Print Report

CEGL001203 Salix exigua / Mesic Graminoids Western Wet Shrubland

Type Concept Sentence: A common riparian shrubland found throughout the western U.S., dominated by Salix exigua with a sparse to dense graminoid understory.


Common (Translated Scientific) Name: Narrowleaf Willow / Mesic Graminoids Western Wet Shrubland

Colloquial Name: No Data Available

Hierarchy Level:  Association

Type Concept: This riparian association is found primarily in the Rocky Mountains and Intermountain West semi-desert regions, and extending into California. The vegetation is characterized by the dominance of Salix exigua in a moderately dense tall-shrub canopy with a dense herbaceous layer dominated by graminoids. It typically occurs as a monoculture of Salix exigua, but can have other woody species, including saplings of Populus deltoides or Salix amygdaloides, and shrubs such as Salix eriocephala, Salix lutea, and Amorpha fruticosa. Tall perennial grasses can appear to codominate the stand when Spartina pectinata, Panicum virgatum or other tall grasses are present. Other mesic graminoids, such as Carex spp., Eleocharis spp., Juncus spp., Pascopyrum smithii, Schoenoplectus pungens, and Sphenopholis obtusata, may be present. Common forb species include Bidens spp., Lobelia siphilitica, Lycopus americanus, Lythrum alatum, Polygonum spp., and Xanthium strumarium. Diagnostic features of this association include the nearly pure stands of Salix exigua shrubs, with a dense herbaceous layer of at least 30% cover of mesic graminoids. It generally occurs along backwater channels and other perennially wet but less scoured sites, such as floodplain swales and irrigation ditches.

Diagnostic Characteristics: Diagnostic features of this association include the nearly pure stands of Salix exigua shrubs, with a dense ground layer of at least 30% cover of graminoids.

Rationale for Nominal Species or Physiognomic Features: No Data Available

Classification Comments: This broadly defined plant association occupies a rather large range and currently includes Salix exigua shrublands with herbaceous layers dominated by the introduced mesic, perennial, sod-grass Poa pratensis. Classification review has determined that western stands belong to Salix exigua Nutt. sensu stricto, and Great Plains stands belong to Salix interior (and their intermediates). Salix interior is an entirely Great Plains and eastward species (Kartesz 1999). In Nebraska, this community intergrades and is a successional stage that appears after both ~Riverine Sand Flats - Bars Sparse Vegetation (CEGL002049)$$ and ~Salix exigua Riparian Wet Shrubland (CEGL001197)$$, which is more frequently disturbed and lacks many of the more mesic herbaceous species.

Similar NVC Types: No Data Available
note: No Data Available

Physiognomy and Structure: Medium to tall multi-stemmed shrubs, ranging from 1 to 5 m in height, often forming dense clumps of clonal growth, with a sparse graminoid understory.

Floristics: This association is characterized by the dominance of Salix exigua in the moderately dense tall-shrub canopy with a dense herbaceous layer dominated by graminoids. Others common woody species may include saplings of Populus deltoides and Salix amygdaloides, and shrubs such as Salix bebbiana, Salix eriocephala, Salix geyeriana, Salix lucida ssp. lasiandra (= Salix lasiandra), Salix lutea, Salix monticola, Salix planifolia, Amorpha fruticosa, or Rosa woodsii. Scattered individuals of Populus fremontii, Salix gooddingii, and Elaeagnus angustifolia may contribute low cover to an emergent canopy layer. Tall perennial graminoids can be dominant in the undergrowth and include Carex pellita (= Carex lanuginosa), Carex nebrascensis, Carex rostrata, Deschampsia cespitosa, Eleocharis palustris, Elymus canadensis, Equisetum spp., Glyceria spp., Juncus arcticus ssp. littoralis (= Juncus balticus), Juncus longistylis, Juncus tenuis, Juncus torreyi, Luzula parviflora, Pascopyrum smithii, Polygonum spp., Schoenoplectus americanus, Schoenoplectus pungens (= Scirpus pungens), Sphenopholis obtusata, and others. The sparse forb cover may include Lobelia siphilitica, Bidens spp., Geum macrophyllum, Lycopus americanus, Lythrum alatum, Mentha arvensis, Typha angustifolia, Veronica americana, and Xanthium strumarium. Introduced forage species such as Agrostis stolonifera, Bromus inermis, Melilotus officinalis (= Melilotus albus), Medicago sativa, Salsola tragus, Poa pratensis, or Phleum pratense may be present to abundant in disturbed stands of this community. Steinauer and Rolfsmeier (1997) note that short-lived stands of Salix eriocephala are found along the Platte River.

