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CEGL004286 Justicia americana Riverbed Vegetation
Type Concept Sentence: No Data Available
Common (Translated Scientific) Name: American Water-willow Riverbed Vegetation
Colloquial Name: Water-willow Rocky Bar & Shore
Hierarchy Level: Association
Type Concept: This widespread association is found primarily in the Piedmont, Central Appalachians, Cumberland Plateau, Interior Low Plateau, Ozarks, Ouachita Mountains, and adjacent provinces. Stands occur on the shoals or bars of rocky streams and riverbeds, where they are subject to frequent high-energy floods. Slopes range from level to moderate but are typically gentle. The substrate is a variable mixture of sand, gravel and cobbles, often with deposits of silt and muck. Rooting occurs in the interstices between cobbles and boulders. Justicia americana is the characteristic dominant. Saururus cernuus is often present and may be codominant. Other herbaceous species that may be present include Bidens spp., Cyperus spp., Diodia teres, Elodea sp., Eleocharis spp., Equisetum arvense, Gratiola brevifolia, Leersia oryzoides, Leersia virginica, Lemna minor, Orontium aquaticum, Physostegia virginiana, Podostemum ceratophyllum, Scirpus spp., Schoenoplectus pungens, Schoenoplectus tabernaemontani, and Xyris difformis var. difformis. Exotics include Lythrum salicaria, Lysimachia vulgaris and Polygonum cespitosum var. longisetum; where present, these are sparse. Some stands have low cover by scattered flood-suppressed trees or an overhanging canopy. Trees in plots include Acer saccharinum, Betula nigra, Carpinus caroliniana, Fraxinus pennsylvanica, and Platanus occidentalis. Scattered shrub seedlings of Salix nigra, Salix caroliniana, Betula nigra, Cephalanthus occidentalis, Acer saccharinum, or Platanus occidentalis may also be present. This association provides habitat in some portions of its range for globally rare dragonflies and herbs.
Diagnostic Characteristics: No Data Available
Rationale for Nominal Species or Physiognomic Features: No Data Available
Classification Comments: In Ohio this type often forms pure patches, but consistent identification may require a simple cutoff rule, such as at least 50% cover of Justicia (Anderson 1982). However, Anderson (1996) no longer recognizes this type.
Similar NVC Types: No Data Available
note: No Data Available
Physiognomy and Structure: No Data Available
Floristics: Justicia americana is the dominant (and sometimes the only) species, forming lawnlike stands in shallow reaches of rivers. Cover by Justicia americana ranges from 40 to 85%. Saururus cernuus is often present and may be codominant. Other herbaceous species may be present but rarely achieve more than 2% cover; they include Bidens spp., Cyperus spp., Diodia teres, Elodea sp., Eleocharis spp., Equisetum arvense, Gratiola brevifolia, Leersia oryzoides, Leersia virginica, Lemna minor, Orontium aquaticum, Physostegia virginiana, Podostemum ceratophyllum, Scirpus spp., Schoenoplectus pungens, Schoenoplectus tabernaemontani, and Xyris difformis var. difformis. Exotics include Lythrum salicaria, Lysimachia vulgaris, and Polygonum cespitosum var. longisetum; they have not been observed in abundance in this community, possibly due to intolerance of high-energy flooding. In some areas, Justicia usually grows in nearly pure patches, so that few other species are associated with it. Cuscuta gronovii, Mimulus ringens, Polygonum spp., Rumex spp., and Salix interior can also occur (Anderson 1982). Some stands have low cover by scattered flood-suppressed trees or an overhanging canopy. Trees in plots include Acer saccharinum, Betula nigra, Carpinus caroliniana, Fraxinus pennsylvanica, and Platanus occidentalis. Scattered shrub seedlings of Salix nigra, Salix caroliniana, Betula nigra, Cephalanthus occidentalis, Acer saccharinum, or Platanus occidentalis may also be present. In the Cumberland Plateau of Alabama, Justicia americana is present in dense patches with some interspersion of other species, including Pilea pumila, Boehmeria cylindrica, Eclipta prostrata (= Eclipta alba), Juncus coriaceus, Mikania scandens, Ludwigia palustris, Leersia sp., and Bidens sp. Schmalzer and DeSelm (1982) discuss Orontium aquaticum growing along streambanks or in shallow riffles "along or with" Justicia americana in the Obed River in the Cumberland Plateau of Tennessee. In 29 plots sampled in the Potomac River watershed (Piedmont, Blue Ridge and Ridge and Valley provinces), Justicia was overwhelmingly dominant (50-75% mean cover), and no associated species occurred in more than 48% of the plots. Vascular plant species richness in sampled plots is low (typically 6-14 taxa).
