Print Report

CEGL005854 Pseudotsuga menziesii / Clintonia uniflora - Xerophyllum tenax Forest

Type Concept Sentence: No Data Available


Common (Translated Scientific) Name: Douglas-fir / Bride''s Bonnet - Common Beargrass Forest

Colloquial Name: No Data Available

Hierarchy Level:  Association

Type Concept: This is a seral, mesic, large-patch to matrix type found in the foothills and montane to lower and even mid-subalpine, relatively cold and dry environments throughout the northern Rocky Mountains and may extend as far west as the Cascade Crest. The associations'' possible elevation range is from 915 to 1800 m (3000-5900 feet), and regardless of the climax series in which it is found, it consistently occurs on south- through west-facing exposures. The range of parent materials is, with the exception of highly unusual substrates like serpentine, literally as great as possible types occurring in the northern Rocky Mountains and northernmost middle Rocky Mountains and may include some ultramafics east of the Cascade Crest. It is difficult to characterize the soils as well, but they are uniformly well-drained and have a low coarse-fragment content, except those sites within the lower to mid-subalpine zone. The overstory is dominated by Pseudotsuga menziesii with a whole host of tree species capable of playing a subordinate role; on warmer sites these include Thuja plicata, Tsuga heterophylla, Abies grandis, and on colder or higher elevation sites are found Abies lasiocarpa, Tsuga mertensiana, and Picea engelmannii. However, the most frequent canopy codominants or associates are the seral species Larix occidentalis, Pinus contorta, and in a restricted portion of the type''s range, Pinus monticola. The tall-shrub component is relatively unimportant, only Alnus viridis ssp. sinuata and Amelanchier alnifolia approach 50% constancy (and have low cover values). The short-shrub layer exhibits greater cover and diversity than the other shrub components with Vaccinium membranaceum, Paxistima myrsinites, Rosa gymnocarpa, Rubus parviflorus, and Spiraea betulifolia being consistently present. Linnaea borealis and Chimaphila umbellata have high constancy in the dwarf-shrub layer. Bromus vulgaris (or Bromus ciliatus) are the only graminoids of note. The diagnostic forbs Clintonia uniflora, Xerophyllum tenax, and Tiarella trifoliata naturally have high constancy and/or cover, however, a number of other forbs also exhibit high constancy, including Arnica latifolia, Aralia nudicaulis, Adenocaulon bicolor, Coptis occidentalis, Cornus canadensis, Galium triflorum, Goodyera oblongifolia, Maianthemum stellatum, Osmorhiza berteroi, Pedicularis racemosa, Orthilia secunda, Thalictrum occidentale, Trillium ovatum, and Viola orbiculata.

Diagnostic Characteristics: No Data Available

Rationale for Nominal Species or Physiognomic Features: No Data Available

Classification Comments: This association defines an extremely broad environmental range due to the approach of defining as unique associations those communities that would formerly have been subsumed within a number of different climax or potential vegetation types; in the case of this association it is a successional stage in environments as mild and wet (defining a Pacific maritime climatic influence) as those supporting Thuja plicata or Tsuga heterophylla / Clintonia uniflora associations (and permutations) to those of the lower subalpine zone characterized by Abies lasiocarpa, Abies grandis, Tsuga mertensiana, and even Picea engelmannii / Clintonia uniflora forest associations. That this association in fact occurs in the states and USFS sections listed derives from this writer''s interpretation of constancy/cover tables of various authors/publications; where Pseudotsuga is listed as a major seral species, the inference has been made that stands quite likely occur on the landscape where this species strongly dominates the upper canopy (often observed by this author throughout northern Idaho and western Montana). It should also be noted that this type is probably less common in zones where Thuja plicata and Tsuga heterophylla are the climax dominants because, following disturbance in these zones, the climax trees are quick to reclaim the site, i.e., they comprise a significant cover of the earliest forested successional stages. The crosswalking for this type is incomplete, and it may well occur west of the Cascade Crest in Oregon and Washington.

