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CEGL001156 Alnus viridis ssp. sinuata / Athyrium filix-femina - Cinna latifolia Wet Shrubland
Type Concept Sentence: No Data Available
Common (Translated Scientific) Name: Sitka Alder / Common Ladyfern - Drooping Woodreed Wet Shrubland
Colloquial Name: No Data Available
Hierarchy Level: Association
Type Concept: This association is found in cool and moist mountainous regions between about 1022-1837 m (3350-6012 feet) elevation, ranging from northern California, along the eastern slope of the Cascades to southern British Columbia, through northeastern Oregon, Washington and Montana and north into the Canadian Rockies. It usually occurs as narrow stringers in moderate to steep, V-shaped valleys in areas of deep or long-lasting snowpacks. Such habitats include floodplains and streambanks of small streams (orders 1 and 2), avalanche chutes, and occasionally springs. These areas often flood during snowmelt and remain wet throughout the summer. Soils vary, but are typically thin silt or sandy loams over alluvial cobble and gravel. Alnus viridis ssp. sinuata forms dense, 3- to 5-m tall thickets with 60 to nearly 100% cover, but less dense stands are also known. Conifers, especially Abies grandis and Picea engelmannii, are sometimes present and may indicate a successional trend toward conifer-dominated associations. Periodic severe flood or avalanche disturbance may be necessary for maintaining the long-term dominance of Alnus viridis ssp. sinuata. The only understory shrubs with greater than 50% constancy (but usually low cover), are Ribes spp. (Ribes hudsonianum or Ribes lacustre), Salix drummondiana, and Rubus parviflorus. Athyrium filix-femina, 30-90 cm tall, is always present in the understory, typically with 20-80% cover, while another fern, Gymnocarpium dryopteris, is sometimes subdominant. Cinna latifolia is often present, but averages only 5% cover (and less than Athyrium filix-femina). Tall forbs, most commonly Maianthemum stellatum, Senecio triangularis, Chamerion angustifolium, Prosartes spp., and Streptopus amplexifolius, have high constancy but usually have less than 10% cover, although they can occasionally be quite abundant, with as much as 50% canopy cover. A lush ground layer composed of species including, but not limited to, Boykinia major, Circaea alpina, Claytonia cordifolia, Galium triflorum, and Mitella spp., is often present beneath the taller Athyrium filix-femina canopy.
Diagnostic Characteristics: No Data Available
Rationale for Nominal Species or Physiognomic Features: No Data Available
Classification Comments: This association is defined as a PNV vegetation type. If it were renamed as a dominance type, the species would include Alnus viridis ssp. sinuata and Ribes hudsonianum (RIHU). This association is a well-documented type known from 64 quantitative plots: 14 plots sampled in northeastern Oregon (Crowe and Clausnitzer 1997, Crowe et al. 2002), approximately 38 plots from eastern Washington (Wooten and Morrison 1995, Kovalchik 2001, WNHP 2002), and 12 plots in central and northern Idaho (IDCDC 2002). Several plots included in the more general Alnus viridis ssp. sinuata community type sampled by Hansen et al. (1995) in northwestern Montana are likely synonymous with this association. Stands are known from Glacier National Park, Montana, but the total number of occurrences is not known (MTNHP 2002a). Cinna latifolia is not present in this community in Montana and rarely has over 10% cover in stands throughout its range. In stands with sparsely vegetated understories, Athyrium filix-femina can have low cover, but it remains the dominant understory species. The following three associations are similar but have yet to be incorporated into the National Vegetation Classification. The related, but distinct Alnus viridis ssp. sinuata / Cinna latifolia (Crowe and Clausnitzer 1997, Crowe et al. 2002) and Alnus viridis ssp. sinuata / Gymnocarpium dryopteris (Kovalchik 2001), which are distinguished by having 100% constancy and moderate cover of their respective diagnostic understory species, but only trace to low cover of Athyrium filix-femina. The Alnus viridis ssp. sinuata / Rubus spectabilis / Athyrium filix-femina association (Diaz and Mellon 1996, Murray 2000, Kovalchik 2001) apparently ranges from the eastern slope of the Cascades west to the Pacific coast and is distinguished by having moderate to high cover and constancy of Rubus spectabilis. If Oplopanax horridus is present with moderate cover, the stand is classified as ~Alnus viridis ssp. sinuata / Oplopanax horridus Shrub Swamp (CEGL001157)$$ (Kovalchik 2001). [SVC sees very little, if any, environmental distinction between ALNVIR/ATHFIL and ALNVIR/MESIC and only the slightest compositional distinction, mostly ATHFIL present with modest cover or not; this may be the case for trying to adopt a type defined elsewhere to local circumstances; recognizing both types does no real "harm," but it is probably an inconsequential distinction.]
