Print Report
CEGL005881 Larix occidentalis / Clintonia uniflora - Xerophyllum tenax Forest
Type Concept Sentence: No Data Available
Common (Translated Scientific) Name: Western Larch / Bride''s Bonnet - Common Beargrass Forest
Colloquial Name: No Data Available
Hierarchy Level: Association
Type Concept: This wholly seral, large-patch to matrix type occupies the relatively cold and dry environments across a number of climax tree series and associated geographic regions; the species defining these series include, but are not limited to, Thuja plicata, Tsuga heterophylla, Tsuga mertensiana, Abies grandis, Abies lasiocarpa, and Picea engelmannii. Thus this mesic type is found throughout the northern Rocky Mountains and may extend as far west as the Cascade Crest on environments characterized as foothills and montane to lower and even mid-subalpine. This association''s possible elevation range is from 915 to 1800 m (3000-5900 feet), and regardless of the climax series in which it is found, it consistently occurs on south- through west-facing exposures. The range of parent materials is, with the exception of highly unusual substrates like serpentine, literally as great as possible types occurring in the northern Rocky Mountains and northernmost middle Rocky Mountains and may include some ultramafics east of the Cascade Crest. It is difficult to characterize the soils as well, but they are uniformly well-drained and have a low coarse-fragment content, except those sites within the lower to mid-subalpine zone. The overstory is dominated by Pseudotsuga menziesii with a whole host of tree species capable of playing a subordinate role; on warmer sites these include Thuja plicata, Tsuga heterophylla, Abies grandis, and on colder or higher elevation sites are found Abies lasiocarpa, Tsuga mertensiana, and Picea engelmannii. However the most frequent canopy codominants or associates are the seral species Larix occidentalis, Pinus contorta, and in a restricted portion of the type''s range, Pinus monticola. The tall-shrub component is relatively unimportant, only Alnus viridis ssp. sinuata and Amelanchier alnifolia approach 50% constancy (and have low cover values). The short-shrub layer exhibits greater cover and diversity than the other shrub components with Vaccinium membranaceum, Paxistima myrsinites, Rosa gymnocarpa, Rubus parviflorus, and Spiraea betulifolia being consistently present. Linnaea borealis and Chimaphila umbellata have high constancy in the dwarf-shrub layer. Bromus vulgaris (or Bromus ciliatus) are the only graminoids of note. The diagnostic forbs Clintonia uniflora, Xerophyllum tenax, and Tiarella trifoliata naturally have high constancy and/or cover; however, a number of other forbs also exhibit high constancy, including Arnica latifolia, Aralia nudicaulis, Adenocaulon bicolor, Coptis occidentalis, Cornus canadensis, Galium triflorum, Goodyera oblongifolia, Maianthemum stellatum, Osmorhiza berteroi, Pedicularis racemosa, Orthilia secunda, Thalictrum occidentale, Trillium ovatum, and Viola orbiculata.
Diagnostic Characteristics: No Data Available
Rationale for Nominal Species or Physiognomic Features: No Data Available
Classification Comments: This association defines an extremely broad environmental range due to the relatively recent approach stipulated by the USNVC of defining as unique associations that would formerly have been subsumed within a number of different climax or potential vegetation types; in the case of this association it is a successional stage in environments as mild and wet (defining a Pacific maritime climatic influence) as those supporting Thuja plicata or Tsuga heterophylla / Clintonia uniflora associations (and permutations) to those of the lower subalpine zone characterized by Abies lasiocarpa / Clintonia uniflora, Abies grandis / Clintonia uniflora, Tsuga mertensiana / Clintonia uniflora, and even Picea engelmannii / Clintonia uniflora forest associations; however, by far the greatest proportion of this type is seral to ~Abies lasiocarpa - Picea engelmannii / Clintonia uniflora - Xerophyllum tenax Forest (CEGL005892)$$. That this association in fact occurs in the states and USFS sections listed derives from this writer''s interpretation of constancy/cover tables of various authors/publications; where Larix occidentalis is listed as a major seral species, the inference has been made that stands quite likely occur on the landscape where this species strongly dominates the upper canopy (often observed by this author throughout northern Idaho and western Montana). It should also be noted that this type is probably less common in zones where Thuja plicata and Tsuga heterophylla are the climax dominants because following disturbance in these environments the climax trees are quick to reclaim the site, i.e., they comprise a significant cover of the earliest forested successional stages. Considering Xerophyllum tenax and Vaccinium membranaceum as ecological analogues may be questionable when applied across the geographic ranges of these species; Vaccinium membranaceum is more broadly distributed, and thus this type could be recognized for northeastern Oregon, where little or no Xerophyllum occurs. The crosswalking for this type is incomplete, and it may well eventually be documented west of the Cascade Crest in Oregon, Washington and British Columbia.
