Print Report

CEGL006545 Aronia arbutifolia - Ilex verticillata - Ilex mucronata / Osmunda cinnamomea Acidic Peatland

Type Concept Sentence: No Data Available


Common (Translated Scientific) Name: Red Chokeberry - Common Winterberry - Catberry / Cinnamon Fern Acidic Peatland

Colloquial Name: Allegheny Mountains Chokeberry - Winterberry Peatland

Hierarchy Level:  Association

Type Concept: This successional shrubland occurs on saturated, temporarily flooded, or semipermanently flooded acidic peatlands in the Allegheny Mountains region of West Virginia and Maryland, at elevations between 770 and 1210 m. It is a small-patch type that occupies flat to gently sloping land (0-2°) in headwater basins. Where conditions are favorable, individual stands may be as large as 12 ha. It is part of the natural disturbance mosaic of beaver-influenced vegetation types at high elevations in the Alleghenies, and also occurs on formerly forested peatlands that were logged or burned within the last 120 years. It is probably more widespread now than in the past, when forests covered much of its current habitat. Where natural succession is unhindered, some stands will likely recover to forested peatlands of high conservation value, although the recovery rate is slow (perhaps a century) in these frost-pocket habitats. Hummock-and-hollow microtopography is well-developed, with rounded peat-covered hummocks formed over woody stem clusters, decayed wood, and roots. Bedrock may be sandstone or, less commonly, limestone. Soil texture is moderately to very poorly drained peat, with a depth greater than one meter, although in earlier successional stands the peat may be only a few centimeters thick. Mean soil pH is 3.8. The unvegetated surface is predominantly litter, with an average of 5% standing water. This community may occur with either tall-shrub or short-shrub physiognomy, depending on the age of the stand. The shrub species favor hummock tops and typically include Aronia melanocarpa, Aronia arbutifolia, Viburnum nudum var. cassinoides, Vaccinium myrtilloides, Rhododendron maximum, Hypericum densiflorum, Ilex verticillata, Ilex mucronata, and Kalmia latifolia. Less frequent, but sometimes locally abundant, shrub species are Vaccinium corymbosum, Alnus incana ssp. rugosa, Vaccinium angustifolium, and Gaylussacia baccata. Dwarf-shrub species with high constancy include Rubus hispidus, Vaccinium oxycoccos, and Vaccinium macrocarpon. The herbaceous layer typically occupies hollows and the sides of hummocks. Species with high cover and constancy are Eriophorum virginicum, Solidago uliginosa, Carex folliculata, and Osmunda cinnamomea var. cinnamomea. Species with lower constancy that are sometimes abundant include Symplocarpus foetidus, Drosera rotundifolia var. rotundifolia, Gentiana linearis, Rhynchospora alba, Carex stipata, Juncus effusus, Lycopodium obscurum, Carex trisperma var. trisperma, Pteridium aquilinum, Gaultheria procumbens, Carex canescens, Carex gynandra, and Juncus brevicaudatus. Nonvascular plants are abundant, dominated by Sphagnum spp. and often including high cover by Polytrichum spp. Mean species richness of all vascular plants and any nonvascular plants with cover >1% is 21 taxa per 400 m2.

Diagnostic Characteristics: No Data Available

Rationale for Nominal Species or Physiognomic Features: No Data Available

Classification Comments: Sixty-three plots (10 occurrences) represent this type, which was classified as part of a 2004-2006 study of high-elevation wetlands in West Virginia''s Allegheny Mountains region (Byers et al. 2007). This type clusters well together considering that it is a fairly broad group. The ordination shows a fairly good grouping in the saturated, Sphagnum-rich portion of species space. The type has been adequately sampled throughout its range in West Virginia, and nine plots have been sampled in Maryland.

Similar NVC Types: No Data Available
note: No Data Available

Physiognomy and Structure: No Data Available

Floristics: This successional shrub peatland occurs in the Allegheny Mountains region of West Virginia and Maryland. It may occur with either tall-shrub or short-shrub physiognomy, depending on the age of the stand. The tall-shrub layer averages 20% cover, growing mostly on hummocks, and typically includes Aronia melanocarpa (= Photinia melanocarpa), Aronia arbutifolia (= Photinia pyrifolia), Viburnum nudum var. cassinoides, Vaccinium myrtilloides, Rhododendron maximum, Hypericum densiflorum, Ilex verticillata, Ilex mucronata (= Nemopanthus mucronatus), and Kalmia latifolia. Less frequent, but sometimes locally abundant, shrub species are Vaccinium corymbosum, Alnus incana ssp. rugosa, Vaccinium angustifolium, and Gaylussacia baccata. The short-shrub stratum averages 35% cover and is similar in composition to the tall-shrub layer. Dwarf-shrub species with high constancy include Rubus hispidus, Vaccinium oxycoccos, and Vaccinium macrocarpon. The herbaceous layer, with mean 60% cover, typically occupies hollows and the sides of hummocks. Species with high cover and constancy are Eriophorum virginicum, Solidago uliginosa, Carex folliculata, and Osmunda cinnamomea var. cinnamomea. Species with lower constancy that are sometimes abundant include Symplocarpus foetidus, Drosera rotundifolia var. rotundifolia, Gentiana linearis, Rhynchospora alba, Carex stipata, Juncus effusus, Lycopodium obscurum, Carex trisperma var. trisperma, Pteridium aquilinum, Gaultheria procumbens, Carex canescens, Carex gynandra, and Juncus brevicaudatus. Nonvascular plants average 70% cover, dominated by Sphagnum spp. (Sphagnum recurvum, Sphagnum fallax, Sphagnum magellanicum, Sphagnum papillosum, Sphagnum capillifolium (= var. capillifolium), Sphagnum flexuosum) and often including high cover by Polytrichum spp. (Polytrichum commune, Polytrichum juniperinum, Polytrichum strictum, Polytrichum ohioense). Mean species richness of all vascular plants and any nonvascular plants with cover >1% is 21 taxa per 400 m2 (Byers et al. 2007).

