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CEGL005880 Larix occidentalis / Clintonia uniflora Forest
Type Concept Sentence: No Data Available
Common (Translated Scientific) Name: Western Larch / Bride''s Bonnet Forest
Colloquial Name: No Data Available
Hierarchy Level: Association
Type Concept: Broadly distributed throughout the northern Rocky Mountains and adjacent terrain, this large-patch to matrix seral community occupies relatively moist (mesic) and warm to cool sites having free air drainage and lacking frost pocket conditions. Elevations range in the north from760 to 1585 m (4500-5200 feet) (extreme outliers at 1710 m (5600 feet)), whereas to the south it ranges from 1060 to 1710 m (3500-5600 feet). It occurs on slopes of all degrees of steepness and aspect orientation, though it is more likely to occur from toeslope through midslope positions (predominantly collecting positions). At the dry extreme of its distribution it is more strongly associated with protected positions such as concave slopes, moist depressions on gentle slopes or plateaus, stringers along perennial stream bottoms, toeslopes and northeastern aspects. Though sites are mesic, verging on hygric, they are inferred to be only briefly or seasonally influenced, if at all, by a high water table; Larix occidentalis occurrence is strongly associated with well-drained positions. A wide variety of parent materials are represented, including those as disparate as granite and limestone, including all manner of glacial-fluvial material. In northern Idaho and northwestern Montana it is routinely found on ash caps, ranging from 3 to 60 cm in depth. The soil textures are predominantly loams and silt loams (reflecting in part the volcanic ash); soils typically have less than 15% coarse fragment content and are well-drained.
This mesic seral association is characterized by Larix occidentalis dominating the upper canopy, though other tree species occur with lesser cover, including both other species considered almost exclusively seral Pinus contorta and Pinus monticola and those capable of functioning as both seral and climax species, including those from warmer environments, Pinus ponderosa (very limited representation), Pseudotsuga menziesii, Thuja plicata, and Tsuga heterophylla and those of colder environments, Abies lasiocarpa, Abies grandis, and Picea engelmannii. The shrub layer may be highly diverse with tall shrubs (e.g., Acer glabrum, Taxus brevifolia, Amelanchier alnifolia), short shrubs (Symphoricarpos albus, Paxistima myrsinites, Rubus parviflorus, Spiraea betulifolia), and dwarf-shrubs (e.g., Chimaphila umbellata, Linnaea borealis, Mahonia repens) abundantly represented. The graminoid component is inconspicuous with no one species exhibiting high constancy, though Bromus vulgaris, Bromus ciliatus, and Calamagrostis rubescens are more consistently present and with greater cover than other graminoids. The cover of the diagnostic forbs Clintonia uniflora and Tiarella trifoliata is greatest when this type occurs in the zones potentially dominated by Thuja plicata and Tsuga heterophylla, up to 30% canopy cover (can even be a dominant forb), whereas in the colder environments characterized by Abies lasiocarpa, Abies grandis, and Picea engelmannii potential dominance cover of these diagnostics and all forbs is generally less. Other forbs of high constancy, at least in some portion of this association''s considerable range, are Aralia nudicaulis, Adenocaulon bicolor, Coptis occidentalis, Cornus canadensis, Galium triflorum, Goodyera oblongifolia, Maianthemum stellatum, Osmorhiza berteroi, Orthilia secunda, Thalictrum occidentale, Trillium ovatum, Viola glabella (or Viola canadensis), and Viola orbiculata.
