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A3132 Pinus cembroides - Pinus discolor - Pinus edulis / Grass Understory Woodland Alliance

Type Concept Sentence: This Madrean open-canopy pinyon-juniper woodland and savanna alliance is composed of Pinus cembroides or Pinus discolor often with Juniperus coahuilensis or Juniperus deppeana and a perennial grass-dominated understory with only sparse cover shrubs. It occurs in foothills and lower montane zones of southeastern Arizona and southern New Mexico, extending into western Texas and Mexico.


Common (Translated Scientific) Name: Mexican Pinyon - Border Pinyon - Two-needle Pinyon / Grass Understory Woodland Alliance

Colloquial Name: Madrean Pinyon - Juniper / Grass Woodland

Hierarchy Level:  Alliance

Type Concept: This Madrean pinyon-juniper woodland alliance occurs in foothills and lower montane zones of southeastern Arizona and southern New Mexico, extending into western Texas and Mexico. The vegetation is characterized by a very open (wooded savanna) to moderately dense tree layer (10-70% cover) composed of pinyon and juniper trees 3-5 m tall. Pinus cembroides, Pinus discolor, or Pinus remota are diagnostic and often dominant species. Juniper trees may be absent to dominant. Juniperus coahuilensis, Juniperus deppeana, or Juniperus pinchotii are differential species. In more northern stands, Pinus edulis and Juniperus monosperma may be present to dominant when diagnostic Madrean species are present. Evergreen oaks such as Quercus arizonica, Quercus emoryi, or Quercus grisea are often present in the tree canopy but have lower cover than conifers. Scattered shrubs may be present but typically do not form a layer. The herbaceous layer dominates the understory with moderate to dense cover (25-80% cover) of perennial graminoids. Characteristic species include Bouteloua eriopoda, Eragrostis intermedia, Muhlenbergia dubia, Muhlenbergia emersleyi, Muhlenbergia montana, Muhlenbergia pauciflora, Muhlenbergia setifolia, Nassella tenuissima, Piptochaetium fimbriatum, Piptochaetium pringlei, and Schizachyrium cirratum. Forbs are typically sparse. Some of the more common forbs include species of Ageratina, Artemisia, Cheilanthes, Chenopodium, Geranium, Hedeoma, Senecio, Thalictrum, and Viguiera. Stands occur on gently to moderately sloping alluvial plains, bajadas and canyon bottoms to moderately steep slopes in canyons, piedmont hills and mountains. Elevation ranges from 1460-2010 m (4780-6600 feet). Soils are shallow, gravelly or stony sandy loams or sandy clay loams, derived from a mixture of alluvium and colluviums mostly associated with igneous substrates.

Diagnostic Characteristics: This alliance includes wooded savannas and open woodlands with a grassy understory. The presence of pinyon trees and abundant cover of perennial graminoids (>25% cover) is diagnostic. Junipers trees are often present to dominant. Diagnostic trees species are Pinus cembroides, Pinus discolor, Pinus remota, Juniperus coahuilensis, Juniperus deppeana, or Juniperus pinchotii. Juniperus monosperma may also be present but is a widespread species and not diagnostic. Pinus edulis may be a dominant species when diagnostic Madrean species are present. Diagnostic Madrean species include Dasylirion leiophyllum, Mimosa aculeaticarpa var. biuncifera, and Nolina microcarpa. The understory may have scattered shrubs present but is characterized by moderate to dense cover (25-80% cover) of perennial graminoids. Characteristic species include Eragrostis intermedia, Muhlenbergia dubia, Muhlenbergia emersleyi, Muhlenbergia pauciflora, Muhlenbergia setifolia, Nassella tenuissima, Piptochaetium fimbriatum, Piptochaetium pringlei, and Schizachyrium cirratum.

Rationale for Nominal Species or Physiognomic Features: No Data Available

Classification Comments: Because of past management actions such as overgrazing by livestock, some stands or portions of stands may have a patchy herbaceous layer with <25% cover of perennial grasses. However, shrub are sparse or absent.

