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CEGL005922 Pinus contorta / Menziesia ferruginea / Clintonia uniflora Forest
Type Concept Sentence: No Data Available
Common (Translated Scientific) Name: Lodgepole Pine / Rusty Menziesia / Bride''s Bonnet Forest
Colloquial Name: No Data Available
Hierarchy Level: Association
Type Concept: This seral, large-patch to matrix lodgepole pine forest occupies the relatively cold and dry environments across a number of climax tree series and associated geographic regions. Thus, this cold mesic type is found throughout the northern Rocky Mountains and may extend as far west as the Cascade Crest on environments characterized as montane to lower and even mid-subalpine. The association''s possible elevation range is from 915 to 1800 m (3000-5700 feet), and it consistently occurs on cool northwest- through east-facing slopes with moderate to extreme degrees of slope. It is generally associated with collecting positions from midslope to toeslope and foot-slopes when it occurs in subalpine zones, but in the more mesic montane zones, it can be found on all positions. The range of parent materials is literally as great as possible types occurring in the northern Rocky Mountains and northernmost middle Rocky Mountains and may include some ultramafics east of the Cascade Crest. The soils are acidic to very acidic, uniformly moderately well-drained to well-drained and have a highly variable coarse-fragment content. Ground surfaces have virtually no exposed rock or bare soil, and duff accumulations vary from moderate to deep. The overstory canopy is often on the cusp between open and closed (defined as 60%) and decidedly dominated by Pinus contorta, but its cover is often less than 40%. A number of other conifers may be present; on warmer sites these include Thuja plicata, Tsuga heterophylla, and Abies grandis, and on colder or higher elevation sites are found Abies lasiocarpa, Tsuga mertensiana, and Picea engelmannii. However, the most frequent canopy codominants or associates are the seral species Larix occidentalis, Pseudotsuga menziesii, and in a restricted portion of the type''s range Pinus monticola. Menziesia ferruginea conspicuously dominates the tall-shrub layer. Alnus viridis ssp. sinuata and Taxus brevifolia are the only other tall shrubs consistently present. The short-shrub layer exhibits greater diversity with Vaccinium membranaceum, Paxistima myrsinites, Rosa gymnocarpa, Rubus parviflorus, and Spiraea betulifolia being consistently present. Linnaea borealis, Chimaphila umbellata, and Vaccinium scoparium have high constancy in the dwarf-shrub layer. Bromus vulgaris (or Bromus ciliatus) are the only graminoids of note. The diagnostic forbs Clintonia uniflora and Tiarella trifoliata have high constancy (both approaching 100%) and/or cover; however, a number of other forbs also exhibit high constancy across this type''s range, including Arnica latifolia, (Arnica cordifolia at lower elevations), Coptis occidentalis, Cornus canadensis, Galium triflorum, Goodyera oblongifolia, Maianthemum stellatum, Osmorhiza berteroi, Orthilia secunda, Thalictrum occidentale, Trillium ovatum, Viola orbiculata, and Xerophyllum tenax. As with the forb layer, the bryoid layer cover apparently varies inversely with the degree of canopy closure.
Diagnostic Characteristics: No Data Available
Rationale for Nominal Species or Physiognomic Features: No Data Available
Classification Comments: With the comparatively recent emphasis on developing descriptions of (and keys to) existing vegetation (Grossman et al. 1998), it has yet to be determined at what cover value forest vegetation types will be distinguished one from another when the canopy tree species are mostly, or exclusively, seral in nature and have a broad environmental range (broad niche). Pfister et al. (1977) recognized Pinus contorta community types only when no other, more shade-tolerant tree species could be found on site. With Pinus contorta, which is exclusively seral except with respect to some subspecies on unusual substrates or atypical environments, researchers in Glacier National Park took the position that this very shade-intolerant, stand-replacing fire-adapted species should have several times the cover of the next most abundant canopy species for a Pinus contorta type to be recognized. What is really being indicated by this approach are areas having experienced stand-replacing fires (or similar catastrophic disturbance, e.g., clearcutting). Another approach could simply recognize the plurality of cover in assigning stands to particular associations treating existing vegetation. The stands representing this type are climax in a number of different tree series, including Abies lasiocarpa, Abies grandis, Tsuga heterophylla, Tsuga mertensiana, and Thuja plicata. That this association in fact occurs in the states and USFS Sections listed derives from this writer''s interpretation of constancy/cover tables of various authors/publications; where Pinus contorta is listed as a major seral species, the inference has been made that stands quite likely occur on the landscape where this species strongly dominates the upper canopy (often observed throughout northern Idaho and western Montana) (S. Cooper pers. comm.). It should also be noted that this type is probably less common in zones where Thuja plicata, Tsuga heterophylla, and Abies amabilis are the climax dominants. This is because, following disturbance in these zones, the climax trees are quick to reclaim the site, i.e., they comprise a significant cover on the earliest forested successional stages (Pinus contorta may seldom be a major seral component on these sites and thus the association is rare as well), and Pinus contorta is favored by stand-replacing fire which is uncommon in these mesic forests.