This mesic seral association is characterized by Larix occidentalis dominating the upper canopy, though other tree species occur with lesser cover, including both other species considered almost exclusively seral Pinus contorta and Pinus monticola and those capable of functioning as both seral and climax species, including those from warmer environments, Pinus ponderosa (very limited representation), Pseudotsuga menziesii, Thuja plicata, and Tsuga heterophylla and those of colder environments, Abies lasiocarpa, Abies grandis, and Picea engelmannii. The shrub layer may be highly diverse with tall shrubs (e.g., Acer glabrum, Taxus brevifolia, Amelanchier alnifolia), short shrubs (Symphoricarpos albus, Paxistima myrsinites, Rubus parviflorus, Spiraea betulifolia), and dwarf-shrubs (e.g., Chimaphila umbellata, Linnaea borealis, Mahonia repens) abundantly represented. The graminoid component is inconspicuous with no one species exhibiting high constancy, though Bromus vulgaris, Bromus ciliatus, and Calamagrostis rubescens are more consistently present and with greater cover than other graminoids. The cover of the diagnostic forbs Clintonia uniflora and Tiarella trifoliata is greatest when this type occurs in the zones potentially dominated by Thuja plicata and Tsuga heterophylla, up to 30% canopy cover (can even be a dominant forb), whereas in the colder environments characterized by Abies lasiocarpa, Abies grandis, and Picea engelmannii potential dominance cover of these diagnostics and all forbs is generally less. Other forbs of high constancy, at least in some portion of this association''s considerable range, are Aralia nudicaulis, Adenocaulon bicolor, Coptis occidentalis, Cornus canadensis, Galium triflorum, Goodyera oblongifolia, Maianthemum stellatum, Osmorhiza berteroi, Orthilia secunda, Thalictrum occidentale, Trillium ovatum, Viola glabella (or Viola canadensis), and Viola orbiculata.
Diagnostic Characteristics: No Data Available
Rationale for Nominal Species or Physiognomic Features: No Data Available
Classification Comments: This association defines an extremely broad environmental range due to the relatively recent approach stipulated by the USNVC of defining as unique associations that would formerly have been subsumed within a number of different climax or potential vegetation types; in the case of this association it is a successional stage in environments as mild and wet (defining a Pacific maritime climatic influence) as those supporting Thuja plicata or Tsuga heterophylla / Clintonia uniflora associations (and permutations) to those of the lower subalpine zone characterized by Abies lasiocarpa, Abies grandis, Tsuga mertensiana, and even Picea engelmannii / Clintonia uniflora forest associations. That this association in fact occurs in the states and USFS sections listed derives from this writer''s interpretation of constancy/cover tables of various authors/publications; where Larix occidentalis is listed as a major seral species, the inference has been made that stands quite likely occur on the landscape where this species strongly dominates the upper canopy (often observed by this author throughout northern Idaho and western Montana). It should also be noted that this type is probably less common in zones where Thuja plicata and Tsuga heterophylla are the climax dominants because, following disturbance in these zones, the climax trees are quick to reclaim the site, i.e., they comprise a significant cover of the earliest forested successional stages. The crosswalking for this type is incomplete, and it may well eventually be documented west of the Cascade Crest in Oregon and Washington.
Similar NVC Types: No Data Available
note: No Data Available
Physiognomy and Structure: No Data Available
Floristics: This mesic seral association is characterized by the dominance of Larix occidentalis in the upper canopy, though other tree species occur with lesser cover, including both species considered almost exclusively seral, Pinus contorta and Pinus monticola, and those capable of functioning as both seral and climax species, including those from warmer environments, Pinus ponderosa (very limited representation), Pseudotsuga menziesii, Thuja plicata, and Tsuga heterophylla and those of colder environments, Abies lasiocarpa, Abies grandis, and Picea engelmannii. The shrub layer may be highly diverse with tall shrubs (e.g., Acer glabrum, Taxus brevifolia, Amelanchier alnifolia), short shrubs (Symphoricarpos albus, Paxistima myrsinites, Rubus parviflorus, Spiraea betulifolia), and dwarf-shrubs (e.g., Chimaphila umbellata, Linnaea borealis, Mahonia repens) abundantly represented. The graminoid component is inconspicuous with no one species exhibiting high constancy, though Bromus vulgaris, Bromus ciliatus, and Calamagrostis rubescens are more consistently present and with greater cover than other graminoids. The cover of the diagnostic forbs Clintonia uniflora and Tiarella trifoliata is greatest when this type occurs in the zones potentially dominated by Thuja plicata and Tsuga heterophylla, up to 30% canopy cover (can even be a dominant forb), whereas in the colder environments characterized by Abies lasiocarpa, Abies grandis, and Picea engelmannii potential dominance cover of these diagnostics and all forbs is generally less. Other forbs of high constancy, at least in some portion of this association''s considerable range, are Aralia nudicaulis, Adenocaulon bicolor, Coptis occidentalis, Cornus canadensis, Galium triflorum, Goodyera oblongifolia, Maianthemum stellatum, Osmorhiza berteroi (= Osmorhiza chilensis), Orthilia secunda (= Pyrola secunda), Thalictrum occidentale, Trillium ovatum, Viola glabella (or Viola canadensis), and Viola orbiculata.