Dynamics:  Flooding during spring periods is common, although scouring is rare. This plant association is considered early-seral typical of recent floodplains and highly disturbed, low, wet areas. The presence of Populus sp. seedlings within this association indicates succession to a cottonwood stand. Overgrazing by livestock will reduce the vigor of Salix exigua and may eventually eliminate it from the site and allow invasion of introduced and non-palatable native species. However, reducing stocking rates will allow Salix exigua to re-establish itself, provided it has not been completely eliminated from the site (Hansen et al. 1995). Tamarix ramosissima has the potential to invade and even dominate stands of this community.

Environmental Description:  This riparian association is found along alluvial terraces of backwater channels and other perennially wet but less scoured sites such as floodplain swales and irrigation ditches. It is found at elevations from 937 to 2700 m (3075-9100 feet). This community is found on temporarily to seasonally flooded sandbars, islands, and shorelines of stream channels and braided rivers. Stands usually occur within 1 m vertical distance of the stream channel on point bars, low floodplains, terraces and along overflow channels. It can also occur away from the stream channel in mesic swales or along the margins of beaver ponds and seeps. Sites are usually relatively flat, though rarely they can have moderate or moderately steep slopes. Soils are derived from alluvium and are quite variable in development, ranging from thin (<1 m) and skeletal with depth (10-50% cobbles) to well-developed Mollisols (Kittel et al. 1999a). Textures are typically loamy sands interspersed with layers of silty clays and alternating with coarse sands. Upper layers (10-30 cm) often have 25-30% organic matter (Kittel et al. 1999a), and organic litter covers most of the unvegetated ground surface.

Geographic Range: This association is found primarily in the Rocky Mountains and Intermountain semi-desert regions, ranging from western Wyoming and Montana west to Idaho, Oregon, Washington, California, and including Nevada, Utah, Arizonica, Colorado and western New Mexico.

Nations: US

States/Provinces:  CA, CO, ID, MT, NM, NV, UT, WY




Confidence Level: High

Confidence Level Comments: No Data Available

Grank: G5

Greasons: No Data Available


Concept Lineage: No Data Available

Predecessors: No Data Available

Obsolete Names: No Data Available

Obsolete Parents: No Data Available

Synonomy: = Salix exigua / Carex sheldonii Association (Crowe et al. 2004)
= Salix exigua / Juncus spp. (Sawyer et al. 2009) [61.209.05]
= Salix exigua / Mesic Graminoids Shrubland (Faber-Langendoen 2001)
= Salix exigua / Mesic graminoids Association (Evens et al. 2014)
= Salix exigua / Mesic Graminoids Community Type (Padgett et al. 1989)
= Salix exigua / Mesic Graminoids Community Type (Jones and Walford 1995)
= Salix exigua / Mesic graminoid Community Type (Padgett et al. 1988b)
= Salix exigua / Mesic graminoid Shrubland (Kittel and Lederer 1993)
= Salix exigua / Mesic graminoid Shrubland (Kittel et al. 1996)
= Salix exigua/Juncus spp. Shrubland [Provisional] (Keeler-Wolf et al. 2012)
? Salix exigua - Poa pratensis Association (Cooper and Cottrell 1990)
? Salix exigua - Poa pratensis Community (Jones and Walford 1995)
= Salix exigua - Poa pratensis Community Type (Walford et al. 2001)
= Salix exigua / Mesic Graminoid Shrubland (Carsey et al. 2003a)
= Salix exigua / Mesic Graminoids Shrubland (Kittel 1994)
< Salix exigua Community Type (Hansen et al. 1995)
= Coyote willow/Mesic Graminoids (Salix exigua/Mesic Graminoids) Plant Association (Kittel et al. 1999a)