Similar NVC Types: No Data Available
note: No Data Available

Physiognomy and Structure: No Data Available

Floristics: The overstory is dominated by Pseudotsuga menziesii with a whole host of tree species capable of playing a subordinate role; on warmer sites these include Thuja plicata, Tsuga heterophylla, Abies grandis, and on colder or higher elevation sites are found Abies lasiocarpa, Tsuga mertensiana, and Picea engelmannii. However, the most frequent canopy codominants or associates are the seral species Larix occidentalis, Pinus contorta, and in a restricted portion of the type''s range, Pinus monticola. The tall-shrub component is relatively unimportant, only Alnus viridis ssp. sinuata and Amelanchier alnifolia approach 50% constancy (and have low cover values). The short-shrub layer exhibits greater cover and diversity than the other shrub components with Vaccinium membranaceum, Paxistima myrsinites, Rosa gymnocarpa, Rubus parviflorus, and Spiraea betulifolia being consistently present. Linnaea borealis and Chimaphila umbellata have high constancy in the dwarf-shrub layer. Bromus vulgaris (or Bromus ciliatus) are the only graminoids of note. The diagnostic forbs Clintonia uniflora, Xerophyllum tenax, and Tiarella trifoliata naturally have high constancy and/or cover, however, a number of other forbs also exhibit high constancy including Arnica latifolia, Aralia nudicaulis, Adenocaulon bicolor, Coptis occidentalis, Cornus canadensis, Galium triflorum, Goodyera oblongifolia, Maianthemum stellatum, Osmorhiza berteroi (= Osmorhiza chilensis), Pedicularis racemosa, Orthilia secunda (= Pyrola secunda), Thalictrum occidentale, Trillium ovatum, and Viola orbiculata.

Dynamics:  No Data Available

Environmental Description:  This seral, large-patch to matrix type occupies the relatively cold and dry environments across a number of climax tree series and associated geographic regions; the species defining these series include, but are not limited to, Thuja plicata, Tsuga heterophylla, Tsuga mertensiana, Abies grandis, Abies lasiocarpa, and Picea engelmannii. Thus this mesic type is found throughout the northern Rocky Mountains and may extend as far west as the Cascade Crest on environments characterized as foothills and montane to lower and even mid subalpine. The associations possible elevation range is from 915 to 1800 m (3000-5900 feet), and regardless of the climax series in which it is found, it consistently occurs on south- through west-facing exposures. These are generally more shedding than collecting positions, occurring in any ridge or hillslope system from midslope up to ridge crest, including level terrain of ridge summits. The range of parent materials is, with the exception of highly unusual substrates like serpentine, literally as great as possible types occurring in the northern Rocky Mountains and northernmost middle Rocky Mountains and may include some ultramafics east of the Cascade Crest. It is difficult to characterize the soils as well, but they are uniformly well-drained and have a low coarse-fragment content, except those sites within the lower to mid-subalpine zone.

Geographic Range: This association occurs from the Blue and Wallowa mountains of northeastern Oregon and southern portion of the Idaho Batholith of central Idaho northward to the Colville National Forest of northeastern Washington, across northern Idaho and into western Montana, predominantly west of the Continental Divide, and as far east as southwestern Alberta. Given opportunity for more complete crosswalking, this type could well be documented from British Columbia and the east slope of the Cascades (the fact that a different subspecies of Pseudotsuga menziesii is distributed west of the Cascades argues for considering those communities as different.

Nations: CA,US

States/Provinces:  AB?, ID, MT, OR, WA




Confidence Level: Low

Confidence Level Comments: No Data Available

Grank: G4G5

Greasons: No Data Available


Concept Lineage: This association (CEGL000307) was based on a Habitat Type concept, including the phases of the H.T. in this association. Phases of the H.T. have now been split out into association elements, as well as the seral components of the H.T.