Similar NVC Types: No Data Available
note: No Data Available
Physiognomy and Structure: No Data Available
Floristics: Alnus viridis ssp. sinuata forms dense, 3- to 5-m tall, thickets with 60% to nearly 100% cover, but less dense stands are also known. Conifers, especially Abies grandis and Picea engelmannii, are sometimes present and may indicate a successional trend toward conifer-dominated associations. Periodic severe flood or avalanche disturbance may be necessary for maintaining the long-term dominance of Alnus viridis ssp. sinuata. The only understory shrubs with greater than 50% constancy (but usually low cover), are Ribes spp. (Ribes hudsonianum or Ribes lacustre) and Rubus parviflorus. Athyrium filix-femina, 30-90 cm tall, is always present in the understory, typically with 20-80% cover, while another fern, Gymnocarpium dryopteris, is sometimes subdominant. Cinna latifolia is often present, but averages only 5% cover (and less than Athyrium filix-femina). Tall forbs, most commonly Maianthemum stellatum, Senecio triangularis, and Streptopus amplexifolius, have high constancy but usually have less than 10% cover each. A lush forb ground layer composed of species including, but not limited to, Boykinia major, Circaea alpina, Claytonia cordifolia, Galium triflorum, and Mitella spp., is often present beneath the taller Athyrium filix-femina canopy.
Dynamics: Alnus viridis ssp. sinuata is considered an early-seral species able to quickly colonize bare mineral soil. It has the ability to resprout after fire, flood, avalanche, ice, or other disturbances to above-ground stems (Kovalchik 2001). Repeated broadcast burning in the Thuja plicata - Tsuga heterophylla zone (such as after logging operations) can increase the frequency and cover of Alnus viridis ssp. sinuata (Hansen et al. 1995), although severe fires may kill root crowns (Crowe and Clausnitzer 1997, Kovalchik 2001). Natural fires are rare to occasional in cool and moist valley bottoms occupied by this type. While it is possible that logging and subsequent slash burning may promote this association in some areas, logging operations may also lead to site desiccation through alteration of the hydrology, soil compaction, and erosion leading to the loss of mesic species such as Athyrium filix-femina (Hansen et al. 1995, Crowe and Clausnitzer 1997).
Environmental Description: This association is apparently limited to moist, mid- to high-elevation mountainous areas with deep or long-lasting snowpacks between about 1022-1837 m (3350-6012 feet) elevation (Crowe and Clausnitzer 1997, Crowe et al. 2002). In these areas the association most frequently develops as narrow stringers in moderate to steep, V-shaped valleys along small streams (orders 1 and 2) and avalanche chutes. Although sediments must remain stable and moist enough to support initial establishment of both Alnus viridis ssp. sinuata and Athyrium filix-femina (Crowe and Clausnitzer 1997), the association may need periodic severe flood or avalanche disturbance to reduce tree invasion and maintain Alnus viridis ssp. sinuata dominance (Hansen et al. 1995, MTNHP 2002a). Alnus viridis ssp. sinuata readily sprouts after severe disturbance and is long-lived.
Geographic Range: This association is known from moist, mid elevations in the inland Pacific Northwest. It is documented in Oregon, Idaho, Washington, British Columbia, Montana, and California.