Similar NVC Types: No Data Available
note: No Data Available
Physiognomy and Structure: No Data Available
Floristics: The overstory canopy is often, especially in subalpine environments, on the cusp between open and closed (currently defined as 60%) and decidedly dominated by Pinus contorta, but its cover is often less than 40%. A whole host of tree species is capable of playing a subordinate role early in the sere; on warmer sites these include Thuja plicata, Tsuga heterophylla, and Abies grandis, and on colder or higher elevation sites are found Abies lasiocarpa, Tsuga mertensiana, and Picea engelmannii. However, the most frequent canopy codominants or associates are the seral species Larix occidentalis, Pseudotsuga menziesii, and in a restricted portion of the type''s range Pinus monticola. Menziesia ferruginea conspicuously dominates the tall-shrub layer, however, its height is quite dependent on environment, exceeding 3.5 m (10 feet) on montane slopes and actually classed as a short shrub (<2 m [6.5 feet]) within much of its subalpine range (where sites are potentially dominated by Abies lasiocarpa, Picea engelmannii, and Tsuga mertensiana); Alnus viridis ssp. sinuata and Taxus brevifolia (predominantly in Idaho and western Montana) are the only other tall shrubs consistently present. The short-shrub layer exhibits greater diversity than the other shrub components with Vaccinium membranaceum, Paxistima myrsinites, Rosa gymnocarpa, Rubus parviflorus, and Spiraea betulifolia being consistently present. In the montane environments Linnaea borealis and Chimaphila umbellata have high constancy in the dwarf-shrub layer, whereas in the subalpine they are considerably reduced, and Vaccinium scoparium is consistently present with up to 50% cover. Bromus vulgaris (or Bromus ciliatus) are the only graminoids of note. The diagnostic forbs Clintonia uniflora and Tiarella trifoliata have high constancy (both approaching 100%) and/or cover; however, a number of other forbs also exhibit high constancy across this type''s range, including Arnica latifolia, (Arnica cordifolia at lower elevations), Coptis occidentalis (peculiar to central and northern Idaho), Cornus canadensis, Galium triflorum, Goodyera oblongifolia, Maianthemum stellatum, Osmorhiza berteroi (= Osmorhiza chilensis), Orthilia secunda (= Pyrola secunda), Thalictrum occidentale, Trillium ovatum, Viola orbiculata, and Xerophyllum tenax. As with the forb layer, bryoid cover apparently varies inversely with the degree of canopy closure; however, tree canopy closure here is never what it is on sites lacking such a prolific tall-shrub layer.
Dynamics: This association is expected to be much more prevalent in high montane and subalpine environments, where stand-replacing fire is the primary mode of stand initiation. In the more mesic montane environments stand-replacing fire is a less frequent mode of stand initiation and, being very shade-intolerant, Pinus contorta does poorly at establishing in these less than full sunlight conditions; this association is expected to be an uncommon type in the Tsuga heterophylla, Thuja plicata, and Abies grandis series. Data from the subalpine zone indicate that because of natural mortality patterns Pinus contorta canopy domination is not expected to last more than 160-180 years following stand initiation (Cooper et al. 1987) and most commonly break-up occurs at 100 to 120 years.
Environmental Description: This seral, large-patch to matrix type occupies relatively cold and moist environments across a number of climax tree series and associated geographic regions; the species defining these series include, but are not limited to, Thuja plicata, Tsuga heterophylla, Tsuga mertensiana, Abies grandis, Abies lasiocarpa, and Picea engelmannii. Thus, this cold mesic type is found throughout the northern Rocky Mountains and may extend as far west as the Cascade Crest on environments characterized as montane to lower and even mid-subalpine. The association''s possible elevation range is from 915 to 1800 m (4000-5700 feet), and regardless of the climax series in which it is found, it consistently occurs on cool northwest- through east-facing slopes with moderate to extreme degrees of slope. It is generally associated with collecting positions from midslope to toeslope and foot-slopes when it occurs in subalpine zones, but in the more mesic montane zones, it can be found on all positions. It has been recorded as low as 910 m (3000 feet) on benches and swales where cold air ponds. The range of parent materials is, with the exception of highly unusual substrates like serpentine, literally as great as possible types occurring in the northern Rocky Mountains and northernmost middle Rocky Mountains and may include some ultramafics east of the Cascade Crest. It is difficult to simply characterize the soils as well, but they are uniformly moderately well-drained to well-drained and have a highly variable coarse-fragment content, but are mostly moderately gravelly throughout (at least when the type occurs in the subalpine zone). Soil reactions vary from acidic to very acidic. Ground surfaces have virtually no exposed rock or bare soil, and duff accumulations vary from moderate to deep.