Concept Author(s): Western Ecology Group

Author of Description: K.A. Schulz, J. Drake, J. Coles and G. Kittel

Acknowledgements: No Data Available

Version Date: 07-14-16

  • Bourgeron, P. S., and L. D. Engelking, editors. 1994. A preliminary vegetation classification of the western United States. Unpublished report. The Nature Conservancy, Western Heritage Task Force, Boulder, CO. 175 pp. plus appendix.
  • CNHP [Colorado Natural Heritage Program]. 2006-2017. Tracked natural plant communities. Colorado Natural Heritage Program, Colorado State University, Fort Collins. [https://cnhp.colostate.edu/ourdata/trackinglist/plant_communities/]
  • Carsey, K., G. Kittel, K. Decker, D. J. Cooper, and D. Culver. 2003a. Field guide to the wetland and riparian plant associations of Colorado. Colorado Natural Heritage Program, Fort Collins, CO.
  • Clark, D., M. Dela Cruz, T. Clark, J. Coles, S. Topp, A. Evenden, A. Wight, G. Wakefield, and J. Von Loh. 2009. Vegetation classification and mapping project report, Capitol Reef National Park. Natural Resource Report NPS/NCPN/NRTR--2009/187. National Park Service, Fort Collins, CO. 882 pp.
  • Cogan, D., J. E. Taylor, and K. Schulz. 2012. Vegetation inventory project: Great Basin National Park. Natural Resource Report NPS/MOJN/NRR--2012/568. National Park Service, Fort Collins, CO. 373 pp.
  • Cogan, D., M. Reid, K. Schulz, and M. Pucherelli. 2004. Zion National Park, Utah 1999-2003. Vegetation Mapping Project. Technical Memorandum 8260-03-01. Remote Sensing and GIS Group Technical Service Center, Bureau of Reclamation, Denver, CO. Appendix F: Vegetation Association Descriptions for Zion.
  • Coles, J., A. Tendick, J. Von Loh, G. Bradshaw, G. Manis, A. Wight, G. Wakefield, and A. Evenden. 2010. Vegetation classification and mapping project report, Black Canyon of the Gunnison National Park. Natural Resource Technical Report NPS/NCPN/NRTR--2010/361. National Park Service, Fort Collins, CO.
  • Coles, J., D. Cogan, D. Salas, A. Wight, G. Wakefield, J. Von Loh, and A. Evenden. 2008a. Vegetation classification and mapping project report, Dinosaur National Monument. Natural Resource Technical Report NPS/NCPN/NRTR-2008/112. National Park Service, Fort Collins, CO. 814 pp.
  • Coles, J., J. Von Loh, A. Evenden, G. Manis, G. Wakefield. and A. Wight. 2008c. Vegetation classification and mapping project report, Natural Bridges National Monument. Natural Resource Technical Report NPS/NCPN/NRTR--2008/077. National Park Service, Fort Collins, CO. 390 pp.
  • Cooper, D. J., and T. R. Cottrell. 1990. Classification of riparian vegetation in the northern Colorado Front Range. Unpublished report prepared for The Nature Conservancy, Colorado Field Office, Boulder. 115 pp.
  • Crowe, E. A., B. L. Kovalchik, and M. J. Kerr. 2004. Riparian and wetland vegetation of central and eastern Oregon. Oregon Natural Heritage Information Center, Institute for Natural Resources, Oregon State University, Portland. 473 pp. [http://oregonstate.edu/ornhic/ publications.html]
  • Evens, J. M., K. Sikes, D. Hastings, and J. Ratchford. 2014. Vegetation alliance descriptions for Lake Mead National Recreation Area, Death Valley National Park and Mojave National Preserve. Unpublished report submitted to USDI National Park Service, Mojave Desert Network Inventory and Monitoring Program. California Native Plant Society, Sacramento, CA.
  • Faber-Langendoen, D., editor. 2001. Plant communities of the Midwest: Classification in an ecological context. Association for Biodiversity Information, Arlington, VA. 61 pp. plus appendix (705 pp.).