Nations: CA?,US
States/Provinces: BC?, CA, ID, MT, OR, WA
Plot Analysis Summary:
http://vegbank.org/natureserve/ELEMENT_GLOBAL.2.683215
Confidence Level: Low
Confidence Level Comments: No Data Available
Grank: G4
Greasons: No Data Available
Type | Name | Database Code | Classification Code |
---|---|---|---|
Class | 2 Shrub & Herb Vegetation Class | C02 | 2 |
Subclass | 2.C Shrub & Herb Wetland Subclass | S44 | 2.C |
Formation | 2.C.4 Temperate to Polar Freshwater Marsh, Wet Meadow & Shrubland Formation | F013 | 2.C.4 |
Division | 2.C.4.Nb Western North American Temperate Freshwater Marsh, Wet Meadow & Shrubland Division | D031 | 2.C.4.Nb |
Macrogroup | 2.C.4.Nb.5 Sitka Alder - Booth''s Willow / Northwest Territory Sedge Montane Marsh, Wet Meadow & Shrubland Macrogroup | M893 | 2.C.4.Nb.5 |
Group | 2.C.4.Nb.5.d Willow species - Alder species - Water Birch Riparian & Seep Shrubland Group | G527 | 2.C.4.Nb.5.d |
Alliance | A3771 Gray Alder - Green Alder Wet Shrubland Alliance | A3771 | 2.C.4.Nb.5.d |
Association | CEGL001156 Sitka Alder / Common Ladyfern - Drooping Woodreed Wet Shrubland | CEGL001156 | 2.C.4.Nb.5.d |
Concept Lineage: No Data Available
Predecessors: No Data Available
Obsolete Names: No Data Available
Obsolete Parents: No Data Available
Synonomy: = Alnus sinuata / Athyrium filix-femina (Crowe and Clausnitzer 1997) [(p.122)]
= Alnus sinuata / Athyrium filix-femina Association (Kovalchik 1993) [(p.133)]
= Alnus sinuata / Cinna latifolia (Crowe and Clausnitzer 1997) [(p.124)]
? Alnus sinuata / Cinna latifolia association (Crowe et al. 2002)
? Alnus viridis ssp. sinuata / Gymnocarpium dryopteris association (Kovalchik 2001)
? Alnus viridis ssp. sinuata / Rubus spectabilis / Athyrium filix-femina association (Kovalchik 2001)
? Alnus viridis ssp. sinuata / Rubus spectabilis / Athyrium filix-femina association (Diaz and Mellen 1996) [apparently ranges from the eastern slope of the Cascades west to the Pacific coast and is distinguished by having moderate to high cover and constancy of Rubus spectabilis.]
< Alnus viridis ssp. sinuata community type (Hansen et al. 1995)
? Alnus viridis ssp. sinuata / Rubus spectabilis / Athyrium filix-femina (Murray 2000)
= Alnus viridus ssp. sinuata / Athyrium filix-femina Association (Crowe et al. 2004)
= Alnus sinuata / Athyrium filix-femina Association (Kovalchik 1993) [(p.133)]
= Alnus sinuata / Cinna latifolia (Crowe and Clausnitzer 1997) [(p.124)]
? Alnus sinuata / Cinna latifolia association (Crowe et al. 2002)
? Alnus viridis ssp. sinuata / Gymnocarpium dryopteris association (Kovalchik 2001)
? Alnus viridis ssp. sinuata / Rubus spectabilis / Athyrium filix-femina association (Kovalchik 2001)
? Alnus viridis ssp. sinuata / Rubus spectabilis / Athyrium filix-femina association (Diaz and Mellen 1996) [apparently ranges from the eastern slope of the Cascades west to the Pacific coast and is distinguished by having moderate to high cover and constancy of Rubus spectabilis.]
< Alnus viridis ssp. sinuata community type (Hansen et al. 1995)
? Alnus viridis ssp. sinuata / Rubus spectabilis / Athyrium filix-femina (Murray 2000)
= Alnus viridus ssp. sinuata / Athyrium filix-femina Association (Crowe et al. 2004)
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