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CEGL005838 Pinus albicaulis - Abies lasiocarpa / Vaccinium scoparium / Xerophyllum tenax Woodland

Type Concept Sentence: No Data Available


Common (Translated Scientific) Name: Whitebark Pine - Subalpine Fir / Grouse Whortleberry / Common Beargrass Woodland

Colloquial Name: No Data Available

Hierarchy Level:  Association

Type Concept: This minor small- to large-patch community is found primarily in upper subalpine habitats throughout the northern Rocky Mountains from central Idaho north and east into west-central Montana, and northward to the Canadian Rockies of southwestern Alberta. It ranges in elevation from 1585 to 2530 m (5200-8300 feet). The lower elevation occurrences are mostly associated with frost-pocket conditions. It usually occupies moisture-shedding sites from midslope to ridge shoulders and ridgetops; it can be found on well-drained benches in cold-air drainages. Degree of slope is highly variable, but warm aspects (from east-facing through west-facing) are predominant. Parent materials are various, from intrusive volcanics of the Idaho Batholith to extrusive volcanics (andesite and basalt) to various sedimentary and metamorphic species, including quartzite, argillite, siltstone, sandstone and schist. Soils are acidic, well-drained, and soil texture is usually on the coarser end of the spectrum (gravelly sandy loams and loams being common). Exposed rock and soil are generally less than 10%, but rock content in the profile often exceeds 20%. The overstory is usually an open canopy (less than 50% cover) dominated by a variable mix of Abies lasiocarpa, Picea engelmannii, Pinus contorta, and the indicator species Pinus albicaulis, which must have at least 5% cover for this type to be recognized. The undergrowth may be quite dense with a layer of Vaccinium scoparium surrounding clumps of Xerophyllum tenax with a thinly distributed Vaccinium membranaceum superimposed; the combined cover of these three species often exceeds 80%. Tall shrubs, if present, could be characterized as accidentals. Spiraea betulifolia and Lonicera utahensis are consistently scattered in the short-shrub layer. Carex geyeri and Calamagrostis rubescens are the only graminoids regularly present, and their cover seldom exceeds 10%. Other than the abundance of the diagnostic Xerophyllum tenax, the forb layer has low cover and is comparatively depauperate with only Arnica latifolia, Anemone piperi, Goodyera oblongifolia, Orthilia secunda, and Viola orbiculata having a consistent presence.

Diagnostic Characteristics: No Data Available

Rationale for Nominal Species or Physiognomic Features: No Data Available

Classification Comments: This association was largely subsumed within the Abies lasiocarpa / Xerophyllum tenax Habitat Type, Vaccinium scoparium Phase (Pfister et al. 1977, Steele et al. 1981, Cooper et al. 1987), in both Idaho and Montana. In accordance with the recent emphasis on creating existing vegetation plant associations, many of the plots used to substantiate the habitat type name cited above would now be classified as ~Pinus contorta / Vaccinium scoparium / Xerophyllum tenax Forest (CEGL005924)$$ or Abies lasiocarpa - Picea engelmannii / Vaccinium scoparium / Xerophyllum tenax Forest (CEGL005914)$$, and a small number have been identified as belonging to the type under consideration. Recognition of a type based on a very minimal cover of a seral tree species (Pinus albicaulis) of broad distribution should provoke questions; the intent of this broadly inclusive type was to capture stands (specifically their locations, particularly in mapping efforts) wherein this highly threatened species occurs or will have once occurred. Considering Vaccinium scoparium and Vaccinium myrtillus as ecological analogues for classification purposes could also create problems, as the ecology of Vaccinium myrtillus is not so well known.

Similar NVC Types: No Data Available
note: No Data Available

Physiognomy and Structure: No Data Available

Floristics: Exposed rock and soil are generally less than 10%, but rock content in the profile often exceeds 20%. The overstory is usually an open canopy (less than 50% cover) dominated by a variable mix of Abies lasiocarpa, Picea engelmannii, Pinus contorta, and the indicator species Pinus albicaulis, which must have at least 5% cover for this type to be recognized as distinctive from ~Abies lasiocarpa - Picea engelmannii / Vaccinium scoparium / Xerophyllum tenax Forest (CEGL005914)$$ or the Pinus contorta-dominated version of this community. The undergrowth may be quite dense with a layer of Vaccinium scoparium surrounding clumps of Xerophyllum tenax with a thinly distributed Vaccinium membranaceum superimposed; the combined cover of these three species often exceeds 80%. Tall shrubs, if present, could be characterized as accidentals. Spiraea betulifolia and Lonicera utahensis are consistently scattered in the short-shrub layer. Carex geyeri and Calamagrostis rubescens are the only graminoids regularly present, and their cover seldom exceeds 10%. Other than the abundance of the diagnostic Xerophyllum tenax, the forb layer has low cover and is comparatively depauperate with only Arnica latifolia, Anemone piperi, Goodyera oblongifolia, Orthilia secunda (= Pyrola secunda), and Viola orbiculata having a consistent presence.