Dynamics: This association is sufficiently mesic to support a host of tree species more shade-tolerant than Larix occidentalis (perhaps least shade-tolerant of western conifers, excepting those of considerably more xeric environments) and therefore the association is wholly a seral community type. Larix occidentalis is a long-lived species (in excess of 700 years in the northern Rocky Mountains) and thus stands fitting this concept are themselves long-persisting; the life of Larix-dominated stands probably does not much exceed 250 years due to various mortality sources and the ingrowth of shade-tolerant species. It has been noted in northern Idaho that following disturbance (particularly logging) in this type that Larix occidentalis often does not succeed itself, the first tree-dominated successional stages being dominated by Pseudotsuga menziesii, Pinus contorta, or less frequently by more shade-tolerant species (Cooper et al. 1987); this response is a consequence of the episodic nature of favorable cone crop years in Larix occidentalis.
Environmental Description: Broadly distributed throughout the northern Rocky Mountains and adjacent Okanogan Highlands of Washington, Blue Mountains of northeastern Oregon, and Wallowa and Seven Devils uplifts of northeastern Oregon and central Idaho, this large-patch to matrix seral community occupies relatively moist (mesic) and warm to cool sites having free air drainage, and lacking frost pocket conditions. Elevations range in the north from 760 to 1585 m (4500-5200 feet) (extreme outliers at 1710 m (5600 feet)), whereas to the south it ranges from 1060 to 1710 m (3500-5600 feet). It occurs on slopes of all degrees of steepness and aspect orientation, though it is more likely to occur from toeslope through midslope positions (predominantly collecting positions). At the dry extreme of its distribution it is more strongly associated with protected positions such as concave slopes, moist depressions on gentle slopes or plateaus, stringers along perennial stream bottoms, toeslopes and northeastern aspects. Though sites are mesic, verging on hygric, they are inferred to be only briefly or seasonally influenced, if at all, by a high water table; Larix occidentalis occurrence is strongly associated with well-drained positions. A wide variety of parent materials are represented, including those as disparate as granite and limestone, including all manner of glacial-fluvial material. In northern Idaho and northwestern Montana it is routinely found on ash caps, ranging from 3 to 60 cm in depth. The soil textures are predominantly loams and silt loams (reflecting in part the volcanic ash); soils typically have less than 15% coarse fragment content and are well-drained.
Geographic Range: This association occurs from the Blue, Wallowa and Seven Devils mountains of northeastern Oregon and southern portion of the Idaho Batholith of central Idaho northward to the Colville National Forest of northeastern Washington, across northern Idaho and into western Montana (almost exclusively west of the Continental Divide). Given the opportunity for more complete crosswalking, this type might well be documented from British Columbia and the east slope of the Cascades; Lillybridge et al. (1995) do document both Clintonia uniflora and Larix occidentalis as common forest components from the eastern slope of Cascades, but the two species do not uniquely overlap in space.
Nations: CA?,US
States/Provinces: ID, MT, OR, WA
Plot Analysis Summary:
http://vegbank.org/natureserve/ELEMENT_GLOBAL.2.728642
Confidence Level: Low
Confidence Level Comments: No Data Available
Grank: GNR
Greasons: No Data Available
Type | Name | Database Code | Classification Code |
---|---|---|---|
Class | 1 Forest & Woodland Class | C01 | 1 |
Subclass | 1.B Temperate & Boreal Forest & Woodland Subclass | S15 | 1.B |
Formation | 1.B.2 Cool Temperate Forest & Woodland Formation | F008 | 1.B.2 |
Division | 1.B.2.Nb Rocky Mountain Forest & Woodland Division | D194 | 1.B.2.Nb |
Macrogroup | 1.B.2.Nb.3 Western Hemlock - Grand Fir - Western Larch Mesic Lower Montane Forest Macrogroup | M500 | 1.B.2.Nb.3 |
Group | 1.B.2.Nb.3.a Grand Fir - Douglas-fir - Western Larch Central Rocky Mountain Forest Group | G211 | 1.B.2.Nb.3.a |
Alliance | A0275 Western Larch Central Rocky Mountain Forest Alliance | A0275 | 1.B.2.Nb.3.a |
Association | CEGL005880 Western Larch / Bride''s Bonnet Forest | CEGL005880 | 1.B.2.Nb.3.a |
Concept Lineage: This association and 3 others replace the placeholder.