Confusion over the taxonomic treatments of Pinus discolor D.K. Bailey & Hawksworth and related taxa Pinus cembroides Zucc. and Pinus remota (Little) D.K. Bailey & Hawksworth has caused confusion in the classification of this alliance. USDA PLANTS Database (USDA NRCS 2013) and Kartesz (1999) both use Bailey and Hawksworth''s (1979) treatment and list the above three names as accepted species. However, Kral''s treatment in Flora of North America (Kral 1993) is based on Little (1971) and includes Pinus remota and Pinus discolor within Pinus cembroides Zucc. (var. remota Little and var. bicolor Little, respectively). Currently, Pinus remota woodlands have been described from the Glass and Del Norte mountains of the Trans-Pecos and the Edwards Plateau in central Texas. Pinus cembroides stands have been reported from the Chicos and Davis mountains of the Trans-Pecos, extending into Coahuila, Mexico. Pinus discolor woodlands have been reported from several mountain ranges in southwestern New Mexico and southeastern Arizona below the Mogollon Rim and include the Animas, Peloncillos, Santa Catalina, Chiricahua, Mule, Dragoon, and Huachuca mountains. The woodlands are predicted to occur throughout the Borderlands of New Mexico, Arizona and Mexico (Muldavin et al. 1998a). It has been suggested that the Pinus discolor associations be renamed Pinus cembroides, but it is not known whether the trees in these associations were misidentified or if there is a problem with the current taxonomy.

Similar NVC Types: No Data Available
note: No Data Available

Physiognomy and Structure: Vegetation included in this alliance has an open-tree canopy that is typically 5-10 m tall, but may reach 15 m. A sparse to moderately dense tree subcanopy (3-10 m tall) is typically present. The canopy is dominated by evergreen needle-leaved trees with the subcanopy dominated by broad-leaved or scale-leaved evergreen trees. Broad-leaved deciduous trees may be present in some stands. Scattered shrubs may be present but do not form a layer. An open to dense herbaceous layer is diagnostic of this alliance and is dominated by perennial graminoids. Perennial forbs have sparse cover. Annual forbs and grasses may be seasonally present.

Floristics: This alliance is characterized by a very open (wooded savanna) to moderately dense tree layer (10-70% cover) composed of pinyon and juniper trees 3-5 m tall. Pinus cembroides, Pinus discolor, or Pinus remota are diagnostic and often dominant species. Juniper trees may be absent to dominant. Juniperus coahuilensis, Juniperus deppeana, Juniperus flaccida, or Juniperus pinchotii are differential species. In more northern stands, Pinus edulis and Juniperus monosperma may be present to dominant when diagnostic Madrean species are present. Evergreen oaks such as Quercus arizonica, Quercus emoryi, or Quercus grisea are often present in the tree canopy, but have lower cover than conifers. Scattered shrubs may be present but typically do not form a layer. The herbaceous layer dominates the understory with moderate to dense cover (25-80% cover) of perennial graminoids. Characteristic herbaceous species include Bouteloua eriopoda, Eragrostis intermedia, Muhlenbergia dubia, Muhlenbergia emersleyi, Muhlenbergia pauciflora, Muhlenbergia setifolia, Nassella tenuissima, Piptochaetium fimbriatum, Piptochaetium pringlei, and Schizachyrium cirratum. Other common graminoids include Aristida schiedeana var. orcuttiana (= Aristida orcuttiana), Bouteloua curtipendula, Bouteloua gracilis, Cyperus fendlerianus, Koeleria macrantha, Muhlenbergia longiligula, Muhlenbergia montana, and Muhlenbergia tenuifolia (= Muhlenbergia monticola). Forbs are typically sparse. Some of the more common forbs include Ageratina herbacea (= Eupatorium herbaceum), Artemisia ludoviciana, Artemisia carruthii, Cheilanthes fendleri, Dysphania graveolens (= Chenopodium incisum), Geranium caespitosum, Hedeoma hyssopifolia, Packera neomexicana (= Senecio neomexicanus), Thalictrum fendleri, and Viguiera spp.

Dynamics:  Stands are typically small and naturally patchy, and reflect the late-successional stage of Sierra Madrean woodlands and savannas. This is a highly diverse alliance with a preponderance of Sierra Madrean elements that have a restricted distribution in the U.S. Grazing and, to a lesser degree, fuelwood harvest continue to impact stands. Altered fire regimes (failed fire suppression followed by increased intensity and extent of fire) may be reducing the number of stands.

Pinyon trees are drought-tolerant and slow-growing (Little 1987, Powell 1988b, Muldavin et al. 1998a). They are also non-sprouting and may be killed by fire (Wright et al. 1979). The effect of fire on a stand is largely dependent on the tree height and density, fine-fuel load on the ground, weather conditions, and season (Dwyer and Pieper 1967, Wright et al. 1979). Trees are more vulnerable in open stands where fires frequently occur in the spring, the relative humidity is low, wind speeds are over 10-20 mph, and there are adequate fine fuels to carry fire (Wright et al. 1979). Under other conditions, burns tend to be spotty with low tree mortality. Large trees are generally not killed unless fine fuels, such as tumbleweeds, have accumulated beneath the tree to provide ladder fuels for the fire to reach the crown (Jameson et al. 1962). Closed-canopy stands rarely burn because they typically do not have enough fine fuels in the understory or wind to carry a fire (Wright et al. 1979).