Dynamics: Stands in some situations may be obliterated by ongoing river channeling. Anderson (1982) describes some of the life-history characteristics of Justicia americana that allow it to persist in river channels.
Environmental Description: This association occurs on the shoals or bars of rocky streams and riverbeds, on bedrock, boulders, cobble, gravel, and sands. They are subject to frequent high-energy floods, and are entirely submerged by most flood events. During extreme low water periods, the substrate can be exposed, showing a varied mixture of sand, gravel and cobbles, often with deposits of silt and muck. Rooting occurs in the interstices between cobbles and boulders. Stands commonly occur on the edge of the river and at the heads and tails of islands and may sometimes occur on deposition bars in the middle of the river. Substrate pH is circumneutral (mean = 6.0) in four samples. Slopes range from level to moderate but are typically gentle. Elevations of West Virginia stands range from 73 to at least 654 m; the type occurs at lower elevations in the Piedmont and Coastal Plain.
Geographic Range: This type is found primarily in the Piedmont, Interior Low Plateau, Cumberland Plateau, Ozarks, Ouachita Mountains, Central Appalachians, and adjacent provinces. It ranges from Alabama, Georgia and the Carolinas west to Arkansas and Oklahoma and north to Ohio, New York, and New Jersey, with possible outliers north to southern Quebec.
Nations: US
States/Provinces: AL, AR, GA, KY, MD, NC, NJ, NY, OH, OK, PA, SC?, TN, VA, WV
Plot Analysis Summary:
http://vegbank.org/natureserve/ELEMENT_GLOBAL.2.685634
Confidence Level: High
Confidence Level Comments: No Data Available
Grank: G4G5
Greasons: No Data Available
Type | Name | Database Code | Classification Code |
---|---|---|---|
Class | 2 Shrub & Herb Vegetation Class | C02 | 2 |
Subclass | 2.C Shrub & Herb Wetland Subclass | S44 | 2.C |
Formation | 2.C.4 Temperate to Polar Freshwater Marsh, Wet Meadow & Shrubland Formation | F013 | 2.C.4 |
Division | 2.C.4.Nd Eastern North American Temperate Freshwater Marsh, Wet Meadow & Shrubland Division | D323 | 2.C.4.Nd |
Macrogroup | 2.C.4.Nd.3 Eastern North American Wet Shoreline Vegetation Macrogroup | M880 | 2.C.4.Nd.3 |
Group | 2.C.4.Nd.3.a Eastern North American Riverine Wetland Vegetation Group | G755 | 2.C.4.Nd.3.a |
Alliance | A1657 American Water-willow Riverbed Alliance | A1657 | 2.C.4.Nd.3.a |
Association | CEGL004286 American Water-willow Riverbed Vegetation | CEGL004286 | 2.C.4.Nd.3.a |
Concept Lineage: No Data Available
Predecessors: No Data Available
Obsolete Names: No Data Available
Obsolete Parents: No Data Available
Synonomy: = Justicia americana - Andropogon gerardii herbaceous channel bed (Walton and Anderson 1997)
= Justicia americana Herbaceous Vegetation (Faber-Langendoen 2001)
= Justicia americana riparian herbaceous vegetation (Vanderhorst 2001b)
< Aquatic Types (Schmalzer and DeSelm 1982)
< IIE3a. Riverside Shoal and Stream Bar Complex (Allard 1990)
? Lizard''s tail emergent bed (Perles et al. 2004)
? Water willow emergent bed (Perles et al. 2004)
= Justicia americana Herbaceous Vegetation (Faber-Langendoen 2001)
= Justicia americana riparian herbaceous vegetation (Vanderhorst 2001b)
< Aquatic Types (Schmalzer and DeSelm 1982)
< IIE3a. Riverside Shoal and Stream Bar Complex (Allard 1990)
? Lizard''s tail emergent bed (Perles et al. 2004)
? Water willow emergent bed (Perles et al. 2004)
- Allard, D. J. 1990. Southeastern United States ecological community classification. Interim report, Version 1.2. The Nature Conservancy, Southeast Regional Office, Chapel Hill, NC. 96 pp.