  • Hansen, P. L., R. D. Pfister, K. Boggs, B. J. Cook, J. Joy, and D. K. Hinckley. 1995. Classification and management of Montana''s riparian and wetland sites. Miscellaneous Publication No. 54. Montana Forest and Conservation Experiment Station, School of Forestry, University of Montana. 646 pp. plus posters.
  • IDCDC [Idaho Conservation Data Center]. 2005. Wetland and riparian plant associations in Idaho. Idaho Conservation Data Center, Idaho Department of Fish and Game, Boise. [http://fishandgame.idaho.gov/tech/CDC/ecology/wetland_riparian_assoc.cfm] (accessed 14 June 2005).
  • Jones, G. 1992b. Wyoming plant community classification (Draft). Wyoming Natural Diversity Database, Laramie, WY. 183 pp.
  • Jones, G. P., and G. M. Walford. 1995. Major riparian vegetation types of eastern Wyoming. Submitted to Wyoming Department of Environmental Quality, Water Quality Division. Wyoming Natural Diversity Database, Laramie, WY. 245 pp.
  • Keeler-Wolf, T., M. Schindel, S. San, P. Moore, and D. Hickson. 2003a. Classification of the vegetation of Yosemite National Park and surrounding environs in Tuolumne, Mariposa, Madera and Mono counties, California. Unpublished report by NatureServe in cooperation with the California Native Plant Society and California Department of Fish and Game, Wildlife and Habitat Data Analysis Branch, Sacramento, CA.
  • Keeler-Wolf, T., P. E. Moore, E. T. Reyes, J. M. Menke, D. N. Johnson, and D. L. Karavidas. 2012. Yosemite National Park vegetation classification and mapping project report. Natural Resource Technical Report NPS/YOSE/NRTR--2012/598. National Park Service, Fort Collins, CO.
  • Kittel, G. M. 1994. Montane vegetation in relation to elevation and geomorphology along the Cache la Poudre River, Colorado. Unpublished thesis, University of Wyoming, Laramie.
  • Kittel, G. M., and N. D. Lederer. 1993. A preliminary classification of the riparian vegetation of the Yampa and San Miguel/Dolores river basins. Unpublished report prepared for Colorado Department of Health and the Environmental Protection Agency by The Nature Conservancy, Colorado Field Office, Boulder.
  • Kittel, G., E. Van Wie, M. Damm, R. Rondeau, S. Kettler, and J. Sanderson. 1999a. A classification of the riparian plant associations of the Rio Grande and Closed Basin watersheds, Colorado. Unpublished report prepared by the Colorado Natural Heritage Program, Colorado State University, Fort Collins.
  • Kittel, G., R. Rondeau, and A. McMullen. 1996. A classification of the riparian vegetation of the Lower South Platte and parts of the Upper Arkansas River basins, Colorado. Submitted to Colorado Department of Natural Resources and the Environmental Protection Agency, Region VIII. Prepared by Colorado Natural Heritage Program, Fort Collins. 243 pp.
  • MTNHP [Montana Natural Heritage Program]. 2002b. List of ecological communities for Montana. Montana Natural Heritage Program, Montana State Library, Helena, MT.
  • Muldavin, E., P. Durkin, M. Bradley, M. Stuever, and P. Mehlhop. 2000a. Handbook of wetland vegetation communities of New Mexico. Volume I: Classification and community descriptions. Final report to the New Mexico Environment Department and the Environmental Protection Agency prepared by the New Mexico Natural Heritage Program, University of New Mexico, Albuquerque.
  • NHNM [Natural Heritage New Mexico]. No date. Unpublished data on file. Natural Heritage New Mexico, University of New Mexico, Albuquerque.