Predecessors: No Data Available
Obsolete Names: No Data Available
Obsolete Parents: No Data Available
Synonomy: >< Abies grandis / Acer glabrum / Clintonia uniflora Plant Association (Williams et al. 1995)
>< Abies grandis / Clintonia uniflora Habitat Type, Clintonia uniflora Phase (Cooper et al. 1987)
>< Abies grandis / Clintonia uniflora Habitat Type, Clintonia uniflora Phase (Pfister et al. 1977)
>< Abies grandis / Clintonia uniflora Plant Association (Johnson and Clausnitzer 1992)
>< Abies grandis / Clintonia uniflora Plant Association (Johnson and Simon 1987)
>< Abies lasiocarpa / Clintonia uniflora Habitat Type, Clintonia uniflora Phase (Cooper et al. 1987)
>< Abies lasiocarpa / Clintonia uniflora Habitat Type, Clintonia uniflora Phase (Pfister et al. 1977)
>< Abies lasiocarpa / Clintonia uniflora Habitat Type, Clintonia uniflora Phase (Steele et al. 1981)
>< Abies lasiocarpa / Clintonia uniflora Plant Association (Johnson and Simon 1987)
>< Abies lasiocarpa / Clintonia uniflora Plant Association (Williams et al. 1995)
>< Larix occidentalis - Betula papyrifera (Populus tremuloides) / Acer glabrum - Alnus sinuata (Leavell 2000)
= Larix occidentalis / Clintonia uniflora Forest (Hop et al. 2007)
>< Thuja plicata / Clintonia uniflora Habitat Type, Clintonia uniflora Phase (Pfister et al. 1977)
>< Thuja plicata / Clintonia uniflora Habitat Type, Clintonia uniflora Phase (Cooper et al. 1987)
>< Tsuga heterophylla / Acer circinatum / Clintonia uniflora (Lillybridge et al. 1995) [TSHE/ACCI/CLUN ASSOCIATION CHS227]
>< Tsuga heterophylla / Clintonia uniflora Habitat Type, Clintonia uniflora Phase (Pfister et al. 1977)
>< Tsuga heterophylla / Clintonia uniflora Habitat Type, Clintonia uniflora Phase (Cooper et al. 1987)
>< Tsuga heterophylla / Pachistima myrsinites / Clintonia uniflora (Lillybridge et al. 1995) [TSHE/PAMY/CLUN ASSOCIATION CHS143]
>< Abies grandis / Clintonia uniflora Habitat Type, Clintonia uniflora Phase (Cooper et al. 1987)
>< Abies grandis / Clintonia uniflora Habitat Type, Clintonia uniflora Phase (Pfister et al. 1977)
>< Abies grandis / Clintonia uniflora Plant Association (Johnson and Clausnitzer 1992)
>< Abies grandis / Clintonia uniflora Plant Association (Johnson and Simon 1987)
>< Abies lasiocarpa / Clintonia uniflora Habitat Type, Clintonia uniflora Phase (Cooper et al. 1987)
>< Abies lasiocarpa / Clintonia uniflora Habitat Type, Clintonia uniflora Phase (Pfister et al. 1977)
>< Abies lasiocarpa / Clintonia uniflora Habitat Type, Clintonia uniflora Phase (Steele et al. 1981)
>< Abies lasiocarpa / Clintonia uniflora Plant Association (Johnson and Simon 1987)
>< Abies lasiocarpa / Clintonia uniflora Plant Association (Williams et al. 1995)
>< Larix occidentalis - Betula papyrifera (Populus tremuloides) / Acer glabrum - Alnus sinuata (Leavell 2000)
= Larix occidentalis / Clintonia uniflora Forest (Hop et al. 2007)
>< Thuja plicata / Clintonia uniflora Habitat Type, Clintonia uniflora Phase (Pfister et al. 1977)
>< Thuja plicata / Clintonia uniflora Habitat Type, Clintonia uniflora Phase (Cooper et al. 1987)
>< Tsuga heterophylla / Acer circinatum / Clintonia uniflora (Lillybridge et al. 1995) [TSHE/ACCI/CLUN ASSOCIATION CHS227]