- Anderson, D. M. 1982. Plant communities of Ohio: A preliminary classification and description. Division of Natural Areas and Preserves, Ohio Department of Natural Resources, Columbus, OH. 182 pp.
- Anderson, D. M. 1996. The vegetation of Ohio: Two centuries of change. Draft. Ohio Biological Survey.
- Edinger, G. J., D. J. Evans, S. Gebauer, T. G. Howard, D. M. Hunt, and A. M. Olivero, editors. 2014a. Ecological communities of New York state. Second edition. A revised and expanded edition of Carol Reschke''s ecological communities of New York state. New York Natural Heritage Program, New York State Department of Environmental Conservation, Albany, NY.
- Evans, M., B. Yahn, and M. Hines. 2009. Natural communities of Kentucky 2009. Kentucky Nature Preserves Commission, Frankfort, KY. 22 pp.
- Faber-Langendoen, D., editor. 2001. Plant communities of the Midwest: Classification in an ecological context. Association for Biodiversity Information, Arlington, VA. 61 pp. plus appendix (705 pp.).
- Fike, J. 1999. Terrestrial and palustrine plant communities of Pennsylvania. Pennsylvania Natural Diversity Inventory. Pennsylvania Department of Conservation and Recreation, Bureau of Forestry, Harrisburg, PA. 86 pp.
- Fleming, G. P. 2007. Ecological communities of the Potomac Gorge in Virginia: Composition, floristics, and environmental dynamics. Natural Heritage Technical Report 07-12. Unpublished report submitted to the National Park Service. Virginia Department of Conservation and Recreation, Division of Natural Heritage, Richmond. 341 pp. plus appendices.
- Fleming, G. P., K. D. Patterson, and K. Taverna. 2017. The natural communities of Virginia: A classification of ecological community groups and community types. Third approximation. Version 3.0. Virginia Department of Conservation and Recreation, Division of Natural Heritage, Richmond, VA. [http://www.dcr.virginia.gov/natural-heritage/natural-communities/]
- Fleming, G. P., and K. Taverna. 2006. Vegetation classification for the National Capitol Region parks, western region. Regional (VA-WVA-MD-DC) analysis prepared for NatureServe and USGS-NPS Vegetation Mapping Program, March 2006. Virginia Department of Conservation and Recreation, Division of Natural Heritage, Richmond.
- Harrison, J. W. 2011. The natural communities of Maryland: 2011 working list of ecological community groups and community types. Unpublished report. Maryland Department of Natural Resources, Wildlife and Heritage Service, Natural Heritage Program, Annapolis. 33 pp.
- Harrison, J. W., compiler. 2004. Classification of vegetation communities of Maryland: First iteration. A subset of the International Classification of Ecological Communities: Terrestrial Vegetation of the United States, NatureServe. Maryland Natural Heritage Program, Maryland Department of Natural Resources, Annapolis. 243 pp.