Similar NVC Types: No Data Available
note: No Data Available
Physiognomy and Structure: No Data Available
Floristics: The overstory is dominated by seral Larix occidentalis with a whole host of tree species capable of playing a subordinate role; on warmer sites these include Thuja plicata, Tsuga heterophylla, Abies grandis, and on colder or higher elevation sites are found Abies lasiocarpa, Tsuga mertensiana, and Picea engelmannii. However, the most frequent canopy codominants or associates are the seral species Pseudotsuga menziesii, Pinus contorta, and in a restricted portion of the type''s range, Pinus monticola. The tall-shrub component is relatively unimportant, only Alnus viridis ssp. sinuata and Amelanchier alnifolia approach 50% constancy (and have low cover values). The short-shrub layer exhibits greater cover and diversity than the other shrub components with Vaccinium membranaceum, Paxistima myrsinites, Rosa gymnocarpa, Rubus parviflorus, and Spiraea betulifolia being consistently present. Linnaea borealis and Chimaphila umbellata have high constancy in the dwarf-shrub layer. Bromus vulgaris (or Bromus ciliatus) are the only graminoids of note. The diagnostic forbs Clintonia uniflora and Tiarella trifoliata between them are 100% constant though their cover seldom exceeds 10% singly or in combination. See Classification Comments for a more in-depth exposition on the reasons both Vaccinium membranaceum and Xerophyllum tenax are used as alternative indications of a particular subset of plots (and distinct environment) than characterized by Clintonia uniflora or Tiarella alone; in any given locality this type should be located at higher elevations and on warmer slopes with better drained soils than say Larix occidentalis / Clintonia uniflora. A number of other forbs also exhibit high constancy include Arnica latifolia, Aralia nudicaulis, Adenocaulon bicolor, Coptis occidentalis, Cornus canadensis, Galium triflorum, Goodyera oblongifolia, Maianthemum stellatum, Osmorhiza berteroi (= Osmorhiza chilensis), Pedicularis racemosa, Orthilia secunda (= Pyrola secunda), Thalictrum occidentale, Trillium ovatum, and Viola orbiculata. Within local landscapes some forbs exhibit high constancy not recorded in other regions, e.g., in northern Idaho Anemone piperi and Pyrola asarifolia are nearly always present.
Dynamics: This association is sufficiently mesic to support a host of tree species more shade-tolerant than Larix occidentalis (perhaps least shade-tolerant of western conifers, excepting those of considerably more xeric environments) and therefore the association is wholly a seral community type. Larix occidentalis is a long-lived species (in excess of 700 years in the northern Rocky Mountains) and thus stands fitting this concept are themselves long-persisting; the life of Larix-dominated stands probably does not much exceed 250 years due to various mortality sources and the ingrowth of shade-tolerant species. It has been noted in northern Idaho that following disturbance (particularly logging) in this type Larix occidentalis does not necessarily succeed itself, the first tree-dominated successional stages being dominated by Pseudotsuga menziesii, Pinus contorta, or less frequently by more shade-tolerant species (Cooper et al. 1987); this response is a consequence of the episodic nature of favorable cone crop years in Larix occidentalis.
Environmental Description: This wholly seral, large-patch to matrix type occupies the relatively cold and dry environments across a number of climax tree series and associated geographic regions; the species defining these series include, but are not limited to, Thuja plicata, Tsuga heterophylla, Tsuga mertensiana, Abies grandis, Abies lasiocarpa, and Picea engelmannii. Thus this mesic type is found throughout the northern Rocky Mountains and may extend as far west as the Cascade Crest on environments characterized as foothills and montane to lower and even mid-subalpine. This association''s possible elevation range is from 915 to 1800 m (3000-5900 feet). Regardless of the climax series in which it is found, it consistently occurs on south- through west-facing exposures. The range of parent materials is, with the exception of highly unusual substrates like serpentine, literally as great as possible types occurring in the northern Rocky Mountains and northernmost middle Rocky Mountains and may include some ultramafics east of the Cascade Crest. It is difficult to characterize the soils as well, but they are uniformly well-drained and have a low coarse-fragment content, except those sites within the lower to mid-subalpine zone.