  • Padgett, W. G., A. P. Youngblood, and A. H. Winward. 1988b. Riparian community type classification of Utah. Publication R4-ECOL-88-01. USDA Forest Service, Forest and Range Experiment Station, Ogden, UT.
  • Padgett, W. G., A. P. Youngblood, and A. H. Winward. 1989. Riparian community type classification of Utah and southeastern Idaho. Research Paper R4-ECOL-89-0. USDA Forest Service, Intermountain Forest and Range Experiment Station, Ogden, UT.
  • Peterson, E. B. 2008. International Vegetation Classification alliances and associations occurring in Nevada with proposed additions. Nevada Natural Heritage Program, Carson City, NV. 348 pp.
  • Romme, W. H., K. D. Heil, J. M. Porter, and R. Fleming. 1993. Plant communities of Capitol Reef National Park, Utah. USDI National Park Service, Technical Report NPS/NAUCARE/NRTER-93/02. Cooperative Park Studies Unit, Northern Arizona University. 37 pp.
  • Sawyer, J. O., T. Keeler-Wolf, and J. Evens. 2009. A manual of California vegetation. Second edition. California Native Plant Society, Sacramento CA. 1300 pp.
  • Schulz, K. A., and M. E. Hall. 2011. Vegetation inventory project: Great Basin National Park. Unpublished report submitted to USDI National Park Service, Mojave Desert Inventory and Monitoring Network. NatureServe, Western Regional Office, Boulder, CO. 30 pp. plus Appendices A-H.
  • Steinauer, G., and S. Rolfsmeier. 1997. Terrestrial natural communities of Nebraska. Draft - October 28, 1997. Nebraska Game and Parks Commission, Lincoln, NE. 117 pp.
  • Stevens, J. E., K. Forrest, S. Neid, and M. Fink. 2007. Bent''s Old Fort National Historic Site: Vegetation classification and mapping. Natural Resource Technical Report NPS/SOPN/NRTR--2007/049. National Park Service, Fort Collins, CO. 115 pp.
  • Tendick, A., J. Coles, K. Decker, M. Hall, J. Von Loh, T. Belote, A. Wight, G. Wakefield, and A. Evenden. 2012. Vegetation classification and mapping project report, Canyonlands National Park. Natural Resource Technical Report NPS/NCPN/NRTR--2012/577. National Park Service, Fort Collins, CO.
  • Tendick, A., J. Coles, P. Williams, G. Bradshaw, G. Manis, A. Wight, G. Wakefield, and A. Evenden. 2010. Vegetation classification and mapping project report, Curecanti National Recreation Area. Natural Resource Technical Report NPS/NCPN/NRTR--2010/408. National Park Service, Fort Collins, CO. 797 pp.
  • Tendick, A., J. Spence, M. Reid, K. Shulz, G. Kittel, K. Green, A. Wight, and G. Wakefield. 2017. Vegetation classification and mapping project report, Glen Canyon National Recreation Area and Rainbow Bridge National Monument. Natural Resource Report NPS/NCPN/NRR—2017/1500. National Park Service, Fort Collins, CO. 1464 pp.
  • Von Loh, J., K. Landgraf, A. Evenden, T. Owens, S. Blauer, and M. Reid. 2007. Vegetation classification and mapping project report, Colorado National Monument. Natural Resource Report NPS/NCPN/NRTR--2007/061. National Park Service, Fort Collins, CO. 564 pp.
  • WNDD [Wyoming Natural Diversity Database]. No date. Unpublished data on file. Wyoming Natural Diversity Database, University of Wyoming, Laramie, WY.
  • Walford, G., G. Jones, W. Fertig, S. Mellman-Brown, and K. Houston. 2001. Riparian and wetland plant community types of the Shoshone National Forest. General Technical Report RMRS-GTR-85. USDA Forest Service, Rocky Mountain Research Station, Fort Collins, CO. 122 pp.
  • Western Ecology Working Group of NatureServe. No date. International Ecological Classification Standard: International Vegetation Classification. Terrestrial Vegetation. NatureServe, Boulder, CO.