- Hoagland, B. 2000. The vegetation of Oklahoma: A classification for landscape mapping and conservation planning. The Southwestern Naturalist 45(4):385-420.
- Hop, K., M. Pyne, T. Foti, S. Lubinski, R. White, and J. Dieck. 2012a. National Park Service vegetation inventory program: Buffalo National River, Arkansas. Natural Resource Report NPS/HTLN/NRR--2012/526. National Park Service, Fort Collins, CO. 340 pp.
- Lea, C. 2000. Plant communities of the Potomac Gorge and their relationship to fluvial factors. M.S. thesis, George Mason University. Fairfax, VA. 219 pp.
- Lea, C. 2004. Draft vegetation types in National Capital Region Parks. Edited by S.C. Gawler and J. Teague. Working draft for review by NatureServe, Virginia Natural Heritage, West Virginia Natural Heritage, Maryland Natural Heritage, and National Park Service. July 2004. 157 pp.
- Major, C. S., C. Bailey, J. Donaldson, R. McCoy, C. Nordman, M. Williams, and D. Withers. 1999. An ecological inventory of selected sites in the Cherokee National Forest. Cost Share Agreement #99-CCS-0804-001. Tennessee Department of Environment and Conservation, Tennessee Division of Natural Heritage, Nashville, TN.
- McCoy, D. A. 1958. Vascular plants of Pontotoc County, Oklahoma. American Midland Naturalist 59:371-396.
- Nelson, J. B. 1986. The natural communities of South Carolina: Initial classification and description. South Carolina Wildlife and Marine Resources Department, Division of Wildlife and Freshwater Fisheries, Columbia, SC. 55 pp.
- Nordman, C. 2004a. Vascular plant community classification for Stones River National Battlefield. NatureServe, Durham, NC. 64 pp. plus appendices and CD.
- Nordman, C., M. Russo, and L. Smart. 2011. Vegetation types of the Natchez Trace Parkway, based on the U.S. National Vegetation Classification. NatureServe Central Databases (International Ecological Classification Standard: Terrestrial Ecological Classifications). Arlington, VA. Data current as of 11 April 2011. 548 pp.
- ONHD [Ohio Natural Heritage Database]. No date. Vegetation classification of Ohio and unpublished data. Ohio Natural Heritage Database, Division of Natural Areas and Preserves, Ohio Department of Natural Resources, Columbus.
- Palmer-Ball, B., Jr., J. J. N. Campbell, M. E. Medley, D. T. Towles, J. R. MacGregor, and R. R. Cicerello. 1988. Cooperative inventory of endangered, threatened, sensitive and rare species, Daniel Boone National Forest, Somerset Ranger District. USDA Forest Service, Daniel Boone National Forest, Berea, KY. 244 pp.
- Peet, R. K., T. R. Wentworth, M. P. Schafale, and A.S. Weakley. No date. Unpublished data of the North Carolina Vegetation Survey. University of North Carolina, Chapel Hill.
- Penfound, W. T. 1953. Plant communities of Oklahoma lakes. Ecology 34:561-583.
- Perles, S. J., G. S. Podniesinski, E. Eastman, L. A. Sneddon, and S. C. Gawler. 2007. Classification and mapping of vegetation and fire fuel models at Delaware Water Gap National Recreation Area. Technical Report NPS/NER/NRTR--2007/076. National Park Service, Philadelphia, PA. 2 volumes.
- Perles, S. J., G. S. Podniesinski, M. Furedi, B. A. Eichelberger, A. Feldmann, G. Edinger, E. Eastman, and L. A. Sneddon. 2008. Vegetation classification and mapping at Upper Delaware Scenic and Recreational River. Technical Report NPS/NER/NRTR--2008/133. National Park Service, Philadelphia, PA. 370 pp.
- Perles, S., G. Podniesinski, and J. Wagner. 2004. Classification, assessment and protection of non-forested floodplain wetlands of the Susquehanna drainage. Report to the U.S. Environmental Protection Agency and Pennsylvania Department of Conservation and Natural Resources. Pennsylvania Natural Heritage Program, Harrisburg. 128 pp.
