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CEGL005836 Pinus albicaulis - Abies lasiocarpa / Menziesia ferruginea / Xerophyllum tenax Woodland

Type Concept Sentence: No Data Available


Common (Translated Scientific) Name: Whitebark Pine - Subalpine Fir / Rusty Menziesia / Common Beargrass Woodland

Colloquial Name: No Data Available

Hierarchy Level:  Association

Type Concept: This minor small- to large-patch community is found primarily in mid to upper subalpine habitats throughout the Nez Perce and Clearwater national forests, north into Idaho''s panhandle and west into west-central Montana. East of the Continental Divide it extends northward to the Canadian Rockies of southwestern Alberta. Its distribution ranges in elevation from 1495 to 2195 m (4900-7200 feet). The lower elevation occurrences are associated with frost-pocket conditions. It usually occupies neutral to collecting positions from toeslopes to ridge shoulders and including ridgetops; it can be found on well-drained benches in cold-air drainages. Degree of slope is highly variable, but cold aspects (from northeast- through east-facing) are predominant. Parent materials are various, from intrusive volcanics of the Idaho Batholith to extrusive volcanics (andesite and basalt) to various sedimentary and metamorphic species, including quartzite, argillite, siltstone, sandstone and schist. Soils are acidic, well-drained, and soil texture is usually on the finer end of the spectrum (loams and silt loams being common). Exposed rock and soil are generally less than 5%, and the rock content in the profile often exceeds 5%, increasing markedly with depth. The overstory is usually an open canopy (less than 50% cover) dominated by a variable mix of Abies lasiocarpa, Picea engelmannii, Pinus contorta, and the indicator species Pinus albicaulis, which must have at least 5% cover for this type to be recognized as distinctive from ~Abies lasiocarpa - Picea engelmannii / Menziesia ferruginea / Xerophyllum tenax Forest (CEGL005895)$$ or the Pinus contorta-dominated version of this community. The undergrowth may be quite dense with a layer of short Menziesia ferruginea overtopping tussocks of Xerophyllum tenax. Vaccinium membranaceum is highly constant and though subordinate in height (and usually in cover) to Menziesia is still regarded a dominant short shrub. The only dwarf-shrub consistently present, Vaccinium scoparium, occurs as scattered patches; if present with greater than 15% cover, then another association is indicated. The combined cover of these four species often constitutes a continuous undulating layer. Spiraea betulifolia and Lonicera utahensis are consistently scattered in the short-shrub layer. There are no graminoids regularly present. Other than the abundance of the diagnostic Xerophyllum tenax, the forb layer has low cover and is comparatively depauperate with only Arnica latifolia, Anemone piperi, Goodyera oblongifolia, Orthilia secunda, and Viola orbiculata having a consistent presence. On moister sites and where the shrub layer is not overwhelming, Valeriana sitchensis and/or Thalictrum occidentale are consistently present with more than 5% cover.

Diagnostic Characteristics: No Data Available

Rationale for Nominal Species or Physiognomic Features: No Data Available

Classification Comments: Many to most of the stands within this association are merely seral representatives of ~Abies lasiocarpa - Picea engelmannii / Menziesia ferruginea / Xerophyllum tenax Forest (CEGL005895)$$; the association under consideration is recognized, at least in the Glacier-Waterton International Peace Park vegetation keys, as having at least 5% canopy cover of Pinus albicaulis. Recognition of a type based on a very minimal cover of a seral tree species (Pinus albicaulis) of broad distribution should provoke questions; the intent of this broadly inclusive type was to capture stands (specifically their locations, particularly in mapping efforts) wherein this highly threatened species occurs or will have once occurred.

Similar NVC Types: No Data Available
note: No Data Available

Physiognomy and Structure: No Data Available

Floristics: The overstory is usually an open canopy (less than 50% cover) dominated by a variable mix of Abies lasiocarpa, Picea engelmannii, Pinus contorta, and the indicator species Pinus albicaulis, which must have at least 5% cover for this type to be recognized as distinctive from ~Abies lasiocarpa - Picea engelmannii / Menziesia ferruginea / Xerophyllum tenax Forest (CEGL005895)$$ or the Pinus contorta-dominated version of this community. The undergrowth may be quite dense with a layer of Menziesia ferruginea (short-shrub height category in this restrictive environment) overtopping tussocks of Xerophyllum tenax. Vaccinium membranaceum is highly constant and though subordinate in height (and usually in cover) to Menziesia is still regarded as a dominant short shrub. In northern Idaho Rhododendron albiflorum may be a conspicuous component by virtue of its height (verging on tall-shrub category) and large white blossoms. The only dwarf-shrub consistently present, Vaccinium scoparium, occurs as scattered patches; if present with greater than 15% cover, then another association is indicated. The combined cover of these four species often constitutes a continuous undulating layer. Spiraea betulifolia and Lonicera utahensis are consistently scattered in the short-shrub layer. There are no graminoids regularly present. Other than the abundance of the diagnostic Xerophyllum tenax, the forb layer has low cover and is comparatively depauperate with only Arnica latifolia, Anemone piperi, Goodyera oblongifolia, Orthilia secunda (= Pyrola secunda), and Viola orbiculata having a consistent presence. On moister sites and where the shrub layer is not overwhelming, Valeriana sitchensis and/or Thalictrum occidentale are consistently present with more than 5% cover.