Geographic Range: This association occurs from the Blue, Wallowa and Seven Devils mountains of northeastern Oregon and southern portion of the Idaho Batholith of central Idaho northward to the Colville National Forest of northeastern Washington, across northern Idaho and into western Montana (almost exclusively west of the Continental Divide). Given the opportunity for more complete crosswalking, this type might well be documented from British Columbia and the east slope of the Cascades; Lillybridge et al. (1995) do document both Clintonia uniflora and Larix occidentalis as common forest components from the eastern slope of the Cascades, but the two species do not uniquely overlap in space.
Nations: US
States/Provinces: ID, MT, OR, WA
Plot Analysis Summary:
http://vegbank.org/natureserve/ELEMENT_GLOBAL.2.728649
Confidence Level: Low
Confidence Level Comments: No Data Available
Grank: GNR
Greasons: No Data Available
Type | Name | Database Code | Classification Code |
---|---|---|---|
Class | 1 Forest & Woodland Class | C01 | 1 |
Subclass | 1.B Temperate & Boreal Forest & Woodland Subclass | S15 | 1.B |
Formation | 1.B.2 Cool Temperate Forest & Woodland Formation | F008 | 1.B.2 |
Division | 1.B.2.Nb Rocky Mountain Forest & Woodland Division | D194 | 1.B.2.Nb |
Macrogroup | 1.B.2.Nb.3 Western Hemlock - Grand Fir - Western Larch Mesic Lower Montane Forest Macrogroup | M500 | 1.B.2.Nb.3 |
Group | 1.B.2.Nb.3.a Grand Fir - Douglas-fir - Western Larch Central Rocky Mountain Forest Group | G211 | 1.B.2.Nb.3.a |
Alliance | A0275 Western Larch Central Rocky Mountain Forest Alliance | A0275 | 1.B.2.Nb.3.a |
Association | CEGL005881 Western Larch / Bride''s Bonnet - Common Beargrass Forest | CEGL005881 | 1.B.2.Nb.3.a |
Concept Lineage: This association and 3 others replace the placeholder.
Predecessors: No Data Available
Obsolete Names: No Data Available
Obsolete Parents: No Data Available
Synonomy: >< Abies grandis / Clintonia uniflora Habitat Type, Xerophyllum tenax Phase (Cooper et al. 1987)
>< Abies grandis / Vaccinium membranaceum / Clintonia uniflora Plant Association (Williams et al. 1995)
>< Abies lasiocarpa / Clintonia uniflora Habitat Type, Xerophyllum tenax Phase (Pfister et al. 1977)
>< Abies lasiocarpa / Clintonia uniflora Habitat Type, Xerophyllum tenax Phase (Cooper et al. 1987)
>< Larix occidentalis - Pseudotsuga menziesii (Pinus contorta) / Vaccinium membranaceum / Xerophyllum tenax (Leavell 2000)
= Larix occidentalis / Clintonia uniflora - Xerophyllum tenax Forest (Hop et al. 2007)
>< Thuja plicata / Clintonia uniflora Habitat Type, Xerophyllum tenax Phase (Cooper et al. 1987)
>< Tsuga heterophylla / Clintonia uniflora Habitat Type, Xerophyllum tenax Phase (Cooper et al. 1987)
>< Tsuga mertensiana / Clintonia uniflora Habitat Type, Xerophyllum tenax Phase (Cooper et al. 1987)
>< Tsuga mertensiana / Vaccinium membranaceum (Lillybridge et al. 1995) [TSME/VAME WEN ASSOCIATION CMS259]
>< Tsuga mertensiana / Xerophyllum tenax - Vaccinium myrtillus (Lillybridge et al. 1995) [TSME/XETE-VAMY ASSOCIATION CMF131]
>< Abies grandis / Vaccinium membranaceum / Clintonia uniflora Plant Association (Williams et al. 1995)
>< Abies lasiocarpa / Clintonia uniflora Habitat Type, Xerophyllum tenax Phase (Pfister et al. 1977)
>< Abies lasiocarpa / Clintonia uniflora Habitat Type, Xerophyllum tenax Phase (Cooper et al. 1987)