- Pyne, M., E. Lunsford Jones, and R. White. 2010. Vascular plant inventory and plant community classification for Mammoth Cave National Park. NatureServe, Durham, NC. 334 pp.
- Salas, D. E., T. Folts-Zettner, R. W. Sanders, and J. Drake. 2010c. Vegetation classification and mapping at Chickasaw National Recreation Area. Natural Resource Technical Report NPS/SOPN/NRTR--2010/286. National Park Service, Fort Collins, CO. 176 pp.
- Schafale, M. P. 2012. Classification of the natural communities of North Carolina, 4th Approximation. North Carolina Department of Environment, Health, and Natural Resources, Division of Parks and Recreation, Natural Heritage Program, Raleigh.
- Schafale, M. P., and A. S. Weakley. 1990. Classification of the natural communities of North Carolina. Third approximation. North Carolina Department of Environment, Health, and Natural Resources, Division of Parks and Recreation, Natural Heritage Program, Raleigh. 325 pp.
- Schmalzer, P. A., and H. R. DeSelm. 1982. Vegetation, endangered and threatened plants, critical plant habitats and vascular flora of the Obed Wild and Scenic River. Unpublished report. USDI National Park Service, Obed Wild and Scenic River. 2 volumes. 369 pp.
- Schotz, A., H. Summer, and R. White, Jr. 2008. Vascular plant inventory and ecological community classification for Little River Canyon National Preserve. NatureServe, Durham, NC. 244 pp.
- Southeastern Ecology Working Group of NatureServe. No date. International Ecological Classification Standard: International Vegetation Classification. Terrestrial Vegetation. NatureServe, Durham, NC.
- TNC and WPC [The Nature Conservancy and Western Pennsylvania Conservancy]. 2004. Classification, assessment, and protection of non-forested floodplain wetlands of the Susquehanna drainage. Pennsylvania Natural Heritage Program, Harrisburg, PA. 128 pp.
- Vanderhorst, J. 2001b. Plant communities of the New River Gorge National River, West Virginia: Northern and southern thirds. Non-game Wildlife and Natural Heritage Program, West Virginia Division of Natural Resources. Elkins. 146 pp.
- Vanderhorst, J. P., B. P. Streets, Z. Arcaro, and S. C. Gawler. 2010. Vegetation classification and mapping at Gauley River National Recreation Area. Technical Report NPS/NER/NRTR--2010/148. National Park Service, Philadelphia, PA.
- Vanderhorst, J. P., J. Jeuck, and S. C. Gawler. 2007. Vegetation classification and mapping of New River Gorge National River, West Virginia. Technical Report NPS/NER/NRTR-2007/092. National Park Service, Philadelphia, PA. 396 pp.
- WVNHP [West Virginia Natural Heritage Program]. No date. Unpublished data. West Virginia Natural Heritage Program, Elkins.
- Walton, D., and M. Anderson. 1997. Critical habitats and associated communities in the riparian zone of the Gauley River. West Virginia Division of Natural Resources, Natural Heritage Program, Elkins.
- Zimmerman, E. A. 2011r. Pennsylvania Natural Heritage Program. Water-willow (Justicia americana) - Smartweed Riverbed Community Factsheet. [http://www.naturalheritage.state.pa.us/Community.aspx?=16017] (accessed February 13, 2012)
- Zimmerman, E. A., T. Davis, M. A. Furedi, B. Eichelberger, J. McPherson, S. Seymour, G. Podniesinski, N. Dewar, and J. Wagner, editors. 2012. Terrestrial and palustrine plant communities of Pennsylvania. Pennsylvania Natural Heritage Program, Pennsylvania Department of Conservation and Natural Resources, Harrisburg. [http://www.naturalheritage.state.pa.us/Communities.aspx]