Geographic Range: This association occurs from the southern portion of the Idaho Batholith of central Idaho northward to the eastern fringes of the Colville National Forest of northeastern Washington and across northern Idaho and into western Montana, predominantly west of the Continental Divide, and as far eastward as southwestern Alberta. Given the opportunity for more complete crosswalking, this type might well be documented from British Columbia and the east slope of the Cascades.
Nations: CA,US
States/Provinces: AB, ID, MT, OR?, WA?
Plot Analysis Summary:
http://vegbank.org/natureserve/ELEMENT_GLOBAL.2.731424
Confidence Level: Low
Confidence Level Comments: No Data Available
Grank: G4G5
Greasons: No Data Available
Type | Name | Database Code | Classification Code |
---|---|---|---|
Class | 1 Forest & Woodland Class | C01 | 1 |
Subclass | 1.B Temperate & Boreal Forest & Woodland Subclass | S15 | 1.B |
Formation | 1.B.2 Cool Temperate Forest & Woodland Formation | F008 | 1.B.2 |
Division | 1.B.2.Nb Rocky Mountain Forest & Woodland Division | D194 | 1.B.2.Nb |
Macrogroup | 1.B.2.Nb.5 Subalpine Fir - Engelmann Spruce - Whitebark Pine Rocky Mountain Forest Macrogroup | M020 | 1.B.2.Nb.5 |
Group | 1.B.2.Nb.5.a Lodgepole Pine Rocky Mountain Forest & Woodland Group | G220 | 1.B.2.Nb.5.a |
Alliance | A3366 Lodgepole Pine Rocky Mountain Forest Alliance | A3366 | 1.B.2.Nb.5.a |
Association | CEGL005922 Lodgepole Pine / Rusty Menziesia / Bride''s Bonnet Forest | CEGL005922 | 1.B.2.Nb.5.a |
Concept Lineage: No Data Available
Predecessors: No Data Available
Obsolete Names: No Data Available
Obsolete Parents: No Data Available
Synonomy: >< Abies grandis / Clintonia uniflora Habitat Type, Menziesia ferruginea Phase (Cooper et al. 1987)
>< Abies lasiocarpa / Clintonia uniflora Habitat Type, Menziesia ferruginea Phase (Cooper et al. 1987)
>< Abies lasiocarpa / Clintonia uniflora Habitat Type, Menziesia ferruginea Phase (Pfister et al. 1977)
>< Abies lasiocarpa / Rhododendron albiflorum / Xerophyllum tenax Plant Association (Williams et al. 1995)
>< Abies lasiocarpa / Rhododendron albiflorum Plant Association (Williams et al. 1995)
>< Picea engelmannii - Abies lasiocarpa / Menziesia ferruginea - Tiarella trifoliata Habitat Type (Ogilvie 1962)
= Pinus contorta / Menziesia ferruginea / Clintonia uniflora Forest (Achuff et al. 2002)
= Pinus contorta / Menziesia ferruginea / Clintonia uniflora Forest (Hop et al. 2007)
>< Thuja plicata / Clintonia uniflora Habitat Type, Menziesia ferruginea Phase (Cooper et al. 1987)
>< Tsuga heterophylla / Clintonia uniflora Habitat Type, Menziesia ferruginea Phase (Cooper et al. 1987)
>< Tsuga mertensiana / Clintonia uniflora Habitat Type, Menziesia ferruginea Phase (Cooper et al. 1987)
>< Abies lasiocarpa / Clintonia uniflora Habitat Type, Menziesia ferruginea Phase (Cooper et al. 1987)
>< Abies lasiocarpa / Clintonia uniflora Habitat Type, Menziesia ferruginea Phase (Pfister et al. 1977)
>< Abies lasiocarpa / Rhododendron albiflorum / Xerophyllum tenax Plant Association (Williams et al. 1995)
>< Abies lasiocarpa / Rhododendron albiflorum Plant Association (Williams et al. 1995)
>< Picea engelmannii - Abies lasiocarpa / Menziesia ferruginea - Tiarella trifoliata Habitat Type (Ogilvie 1962)
= Pinus contorta / Menziesia ferruginea / Clintonia uniflora Forest (Achuff et al. 2002)
= Pinus contorta / Menziesia ferruginea / Clintonia uniflora Forest (Hop et al. 2007)
>< Thuja plicata / Clintonia uniflora Habitat Type, Menziesia ferruginea Phase (Cooper et al. 1987)
>< Tsuga heterophylla / Clintonia uniflora Habitat Type, Menziesia ferruginea Phase (Cooper et al. 1987)
>< Tsuga mertensiana / Clintonia uniflora Habitat Type, Menziesia ferruginea Phase (Cooper et al. 1987)
- ANHIC [Alberta Natural Heritage Information Centre]. 2018. Community database files. Alberta Natural Heritage Information Centre, Parks and Protected Areas Division, Alberta Community Development, Edmonton.
