Print Report

CEGL004103 Carex torta Riverbed Vegetation

Type Concept Sentence: No Data Available


Common (Translated Scientific) Name: Twisted Sedge Riverbed Vegetation

Colloquial Name: Rocky Bar & Shore (Twisted Sedge Type)

Hierarchy Level:  Association

Type Concept: This association accommodates herbaceous alluvial wetlands in which Carex torta is a dominant or characteristic species. It occupies sand, gravel, and rock bars along small rivers and streams in valleys and gorges in the Southern Appalachians, ranging west into the Cumberland Plateau and the Interior Low Plateau, and north into the Central Appalachians, Allegheny Mountains, and Piedmont. This association is characterized by light-requiring, tough-rooted herbaceous perennials tolerant of frequent inundation and flood-scouring. Carex torta often forms dense, extensive colonies. Associated species vary with geography but can include Polygonum sagittatum, Dichanthelium clandestinum, Solidago rugosa ssp. aspera, Juncus effusus, Equisetum arvense, Onoclea sensibilis, Vernonia noveboracensis, Lycopus virginicus, Lobelia cardinalis, Symphyotrichum dumosum, Lycopus virginicus, Osmunda regalis, Hypericum mutilum, Eutrochium fistulosum, Solidago patula, Boehmeria cylindrica, Amphicarpaea bracteata, Acalypha rhomboidea, Impatiens capensis, Leersia oryzoides, and Symphyotrichum lateriflorum. Physiognomy of this type varies from strictly herbaceous to wooded herbaceous to shrubby. Scattered shrubs and small, battered specimens of Platanus occidentalis, Betula nigra, Cornus amomum, Alnus serrulata, and Carpinus caroliniana are present in some stands. An overhanging canopy may include Platanus occidentalis, Betula alleghaniensis var. alleghaniensis, or Acer rubrum. Overhanging shrubs often include Rhododendron maximum. Stands in disturbed landscapes may be heavily invaded by Microstegium vimineum, Polygonum cespitosum var. longisetum, and other weedy exotics.

Diagnostic Characteristics: No Data Available

Rationale for Nominal Species or Physiognomic Features: No Data Available

Classification Comments: No Data Available

Similar NVC Types: No Data Available
note: No Data Available

Physiognomy and Structure: No Data Available

Floristics: Vegetation is characterized by light-demanding, tough-rooted herbaceous perennials tolerant of frequent inundation and flood-scouring. Carex torta is usually the dominant species and typically forms dense, extensive colonies. Associated species vary with geography. Polygonum sagittatum, Solidago rugosa, and Dichanthelium clandestinum are typical; others include Acalypha rhomboidea, Amphicarpaea bracteata, Boehmeria cylindrica, Equisetum arvense, Eutrochium fistulosum (= Eupatorium fistulosum), Euthamia graminifolia var. graminifolia, Juncus effusus, Hypericum mutilum, Hypericum perforatum, Impatiens capensis, Leersia oryzoides, Lobelia cardinalis, Lycopus virginicus, Onoclea sensibilis, Osmunda regalis, Prunella vulgaris, Solidago patula, Symphyotrichum dumosum (= Aster dumosus), Symphyotrichum lateriflorum (= Aster lateriflorus), Symphyotrichum prenanthoides, Thalictrum spp., Trautvetteria caroliniensis var. caroliniensis, Verbesina alternifolia, and Vernonia noveboracensis. Cover by nonvascular plants is insignificant. Physiognomy of this type varies from strictly herbaceous to wooded herbaceous to shrubby. Scattered shrubs and small, battered specimens of Platanus occidentalis, Betula nigra, Cornus amomum, Alnus serrulata, and Carpinus caroliniana are present in some stands. An overhanging canopy with an average cover of 20% may include Platanus occidentalis, Betula alleghaniensis var. alleghaniensis, or Acer rubrum. Sparse overhanging shrubs often include Rhododendron maximum. Stands in disturbed landscapes may be heavily invaded by Microstegium vimineum, Polygonum cespitosum var. longisetum, and other weedy exotics. Mean species richness of vascular plants in six West Virginia plots was 36 taxa per 400 m2, with most of the diversity in the herbaceous stratum.

Dynamics:  Periodic large or severe floods transport and deposit large numbers of cobbles, stones, and even boulders in characteristic bars both within the channel (islands) and along its edges (streambanks). These bars provide a matrix for deposition of finer alluvium and habitats for the establishment of tough, adaptable herbaceous plants, which in turn stabilize the bars with massive networks of perennial rootstocks. Regular flood-scouring batters or removes woody plants which take root in these habitats, maintaining open-canopy conditions. These habitats are highly dynamic, with conditions more or less constantly shifting in response to an irregular but powerful disturbance regime. While some of the bar habitats may be damaged or removed by severe floods, others may accrete or be newly deposited during the same events (Hupp 1982). Successionally, this unit can be considered a pioneering type on new, coarse alluvial land, but it is also more or less permanently maintained by natural disturbances.