Dynamics:  Though there a number of stands supporting this type, it is expected to be a relatively rare type based on the fact that it is found primarily on cool to cold exposures. These sites are not chosen (explicitly avoided) when corvid birds bury Pinus albicaulis seed caches; these unexploited seed caches are the primary means by which this predominantly seral pine disperses to disturbed sites. This pine has also been documented to have become ecologically extinct in some ecosystems, such as Glacier National Park, due to the apparent virulence of white pine blister rust (Cronartium ribicola); there is little hope for natural recovery due to a very low rate of natural resistance.

Environmental Description:  This minor small- to large-patch community is found primarily in mid to upper subalpine habitats throughout the Nez Perce and Clearwater national forests, north into Idaho''s panhandle and west into west-central Montana; east of the Continental Divide it extends northward to the Canadian Rockies of southwestern Alberta. In the southern portion of its distribution elevations range from 1555 to 2105 m (5100-6900 feet), whereas in the north it ranges between 1495 to 1830 m (4900-6000 feet) and is really only regularly present above 1800 m (5900 feet), the lower elevation occurrences being associated with frost-pocket conditions. In Montana it is found primarily between 1675 and 2195 m (5500-7200 feet). It usually occupies neutral to collecting positions from toeslopes to ridge shoulders and including ridgetops; it can be found on well-drained benches in cold-air drainages. Degree of slope is highly variable, but cold aspects (from northeast- through east-facing) are predominant. Parent materials are various, from intrusive volcanics of the Idaho Batholith to extrusive volcanics (andesite and basalt) to various sedimentary and metamorphic species, including quartzite, argillite, siltstone, sandstone and schist. Soils are acidic (mean pH in 4.6 to 4.9 range), well-drained, and soil texture is usually on the finer end of the spectrum (loams and silt loams being common). Exposed rock and soil are generally less than 5%, and the rock content in the profile often exceeds 5% and increases markedly with depth.

Geographic Range: This minor small- to large-patch community is found primarily in mid to upper subalpine habitats throughout the Nez Perce and Clearwater national forests, north into Idaho''s panhandle and west into west-central Montana. East of the Continental Divide it extends northward to the Canadian Rockies of southwestern Alberta.

Nations: CA,US

States/Provinces:  AB, ID, MT




Confidence Level: Low

Confidence Level Comments: No Data Available

Grank: G3?

Greasons: No Data Available


Concept Lineage: No Data Available

Predecessors: No Data Available

Obsolete Names: No Data Available

Obsolete Parents: No Data Available

Synonomy: >< Abies lasiocarpa / Menziesia ferruginea Habitat Type, Xerophyllum tenax Phase (Cooper et al. 1987)
>< Abies lasiocarpa / Menziesia ferruginea Habitat Type (Pfister et al. 1977)
>< Picea engelmannii - Abies lasiocarpa / Menziesia ferruginea Habitat Type (Ogilvie 1962)
= Pinus albicaulis - Abies lasiocarpa / Menziesia ferruginea / Xerophyllum tenax Woodland (Hop et al. 2007)
>< C27: Picea engelmannii - Abies lasiocarpa / Menziesia ferruginea / Vaccinium membranaceum - Xerophyllum tenax Vegetation Type (Achuff et al. 2002)

Concept Author(s): Hop et al. (2007)

Author of Description: S.V. Cooper

Acknowledgements: No Data Available

Version Date: 02-17-04

  • ANHIC [Alberta Natural Heritage Information Centre]. 2018. Community database files. Alberta Natural Heritage Information Centre, Parks and Protected Areas Division, Alberta Community Development, Edmonton.
  • Achuff, P. L., R. L. McNeil, M. L. Coleman, C. Wallis and C. Wershler. 2002. Ecological land classification of Waterton Lakes National Park, Alberta. Volume I: Integrated resource description. Parks Canada, Waterton Lakes National Park, Alberta. 226 pp.
  • Cooper, S. V., K. E. Neiman, R. Steele, and D. W. Roberts. 1987. Forest habitat types of northern Idaho: A second approximation. General Technical Report INT-236.USDA Forest Service, Intermountain Research Station, Ogden, UT. 135 pp. [reprinted in 1991]
  • Hop, K., M. Reid, J. Dieck, S. Lubinski, and S. Cooper. 2007. U.S. Geological Survey-National Park Service Vegetation Mapping Program: Waterton-Glacier International Peace Park. U.S. Geological Survey, Upper Midwest Environmental Sciences Center, La Crosse, WI. 131 pp. plus Appendices A-L.
  • Ogilvie, R. T. 1962. Ecology of spruce forests on the east slope of the Rocky Mountains in Alberta. Unpublished dissertation, Washington State University, Pullman. 189 pp.
  • Pfister, R. D., B. L. Kovalchik, S. F. Arno, and R. C. Presby. 1977. Forest habitat types of Montana. General Technical Report INT-34. USDA Forest Service, Intermountain Forest and Range Experiment Station, Ogden, UT. 174 pp.
  • Reid, M. S., S. V. Cooper, and G. Kittel. 2004. Vegetation classification of Waterton-Glacier International Peace Park. Final report for USGS-NPS Vegetation Mapping Program, International Peace Park Mapping Project. NatureServe, Arlington VA.
  • Western Ecology Working Group of NatureServe. No date. International Ecological Classification Standard: International Vegetation Classification. Terrestrial Vegetation. NatureServe, Boulder, CO.