>< Tsuga heterophylla / Clintonia uniflora Habitat Type, Clintonia uniflora Phase (Pfister et al. 1977)
>< Tsuga heterophylla / Clintonia uniflora Habitat Type, Clintonia uniflora Phase (Cooper et al. 1987)
>< Tsuga heterophylla / Pachistima myrsinites / Clintonia uniflora (Lillybridge et al. 1995) [TSHE/PAMY/CLUN ASSOCIATION CHS143]
- Cooper, S. V., K. E. Neiman, R. Steele, and D. W. Roberts. 1987. Forest habitat types of northern Idaho: A second approximation. General Technical Report INT-236.USDA Forest Service, Intermountain Research Station, Ogden, UT. 135 pp. [reprinted in 1991]
- Daubenmire, R. F., and J. B. Daubenmire. 1968. Forest vegetation of eastern Washington and northern Idaho. Washington State University Agricultural Experiment Station Technical Bulletin No. 60. 104 pp.
- Hop, K., M. Reid, J. Dieck, S. Lubinski, and S. Cooper. 2007. U.S. Geological Survey-National Park Service Vegetation Mapping Program: Waterton-Glacier International Peace Park. U.S. Geological Survey, Upper Midwest Environmental Sciences Center, La Crosse, WI. 131 pp. plus Appendices A-L.
- Johnson, C. G., Jr., and S. A. Simon. 1987. Plant associations of the Wallowa-Snake Province Wallowa-Whitman National Forest. Technical Paper R6-ECOL-TP-255A-86. USDA Forest Service, Pacific Northwest Region, Wallowa-Whitman National Forest. 399 pp. plus appendices.
- Johnson, C. G., and R. R. Clausnitzer. 1992. Plant associations of the Blue and Ochoco mountains. R6-ERW-TP-036-92. USDA Forest Service, Pacific Northwest Region, Wallowa-Whitman National Forest. 163 pp. plus appendices.
- Leavell, D. 2000. Vegetation and process of the Kootenai National Forest. Ph.D. dissertation, University of Montana, Missoula. 508 pp.
- Lillybridge, T. R., B. L. Kovalchik, C. K. Williams, and B. G. Smith. 1995. Field guide for forested plant associations of the Wenatchee National Forest. General Technical Report PNW-GTR-359. USDA Forest Service, Pacific Northwest Research Station, Portland, OR. 335 pp.
- Pfister, R. D., B. L. Kovalchik, S. F. Arno, and R. C. Presby. 1977. Forest habitat types of Montana. General Technical Report INT-34. USDA Forest Service, Intermountain Forest and Range Experiment Station, Ogden, UT. 174 pp.
- Reid, M. S., S. V. Cooper, and G. Kittel. 2004. Vegetation classification of Waterton-Glacier International Peace Park. Final report for USGS-NPS Vegetation Mapping Program, International Peace Park Mapping Project. NatureServe, Arlington VA.
- Steele, R., R. D. Pfister, R. A. Ryker, and J. A. Kittams. 1981. Forest habitat types of central Idaho. General Technical Report INT-114. USDA Forest Service, Intermountain Forest and Range Experiment Station, Ogden, UT. 138 pp.
- Western Ecology Working Group of NatureServe. No date. International Ecological Classification Standard: International Vegetation Classification. Terrestrial Vegetation. NatureServe, Boulder, CO.
- Williams, C. K., B. F. Kelly, B. G. Smith, and T. R. Lillybridge. 1995. Forest plant associations of the Colville National Forest. General Technical Report PNW-GTR-360. USDA Forest Service, Pacific Northwest Region, Portland, OR. 140 pp.