Environmental Description:  This association occupies boulder and gravel bars on the frequently flooded, active channel shelves of high-gradient streams and small rivers, often forming small, discontinuous, linear patches. It is subject to frequent, high-energy inundation and flood-scouring, which can occur at any time of year. Flooding duration is probably similar to that documented along Passage Creek in Shenandoah County, Virginia, by Hupp (1982). In that drainage, the channel shelf was inundated approximately 15% of the time. Elevations in West Virginia range from 390-1200 m. Soils are moderately to poorly drained sand with pH averaging 6.5 (n=5). Organic soils are not present. Soil chemistry is characterized by high Cu, Mn; moderate B, Ca, Mg; and low Al, ENR, Fe, H, K, Na, P, S, TEC, Zn, and organic matter (n=3).

Geographic Range: This community is found as small-patch linear occurrences along waterways in the Southern Appalachians, west into the Cumberland Plateau and the Interior Low Plateau, and north into the Central Appalachians, Allegheny Mountains, and Piedmont.

Nations: US

States/Provinces:  AL, DE, GA, KY, MD, NC, PA, SC, TN, VA, WV




Confidence Level: Moderate

Confidence Level Comments: No Data Available

Grank: G3G4

Greasons: No Data Available


Concept Lineage: No Data Available

Predecessors: No Data Available

Obsolete Names: No Data Available

Obsolete Parents: No Data Available

Synonomy: ? Carex torta Association (Fleming and Moorhead 1996)
= Carex torta Herbaceous Vegetation (Fleming and Taverna 2006)
= Carex torta Herbaceous Vegetation (Fleming and Coulling 2001)
= Carex torta Herbaceous Vegetation (Bowman 2000)
= Carex torta Riverscour Prairie (Byers et al. 2007)
= Carex torta riparian herbaceous vegetation (Vanderhorst 2001b)
< IIE3a. Riverside Shoal and Stream Bar Complex (Allard 1990)
= Sedge - spotted joe pye weed riverine herbaceous vegetation (Perles et al. 2004)
? Torturous sedge gravel rivershore (CAP pers. comm. 1998)

Concept Author(s): G. Fleming and P. Coulling (2001)