Dynamics:  Prior to the advent of white pine blister rust (Cronartium ribicola) in the ecosystems containing this association, this type would have been considered a long-persisting seral community because the diagnostic species Pinus albicaulis is quite capable of attaining ages in excess of 400 years. Persistence to these extreme ages would have assured this species being present (say within a drainage) given normal fire-return intervals. However, some disturbance (most usually fire) is needed to reinitiate this association once the Pinus albicaulis has died out. Dispersal by animal vectors is its normal mode of reestablishing; corvid birds caching seeds on recently denuded slopes is the dominant mode of dispersal (unrecovered seeds germinate to form seedling clusters). Squirrels also harvest cones, which are cached in the forest; though caches are unlikely to escape harvest (by squirrels or bears), should unharvested seeds remain to germinate, they would do so well within an established stand. These seedlings would be subject to canopy shading, a light regime inimical to their establishment and survival. Given the virulence of white pine blister rust within the geographic area supporting this association, it is probable that the term ecologically extinct applied to Pinus albicaulis in Glacier National Park will become operative for much of this type''s distribution.

Environmental Description:  This minor small- to large-patch community is found primarily in upper subalpine habitats throughout the northern portion of central Idaho and northward to the Nez Perce and Clearwater national forests and west into west-central Montana (where it is areally most extensive); east of the Continental Divide it extends northward to the Canadian Rockies of southwestern Alberta. In the southern portion of its distribution elevations range from 1980 to 2530 m (6500-8300 feet), whereas in the north it ranges between 1585 and 2105 m (6500-8300 feet) and is really only regularly present above 1800 m (5900 feet), the lower elevation occurrences being associated with frost-pocket conditions. It usually occupies moisture-shedding sites from midslope to ridge shoulders and ridgetops; it can be found on well-drained benches in cold-air drainages. Degree of slope is highly variable, but warm aspects (from east-facing through west-facing) are predominant. Parent materials are various, from intrusive volcanics of the Idaho Batholith to extrusive volcanics (andesite and basalt) to various sedimentary and metamorphic species, including quartzite, argillite, siltstone, sandstone and schist. Soils are acidic (mean pH in 4.4 to 4.6 range), well-drained, and soil texture is usually on the coarser end of the spectrum (gravelly sandy loams and loams being common). Exposed rock and soil are generally less than 10%, but rock content in the profile often exceeds 20%.

Geographic Range: No Data Available

Nations: CA,US

States/Provinces:  AB, ID, MT, WA?




Confidence Level: Low

Confidence Level Comments: No Data Available

Grank: G3?

Greasons: No Data Available


Concept Lineage: No Data Available

Predecessors: No Data Available

Obsolete Names: No Data Available

Obsolete Parents: No Data Available

Synonomy: >< Abies lasiocarpa / Xerophyllum tenax Habitat Type, Vaccinium scoparium Phase (Pfister et al. 1977)
>< Abies lasiocarpa / Xerophyllum tenax Habitat Type, Vaccinium scoparium Phase (Cooper et al. 1987)
>< Abies lasiocarpa / Xerophyllum tenax Habitat Type, Vaccinium scoparium Phase (Steele et al. 1981)
= Pinus albicaulis - Abies lasiocarpa / Vaccinium scoparium / Xerophyllum tenax Woodland (Hop et al. 2007)
>< Tsuga mertensiana / Xerophyllum tenax Habitat Type (Pfister et al. 1977)
>< Tsuga mertensiana / Xerophyllum tenax Habitat Type, Vaccinium scoparium Phase (Cooper et al. 1987)

Concept Author(s): Hop et al. (2007)

Author of Description: S.V. Cooper

Acknowledgements: No Data Available

Version Date: 02-17-04

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