>< Larix occidentalis - Pseudotsuga menziesii (Pinus contorta) / Vaccinium membranaceum / Xerophyllum tenax (Leavell 2000)
= Larix occidentalis / Clintonia uniflora - Xerophyllum tenax Forest (Hop et al. 2007)
>< Thuja plicata / Clintonia uniflora Habitat Type, Xerophyllum tenax Phase (Cooper et al. 1987)
>< Tsuga heterophylla / Clintonia uniflora Habitat Type, Xerophyllum tenax Phase (Cooper et al. 1987)
>< Tsuga mertensiana / Clintonia uniflora Habitat Type, Xerophyllum tenax Phase (Cooper et al. 1987)
>< Tsuga mertensiana / Vaccinium membranaceum (Lillybridge et al. 1995) [TSME/VAME WEN ASSOCIATION CMS259]
>< Tsuga mertensiana / Xerophyllum tenax - Vaccinium myrtillus (Lillybridge et al. 1995) [TSME/XETE-VAMY ASSOCIATION CMF131]
- Cooper, S. V., K. E. Neiman, R. Steele, and D. W. Roberts. 1987. Forest habitat types of northern Idaho: A second approximation. General Technical Report INT-236.USDA Forest Service, Intermountain Research Station, Ogden, UT. 135 pp. [reprinted in 1991]
- Daubenmire, R. F., and J. B. Daubenmire. 1968. Forest vegetation of eastern Washington and northern Idaho. Washington State University Agricultural Experiment Station Technical Bulletin No. 60. 104 pp.
- Hop, K., M. Reid, J. Dieck, S. Lubinski, and S. Cooper. 2007. U.S. Geological Survey-National Park Service Vegetation Mapping Program: Waterton-Glacier International Peace Park. U.S. Geological Survey, Upper Midwest Environmental Sciences Center, La Crosse, WI. 131 pp. plus Appendices A-L.
- Johnson, C. G., Jr., and S. A. Simon. 1987. Plant associations of the Wallowa-Snake Province Wallowa-Whitman National Forest. Technical Paper R6-ECOL-TP-255A-86. USDA Forest Service, Pacific Northwest Region, Wallowa-Whitman National Forest. 399 pp. plus appendices.
- Johnson, C. G., and R. R. Clausnitzer. 1992. Plant associations of the Blue and Ochoco mountains. R6-ERW-TP-036-92. USDA Forest Service, Pacific Northwest Region, Wallowa-Whitman National Forest. 163 pp. plus appendices.
- Leavell, D. 2000. Vegetation and process of the Kootenai National Forest. Ph.D. dissertation, University of Montana, Missoula. 508 pp.
- Lillybridge, T. R., B. L. Kovalchik, C. K. Williams, and B. G. Smith. 1995. Field guide for forested plant associations of the Wenatchee National Forest. General Technical Report PNW-GTR-359. USDA Forest Service, Pacific Northwest Research Station, Portland, OR. 335 pp.
- Pfister, R. D., B. L. Kovalchik, S. F. Arno, and R. C. Presby. 1977. Forest habitat types of Montana. General Technical Report INT-34. USDA Forest Service, Intermountain Forest and Range Experiment Station, Ogden, UT. 174 pp.
- Reid, M. S., S. V. Cooper, and G. Kittel. 2004. Vegetation classification of Waterton-Glacier International Peace Park. Final report for USGS-NPS Vegetation Mapping Program, International Peace Park Mapping Project. NatureServe, Arlington VA.
- Western Ecology Working Group of NatureServe. No date. International Ecological Classification Standard: International Vegetation Classification. Terrestrial Vegetation. NatureServe, Boulder, CO.
- Williams, C. K., B. F. Kelly, B. G. Smith, and T. R. Lillybridge. 1995. Forest plant associations of the Colville National Forest. General Technical Report PNW-GTR-360. USDA Forest Service, Pacific Northwest Region, Portland, OR. 140 pp.