Dynamics:  This is a small-patch successional shrubland community. It is maintained by beaver activity, high water tables in flat-lying headwater basins, and rainfall. Nutrient cycling occurs from the decay of litter or woody vegetation.

Environmental Description:  This successional shrubland occurs on saturated, temporarily flooded, or semipermanently flooded acidic peatlands in the Allegheny Mountains region of West Virginia and Maryland, at elevations between 770 and 1210 m. It is a small-patch type that occupies flat to gently sloping land (0-2°) in headwater basins. Where conditions are favorable, individual stands may be as large as 12 ha. It is part of the natural disturbance mosaic of beaver-influenced vegetation types at high elevations in the Alleghenies, and also occurs on formerly forested peatlands that were logged or burned within the last 120 years. It is probably more widespread now than in the past, when forests covered much of its current habitat. Where natural succession is unhindered, some stands will likely recover to forested peatlands of high conservation value, although the recovery rate is slow (perhaps a century) in these frost-pocket habitats. Hummock-and-hollow microtopography is well-developed, with rounded peat-covered hummocks formed over woody stem clusters, decayed wood, and roots. Bedrock may be sandstone or less commonly limestone. Soil texture is moderately to very poorly drained peat, with a depth greater than one meter, although in earlier successional stands the peat may be only a few centimeters thick (Byers et al. 2007).

Geographic Range: This community is known from the Allegheny Mountains region of West Virginia and Maryland, at elevations between 770 and 1210 m.

Nations: US

States/Provinces:  MD, WV




Confidence Level: Moderate

Confidence Level Comments: No Data Available

Grank: GNR

Greasons: No Data Available


Concept Lineage: No Data Available

Predecessors: No Data Available

Obsolete Names: No Data Available

Obsolete Parents: No Data Available

Synonomy: = Photinia (melanocarpa, pyrifolia) - Viburnum nudum var. cassinoides / Eriophorum virginicum / Sphagnum spp. Shrub Peatland (Byers et al. 2007)
< Polytrichum - Pyrus melanocarpa - Vaccinium myrtilloides hummock bog (Fortney 1975)
= Polytrichum - shrub community (Wieder et al. 1981)
< Pyrus melanocarpa - Vaccinium myrtilloides - Hypericum densiflorum low shrub community (Robinette 1964)
= Viburnum cassinoides - Rubus hispidus - Pyrus melanocarpa hummock community (Edens 1973)

Concept Author(s): Eastern Ecology Group

Author of Description: E.A. Byers

Acknowledgements: No Data Available

Version Date: 05-11-07

  • Byers, E. A., J. P. Vanderhorst, and B. P. Streets. 2007. Classification and conservation assessment of high elevation wetland communities in the Allegheny Mountains of West Virginia. West Virginia Natural Heritage Program, West Virginia Division of Natural Resources, Elkins.
  • CVI [Canaan Valley Institute]. 2005. Wetland assessment. Unpublished report.
  • Eastern Ecology Working Group of NatureServe. No date. International Ecological Classification Standard: International Vegetation Classification. Terrestrial Vegetation. NatureServe, Boston, MA.
  • Edens, D. L. 1973. The ecology and succession of Cranberry Glades, WV. Ph.D. dissertation, North Carolina State University, Raleigh.
  • Fortney, R. H. 1975. The vegetation of Canaan Valley, West Virginia: A taxonomic and ecological study. Ph.D. dissertation, University of West Virginia, Morgantown.
  • Robinette, S. L. 1964. Plant ecology of an Allegheny mountain swamp. M.S. thesis, West Virginia University, Morgantown.
  • WVNHP [West Virginia Natural Heritage Program]. No date. Unpublished data. West Virginia Natural Heritage Program, Elkins.
  • Walbridge, M. R. 1982. Vegetation patterning and community distribution in four high-elevation headwater wetlands in West Virginia. M.S. thesis, West Virginia University, Morgantown.
  • Wieder, R. K., A. M. McCormick, and G. E. Lang. 1981. Vegetational analysis of Big Run Bog, a nonglaciated sphagnum bog in West Virginia. Castanea 46:16-29.