Author of Description: G. Fleming, P. Coulling, S.C. Gawler

Acknowledgements: No Data Available

Version Date: 12-18-06

  • Allard, D. J. 1990. Southeastern United States ecological community classification. Interim report, Version 1.2. The Nature Conservancy, Southeast Regional Office, Chapel Hill, NC. 96 pp.
  • Bowman, P. 2000. Draft classification for Delaware. Unpublished draft. Delaware Natural Heritage Program, Smyrna, DE.
  • Byers, E. A., J. P. Vanderhorst, and B. P. Streets. 2007. Classification and conservation assessment of high elevation wetland communities in the Allegheny Mountains of West Virginia. West Virginia Natural Heritage Program, West Virginia Division of Natural Resources, Elkins.
  • CAP [Central Appalachian Forest Working Group]. 1998. Central Appalachian Working group discussions. The Nature Conservancy, Boston, MA.
  • Coxe, R. 2009. Guide to Delaware vegetation communities. Spring 2009 edition. State of Delaware, Division of Fish and Wildlife, Delaware Natural Heritage Program, Smyrna.
  • Eyre, F. H., editor. 1980. Forest cover types of the United States and Canada. Society of American Foresters, Washington, DC. 148 pp.
  • Fleming, G. P., K. D. Patterson, and K. Taverna. 2017. The natural communities of Virginia: A classification of ecological community groups and community types. Third approximation. Version 3.0. Virginia Department of Conservation and Recreation, Division of Natural Heritage, Richmond, VA. [http://www.dcr.virginia.gov/natural-heritage/natural-communities/]
  • Fleming, G. P., and K. Taverna. 2006. Vegetation classification for the National Capitol Region parks, western region. Regional (VA-WVA-MD-DC) analysis prepared for NatureServe and USGS-NPS Vegetation Mapping Program, March 2006. Virginia Department of Conservation and Recreation, Division of Natural Heritage, Richmond.
  • Fleming, G. P., and P. P. Coulling. 2001. Ecological communities of the George Washington and Jefferson national forests, Virginia. Preliminary classification and description of vegetation types. Virginia Department of Conservation and Recreation, Division of Natural Heritage, Richmond, VA. 317 pp.
  • Fleming, G. P., and W. H. Moorhead, III. 1996. Ecological land units of the Laurel Fork Area, Highland County, Virginia. Natural Heritage Technical Report 96-08. Virginia Department of Conservation and Recreation, Division of Natural Heritage, Richmond. 114 pp. plus appendices.
  • Harrison, J. W., compiler. 2004. Classification of vegetation communities of Maryland: First iteration. A subset of the International Classification of Ecological Communities: Terrestrial Vegetation of the United States, NatureServe. Maryland Natural Heritage Program, Maryland Department of Natural Resources, Annapolis. 243 pp.
  • Hupp, C. R. 1982. Stream-grade variation and riparian forest ecology along Passage Creek, Virginia. Bulletin of the Torrey Botanical Club 109:488-499.
  • NatureServe Ecology - Southeastern United States. No date. Unpublished data. NatureServe, Durham, NC.
  • Nelson, J. B. 1986. The natural communities of South Carolina: Initial classification and description. South Carolina Wildlife and Marine Resources Department, Division of Wildlife and Freshwater Fisheries, Columbia, SC. 55 pp.
  • PNHP [Pennsylvania Natural Heritage Program]. 2004. Classification, assessment and protection of non-forested floodplain wetlands of the Susquehanna Drainage. U.S. EPA Wetlands Protection State Development Grant no. CD-98337501. Report to the U.S. Environmental Protection Agency and the Pennsylvania Department of Conservation and Natural Resources, Bureau of Forestry, Ecological Services Section, Harrisburg, PA.
  • Palmer-Ball, B., Jr., J. J. N. Campbell, M. E. Medley, D. T. Towles, J. R. MacGregor, and R. R. Cicerello. 1988. Cooperative inventory of endangered, threatened, sensitive and rare species, Daniel Boone National Forest, Somerset Ranger District. USDA Forest Service, Daniel Boone National Forest, Berea, KY. 244 pp.
  • Peet, R. K., T. R. Wentworth, M. P. Schafale, and A.S. Weakley. No date. Unpublished data of the North Carolina Vegetation Survey. University of North Carolina, Chapel Hill.
  • Perles, S., G. Podniesinski, and J. Wagner. 2004. Classification, assessment and protection of non-forested floodplain wetlands of the Susquehanna drainage. Report to the U.S. Environmental Protection Agency and Pennsylvania Department of Conservation and Natural Resources. Pennsylvania Natural Heritage Program, Harrisburg. 128 pp.
  • Schafale, M. P. 2012. Classification of the natural communities of North Carolina, 4th Approximation. North Carolina Department of Environment, Health, and Natural Resources, Division of Parks and Recreation, Natural Heritage Program, Raleigh.
  • Schafale, M. P., and A. S. Weakley. 1990. Classification of the natural communities of North Carolina. Third approximation. North Carolina Department of Environment, Health, and Natural Resources, Division of Parks and Recreation, Natural Heritage Program, Raleigh. 325 pp.
  • Schafale, Mike P. Personal communication. Ecologist, North Carolina Department of Environment, Health, and Natural Resources, Division of Parks and Recreation, Natural Heritage Program, Raleigh.
  • Southeastern Ecology Working Group of NatureServe. No date. International Ecological Classification Standard: International Vegetation Classification. Terrestrial Vegetation. NatureServe, Durham, NC.
  • Vanderhorst, J. 2001b. Plant communities of the New River Gorge National River, West Virginia: Northern and southern thirds. Non-game Wildlife and Natural Heritage Program, West Virginia Division of Natural Resources. Elkins. 146 pp.
  • Vanderhorst, J. P., J. Jeuck, and S. C. Gawler. 2007. Vegetation classification and mapping of New River Gorge National River, West Virginia. Technical Report NPS/NER/NRTR-2007/092. National Park Service, Philadelphia, PA. 396 pp.
  • WVNHP [West Virginia Natural Heritage Program]. No date. Unpublished data. West Virginia Natural Heritage Program, Elkins.
  • Zimmerman, E. A. 2011p. Pennsylvania Natural Heritage Program. Twisted Sedge (Carex torta) Stream Margin Factsheet. [http://www.naturalheritage.state.pa.us/Community.aspx?=30021] (accessed February 16, 2012)
  • Zimmerman, E. A., T. Davis, M. A. Furedi, B. Eichelberger, J. McPherson, S. Seymour, G. Podniesinski, N. Dewar, and J. Wagner, editors. 2012. Terrestrial and palustrine plant communities of Pennsylvania. Pennsylvania Natural Heritage Program, Pennsylvania Department of Conservation and Natural Resources, Harrisburg. [http://www.naturalheritage.state.pa.us/Communities.aspx]
  • Zimmerman, E., and G. Podniesinski. 2008. Classification, assessment and protection of floodplain wetlands of the Ohio Drainage. U.S. EPA Wetlands Protection State Development Grant no. CD-973081-01-0. Report submitted to the U.S. Environmental Protection Agency and the Pennsylvania Department of Conservation and Natural Resources, Office of Conservation Science. Pennsylvania Natural Heritage Program, Western Pennsylvania Conservancy, Pittsburgh, PA.