- Achuff, P. L., R. L. McNeil, M. L. Coleman, C. Wallis and C. Wershler. 2002. Ecological land classification of Waterton Lakes National Park, Alberta. Volume I: Integrated resource description. Parks Canada, Waterton Lakes National Park, Alberta. 226 pp.
- Cooper, S. V., K. E. Neiman, R. Steele, and D. W. Roberts. 1987. Forest habitat types of northern Idaho: A second approximation. General Technical Report INT-236.USDA Forest Service, Intermountain Research Station, Ogden, UT. 135 pp. [reprinted in 1991]
- Cooper, Steve. Personal communication. Ecologist, Montana Natural Heritage Program, Helena, MT.
- Daubenmire, R. F., and J. B. Daubenmire. 1968. Forest vegetation of eastern Washington and northern Idaho. Washington State University Agricultural Experiment Station Technical Bulletin No. 60. 104 pp.
- Grossman, D. H., D. Faber-Langendoen, A. S. Weakley, M. Anderson, P. Bourgeron, R. Crawford, K. Goodin, S. Landaal, K. Metzler, K. D. Patterson, M. Pyne, M. Reid, and L. Sneddon. 1998. International classification of ecological communities: Terrestrial vegetation of the United States. Volume I. The national vegetation classification system: Development, status, and applications. The Nature Conservancy, Arlington, VA.
- Hop, K., M. Reid, J. Dieck, S. Lubinski, and S. Cooper. 2007. U.S. Geological Survey-National Park Service Vegetation Mapping Program: Waterton-Glacier International Peace Park. U.S. Geological Survey, Upper Midwest Environmental Sciences Center, La Crosse, WI. 131 pp. plus Appendices A-L.
- Johnson, C. G., Jr., and S. A. Simon. 1987. Plant associations of the Wallowa-Snake Province Wallowa-Whitman National Forest. Technical Paper R6-ECOL-TP-255A-86. USDA Forest Service, Pacific Northwest Region, Wallowa-Whitman National Forest. 399 pp. plus appendices.
- Lillybridge, T. R., B. L. Kovalchik, C. K. Williams, and B. G. Smith. 1995. Field guide for forested plant associations of the Wenatchee National Forest. General Technical Report PNW-GTR-359. USDA Forest Service, Pacific Northwest Research Station, Portland, OR. 335 pp.
- Ogilvie, R. T. 1962. Ecology of spruce forests on the east slope of the Rocky Mountains in Alberta. Unpublished dissertation, Washington State University, Pullman. 189 pp.
- Pfister, R. D., B. L. Kovalchik, S. F. Arno, and R. C. Presby. 1977. Forest habitat types of Montana. General Technical Report INT-34. USDA Forest Service, Intermountain Forest and Range Experiment Station, Ogden, UT. 174 pp.
- Reid, M. S., S. V. Cooper, and G. Kittel. 2004. Vegetation classification of Waterton-Glacier International Peace Park. Final report for USGS-NPS Vegetation Mapping Program, International Peace Park Mapping Project. NatureServe, Arlington VA.
- Western Ecology Working Group of NatureServe. No date. International Ecological Classification Standard: International Vegetation Classification. Terrestrial Vegetation. NatureServe, Boulder, CO.
- Williams, C. K., B. F. Kelly, B. G. Smith, and T. R. Lillybridge. 1995. Forest plant associations of the Colville National Forest. General Technical Report PNW-GTR-360. USDA Forest Service, Pacific Northwest Region, Portland, OR. 140 pp.