Predecessors: No Data Available

Obsolete Names: No Data Available

Obsolete Parents: No Data Available

Synonomy: >< Abies grandis / Clintonia uniflora Habitat Type, Xerophyllum tenax Phase (Cooper et al. 1987)
>< Abies grandis / Vaccinium membranaceum / Clintonia uniflora Plant Association (Williams et al. 1995)
>< Abies lasiocarpa / Clintonia uniflora Habitat Type, Xerophyllum tenax Phase (Pfister et al. 1977)
>< Abies lasiocarpa / Clintonia uniflora Habitat Type, Xerophyllum tenax Phase (Cooper et al. 1987)
= Pseudotsuga menziesii / Clintonia uniflora - Xerophyllum tenax Forest (Hop et al. 2007)
>< Thuja plicata / Clintonia uniflora Habitat Type, Xerophyllum tenax Phase (Cooper et al. 1987)
>< Tsuga heterophylla / Clintonia uniflora Habitat Type, Xerophyllum tenax Phase (Cooper et al. 1987)
>< Tsuga mertensiana / Clintonia uniflora Habitat Type, Xerophyllum tenax Phase (Cooper et al. 1987)
>< Tsuga mertensiana / Vaccinium membranaceum (Lillybridge et al. 1995) [TSME/VAME WEN ASSOCIATION CMS259]

Concept Author(s): Hop et al. (2007)

Author of Description: Western Ecology Group

Acknowledgements: No Data Available

Version Date: 02-19-04

  • ANHIC [Alberta Natural Heritage Information Centre]. 2018. Community database files. Alberta Natural Heritage Information Centre, Parks and Protected Areas Division, Alberta Community Development, Edmonton.
  • Achuff, P. L., R. L. McNeil, M. L. Coleman, C. Wallis and C. Wershler. 2002. Ecological land classification of Waterton Lakes National Park, Alberta. Volume I: Integrated resource description. Parks Canada, Waterton Lakes National Park, Alberta. 226 pp.
  • Cooper, S. V., K. E. Neiman, R. Steele, and D. W. Roberts. 1987. Forest habitat types of northern Idaho: A second approximation. General Technical Report INT-236.USDA Forest Service, Intermountain Research Station, Ogden, UT. 135 pp. [reprinted in 1991]
  • Daubenmire, R. F., and J. B. Daubenmire. 1968. Forest vegetation of eastern Washington and northern Idaho. Washington State University Agricultural Experiment Station Technical Bulletin No. 60. 104 pp.
  • Hop, K., M. Reid, J. Dieck, S. Lubinski, and S. Cooper. 2007. U.S. Geological Survey-National Park Service Vegetation Mapping Program: Waterton-Glacier International Peace Park. U.S. Geological Survey, Upper Midwest Environmental Sciences Center, La Crosse, WI. 131 pp. plus Appendices A-L.
  • Johnson, C. G., Jr., and S. A. Simon. 1987. Plant associations of the Wallowa-Snake Province Wallowa-Whitman National Forest. Technical Paper R6-ECOL-TP-255A-86. USDA Forest Service, Pacific Northwest Region, Wallowa-Whitman National Forest. 399 pp. plus appendices.
  • Lillybridge, T. R., B. L. Kovalchik, C. K. Williams, and B. G. Smith. 1995. Field guide for forested plant associations of the Wenatchee National Forest. General Technical Report PNW-GTR-359. USDA Forest Service, Pacific Northwest Research Station, Portland, OR. 335 pp.
  • Pfister, R. D., B. L. Kovalchik, S. F. Arno, and R. C. Presby. 1977. Forest habitat types of Montana. General Technical Report INT-34. USDA Forest Service, Intermountain Forest and Range Experiment Station, Ogden, UT. 174 pp.
  • Reid, M. S., S. V. Cooper, and G. Kittel. 2004. Vegetation classification of Waterton-Glacier International Peace Park. Final report for USGS-NPS Vegetation Mapping Program, International Peace Park Mapping Project. NatureServe, Arlington VA.
  • Western Ecology Working Group of NatureServe. No date. International Ecological Classification Standard: International Vegetation Classification. Terrestrial Vegetation. NatureServe, Boulder, CO.
  • Williams, C. K., B. F. Kelly, B. G. Smith, and T. R. Lillybridge. 1995. Forest plant associations of the Colville National Forest. General Technical Report PNW-GTR-360. USDA Forest Service, Pacific Northwest Region, Portland, OR. 140 pp.