Clintonia uniflora Habitat Type, Clintonia uniflora Phase (Cooper et al. 1987)
>< Tsuga heterophylla / Acer circinatum / Clintonia uniflora (Lillybridge et al. 1995) [TSHE/ACCI/CLUN ASSOCIATION CHS227]
>< Tsuga heterophylla / Clintonia uniflora Habitat Type, Clintonia uniflora Phase (Cooper et al. 1987)
>< Tsuga heterophylla / Clintonia uniflora Habitat Type, Clintonia uniflora Phase (Pfister et al. 1977)
>< Tsuga heterophylla / Pachistima myrsinites / Clintonia uniflora (Lillybridge et al. 1995) [TSHE/PAMY/CLUN ASSOCIATION CHS143]
- ANHIC [Alberta Natural Heritage Information Centre]. 2018. Community database files. Alberta Natural Heritage Information Centre, Parks and Protected Areas Division, Alberta Community Development, Edmonton.
- Achuff, P. L., R. L. McNeil, M. L. Coleman, C. Wallis and C. Wershler. 2002. Ecological land classification of Waterton Lakes National Park, Alberta. Volume I: Integrated resource description. Parks Canada, Waterton Lakes National Park, Alberta. 226 pp.
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- Cooper, Steve. Personal communication. Ecologist, Montana Natural Heritage Program, Helena, MT.
- Daubenmire, R. F., and J. B. Daubenmire. 1968. Forest vegetation of eastern Washington and northern Idaho. Washington State University Agricultural Experiment Station Technical Bulletin No. 60. 104 pp.
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- Johnson, C. G., Jr., and S. A. Simon. 1987. Plant associations of the Wallowa-Snake Province Wallowa-Whitman National Forest. Technical Paper R6-ECOL-TP-255A-86. USDA Forest Service, Pacific Northwest Region, Wallowa-Whitman National Forest. 399 pp. plus appendices.
- Lillybridge, T. R., B. L. Kovalchik, C. K. Williams, and B. G. Smith. 1995. Field guide for forested plant associations of the Wenatchee National Forest. General Technical Report PNW-GTR-359. USDA Forest Service, Pacific Northwest Research Station, Portland, OR. 335 pp.
- Lotan, J. E., and D. A. Perry. 1983. Ecology and regeneration of lodgepole pine. Agriculture Handbook 606. USDA Forest Service, Washington, DC. 51 pp.
- Pfister, R. D., B. L. Kovalchik, S. F. Arno, and R. C. Presby. 1977. Forest habitat types of Montana. General Technical Report INT-34. USDA Forest Service, Intermountain Forest and Range Experiment Station, Ogden, UT. 174 pp.
- Reid, M. S., S. V. Cooper, and G. Kittel. 2004. Vegetation classification of Waterton-Glacier International Peace Park. Final report for USGS-NPS Vegetation Mapping Program, International Peace Park Mapping Project. NatureServe, Arlington VA.
- Steele, R., R. D. Pfister, R. A. Ryker, and J. A. Kittams. 1981. Forest habitat types of central Idaho. General Technical Report INT-114. USDA Forest Service, Intermountain Forest and Range Experiment Station, Ogden, UT. 138 pp.
- Western Ecology Working Group of NatureServe. No date. International Ecological Classification Standard: International Vegetation Classification. Terrestrial Vegetation. NatureServe, Boulder, CO.
- Williams, C. K., B. F. Kelly, B. G. Smith, and T. R. Lillybridge. 1995. Forest plant associations of the Colville National Forest. General Technical Report PNW-GTR-360. USDA Forest Service, Pacific Northwest Region, Portland, OR. 140 pp.
- Williams, C. K., and T. R. Lillybridge. 1983. Forested plant associations of the Okanogan National Forest. R6-Ecol-132b-1983. USDA Forest Service, Pacific Northwest Region, Portland, OR. 140 pp.