Although pinyon trees are drought-tolerant, prolonged droughts will weaken trees and promote mortality by secondary agents. Periodic die-offs of pinyon pine caused by insects, such as the pinyon ips beetle (Ips confusus), or fungal agents, such as blackstain root-rot (Leptographium wageneri), tend to be correlated with droughts (Anhold 2005). These mortality events may be localized or widespread but can result in 50 to 90% mortality of Pinus edulis (Harrington and Cobb 1988).

In addition, altered fire regimes, cutting trees for fencing or firewood, and improper grazing by livestock have significant impacts on the quality of sites. Grazing by livestock can modify the fire regime by removing the fine fuels that carry fire. Fire, livestock grazing, and trampling by recreationalists and vehicles disturb cryptogamic soil crusts that help maintain soil structure, reduce soil erosion, provide habitat for plants and preserve biological diversity (Ladyman and Muldavin 1996). More study is needed to understand and manage these woodlands ecologically. Climatic and other factors have resulted in denser and expanded pinyon-juniper stands and has likely become denser in much of the Madrean region. Denser stands are more susceptible to attack by insects and disease (Anhold 2005).

Environmental Description:  This Madrean pinyon-juniper woodland alliance occurs in foothills and lower montane zones of southeastern Arizona and southern New Mexico, extending into western Texas and Mexico. Elevation ranges from 1460-2010 m (4780-6600 feet). Stands occur on gently to moderately sloping alluvial plains, bajadas and canyon bottoms to moderately steep slopes in canyons, piedmont hills, mesas and mountains. Climate is semi-arid. Summers are generally hot and winters mild with cold periods and occasional snows. The mean annual precipitation ranges from 45-50 cm, mostly occurring during the Arizona monsoon season from July to September. Soils are shallow, gravelly or stony sandy loams or sandy clay loams, derived from a mixture of alluvium and colluvium. Aspect is variable, but stands are typically on the less xeric north slopes. Soils are shallow, lithic, gravelly loams derived from colluvium and residuum, but include deeper loamy soils, especially on sites where trees have recently colonized. Parent materials include alluvium, andesite, limestone, rhyolite and tuff.

Adjacent vegetation at higher elevations is typically woodlands or forests dominated by Pinus ponderosa, Pinus engelmannii, or Pinus leiophylla with a subcanopy of Quercus spp. Adjacent vegetation at lower elevations is often woodlands or savannas dominated by Pinus discolor, Pinus edulis, or Juniperus spp. or chaparral dominated by Arctostaphylos spp. or Quercus turbinella.

Geographic Range: This Madrean pinyon-juniper savanna and woodland group occurs in lower foothills and plains of southeastern Arizona and southern New Mexico, extending into western Texas and northern Mexican states of Chihuahua and Sonora.

Nations: MX,US

States/Provinces:  AZ, MXCHH, MXCOA, MXSON, NM, TX




Confidence Level: Low

Confidence Level Comments: No Data Available

Grank: GNR

Greasons: No Data Available


Concept Lineage: In part A.392, A.510, A.516, A.523, and A.538. This alliance was created from associations with perennial grass dominated understory selected from the following Old Alliances II.A.4.N.a. Pinus cembroides Woodland Alliance (A.510), II.A.4.N.a. Pinus discolor Woodland Alliance (A.538), II.A.4.N.a. Pinus edulis - (Juniperus spp.) Woodland Alliance (A.516).

Predecessors: No Data Available

Obsolete Names: No Data Available

Obsolete Parents: No Data Available

Synonomy: > Pinus discolor / Piptochaetium fimbriatum Woodland (Bassett et al. 1987)
> Pinus discolor-Nolina microcarpa-Muhlenbergia emersleyi Habitat Type (Willing 1987) [Need to review reference to determine if this type has a significant shrub layer. Relationship uncertain.]
> Pinus discolor/Muhlenbergia emersleyi Habitat Type (Bassett et al. 1987) [relationship uncertain]
> Pinus edulis / Muhlenbergia pauciflora Plant Association (Kennedy 1983a)
< Colorado Pinyon-Alligator Juniper Series (Dick-Peddie 1993) [relationship uncertain]
> Pinyon Pine/New Mexico Muhly PA (Muldavin et al. 2000b)

Concept Author(s): K.A. Schulz, in Faber-Langendoen et al. (2013)

Author of Description: K.A. Schulz

Acknowledgements: No Data Available

Version Date: 01-08-14

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